Parena (Crossoglossa) levata Andrewes, 1931

Shi, Hongliang & Liang, Hongbin, 2023, Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina), Zootaxa 5286 (1), pp. 1-144 : 36

[3] Parena (Crossoglossa) levata Andrewes, 1931

Habitus: Figs 17E, 17F .

Andrewes, 1931a: 77 (type locality: Sumatra: Sibolangit ; holotype in NHML); Jedlička, 1952: 210 .

Type material examined. Parena levata Andrewes : Holotype (NHML, Fig. 17E ), female, body length = 9.6 mm, " 8.x.1925 / Sibolangit / Kamy ", "Type"[red label], " Parena / levata /Type Andr. /H.E. Andrewes det.", "H.E. Andrewes Coll. / B.M. 1945-97".

Non-type material examined. 1 female (IZAS), "Boreno: Sahah, Keningau district, Jungle Girl Camp. N5.4430, E116.4512, 1182 m; Shi H.L. & Liu Y. lgt. light trap, Ins. Zoo., CAS. 2016.IV.29 N" < Figs 14C , 17F >.

Comparisons. P. levata is distinguishable from the other two species in this species group by its elytra disc not depressed near the middle of intervals 3 to 6. Despite this, the external features of P. levata is almost identical to specimens of P. laesipennis from Southeast Asia.

Description. Body length 9.5–9.6 mm. Dorsum reddish brown; basal three antennomeres, and basal half of antennomere 4 brown, rest parts of antennae black; venter brown. Pronotum strongly transverse, PW/PL = 1.55–1.60, slightly wider than head, PW/HW = 1.05–1.07; lateral explanations very wide. Elytra without microsculpture; striae not incised, replaced by lines of fine punctures; intervals completely flat, very finely and sparsely punctate; disc not depressed near middle of intervals 3 to 6; lateral sides depressed near anterior third shallower than in other species; apical truncation distinct, straight, outer apical angles weakly angulate, forming very blunt obtuse angles; sutural angles sharply pointed, forming short denticles. Male genitalia unknown. Gonocoxite II of ovipositor dichotomous, inner and outer branches each with two ensiform setae apically, setae on outer branch much shorter than those on the inner one.

Distribution ( Map 3 , blue). Sumatra, Borneo (Sabah).

Figures

MAP 3. Distributions for species in the Parena (Crossoglossa) laesipennis group.

FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong.

FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm.