Agnosthaetus rodmani Clarke

Clarke, Dave J., 2011, A Revision of the New Zealand Endemic Rove Beetle Genus Agnosthaetus Bernhauer (Coleoptera: Staphylinidae), The Coleopterists Bulletin (mo 10) 2011, pp. 1-118: 24-26

publication ID

http://doi.org/ 10.1649/0010-065X-65.mo4.1

publication LSID

lsid:zoobank.org:pub:0818A3A2-AB42-43D8-8F76-4F65F367C584

persistent identifier

http://treatment.plazi.org/id/EC6E9FBC-424A-4C49-9EBE-33CF69D6B11F

taxon LSID

lsid:zoobank.org:act:EC6E9FBC-424A-4C49-9EBE-33CF69D6B11F

treatment provided by

Carolina

scientific name

Agnosthaetus rodmani Clarke
status

new species

(4) Agnosthaetus rodmani Clarke   , new species

( Figs. 11 View Figs , 66 View Figs , 70 View Figs , 81 View Figs , 92 View Figs , 97 View Figs , 101 View Figs , Map 1 View Map 1 )

Type Material. Holotype. ♂, with four labels: “ NEW ZEALAND [ South Island ], WD| Totara Valley | Weir Ck 122 m | 10 Jun 1983 | H.P. McColl / Litter| 11/83/ FMNH-INS 0000 048 042 / HOLOTYPE Agnosthaetus rodmani Clarke   , ♂, design. D. Clarke 2011”, in NZAC   . Paratypes. 39 specimens (21♂, 18♀). NEW ZEALAND: South Island: WD: Fox Glacier , 43°28.062′S, 170°1.188′E [coord.=town], 12.xi.1968, 68/171, litter, J.I. Townsend GoogleMaps   , 1♀, FMNH-INS 48043 (in NZAC)   ; Fox Glacier, Minehaha Walk , 180 m, 43°28.56′S, 170°0.72′E, 12.ii.2007, moss & litter, J.T. Nunn GoogleMaps   , 1♂, FMNH-INS 19308 (in JTNC)   ; Franz Josef Glacier, Head of Alex Knob (2.7 km S), 150 m, 43°25′S, 170°10′E, coastal forest, 15.i.1998, NZEA 1L C98 063, berl., leaf litter, R. Leschen & C. Carlton GoogleMaps   , 1♂, KUNHM-ENT SM0365078 (in KSEM)   ; Lake Mahinapua , 20 m, 42°47.706′S, 170°55.002′E, punga-broadleaf forest, 17.i.1982, sweeping ferns/ ground cover, J.W. Early GoogleMaps   , 1♀, FMNH-INS 19084 (in LUNZ)   ; Lake Mahinapua, Jum Michel Trk. , 42°47′S, 170°55′E, 12.v.1999, RL 305, Berl., leaf litter/fungusy logs, R. Leschen & R. Hoare GoogleMaps   , 1♂ [date as 12-II-1999, i.e., not May], KUNHM-ENT SM0673914 (in KSEM)   ; 1♂, FMNH-INS 48041 (in NZAC)   ; Mananui Bush, Ruatapu (3 km NNE), 10 m, ANMT 733, 42°47′S, 170°54′E, coastal hdwd- Dicksonia   dune forest, 9.i.1985, FMHD#85-466, litter, forest, Berl., A. Newton & M. Thayer GoogleMaps   , 4♂ 1♂, FMNH-INS 48044 (in FMNH)   ; Mt. Hercules Scen. Res., Harihari (12.6 km WSW), 75 m, ANMT 734, 43°11′S, 170°27′E, hardwood-podocarp forest, 9.i.1985, FMHD#85-467, litter, forest, Berl., A. Newton & M. Thayer GoogleMaps   , 1♀, FMNH-INS 48045 (in FMNH)   ; Waiho R. Valley , 200 m, 6.ii.1984, leaf litter, P.M. Hammond   , 11♂, 11♀ FMNH-INS 19044, FMNH- INS 38176 View Materials –185, 187–197 (in BMNH)   ; 6.ii.1984, Nothofagus   forest litter, P.M. Hammond, 3♀, FMNH-INS 38186, 198–199 (in BMNH)   ; Westland N.P., adj. Canavans Knob , 140 m, 43°22.968′S, 170°9.828′E, totara/matai forest, 4.ii.1982, leaf litter in totara/matai forest, A.B. Miller GoogleMaps   , 1♀, FMNH- INS 19082 View Materials (in LUNZ)   ; Westland N.P., Fox Glacier (8.8 km NE), 315 m, ANMT 742, 43°26′S, 170°5′E, low hardwood forest, 18.i.1985, FMHD#85-475, litter, forest, Berl., A. Newton & M. Thayer GoogleMaps   , 1♂, FMNH-INS 48047 (in FMNH)   .

Diagnosis. Agnosthaetus rodmani   can be distinguished from all other Agnosthaetus species   by the combination of the distinct reticulate microsculpture on the dorsum of the head ( Fig. 11 View Figs ), the basal mental tooth ( Fig. 66 View Figs , arrow), the distinctive chromosome-shaped medial pronotal sulci with the apices deflected obliquely and separate from the anterior punctures ( Fig. 70 View Figs , ap), and the complete lateral pronotal carina (continuous with anterior pronotal margin: Fig. 81 View Figs , pc).

Description. Color: More or less uniformly yellowish to reddish brown, occasionally with darker elytra and terminal abdominal segments. Head ( Fig. 11 View Figs ): Frontal ridge present (cf. Fig. 12 View Figs , fr). Dorsum sparsely punctate; with punctures distributed over more or less entire surface except for small posterior impunctate region. Punctures deep, but indistinctly defined; diameter subequal to or slightly greater than diameter of eye facet; interpuncture distance mostly less than half puncture diameter. Dorsal microsculpture present on entire or most of surface; distinctly reticulate ( Fig. 11 View Figs ). Dorsal tentorial sulcus (cf. Figs. 10–11 View Figs , dt) distinctly slit-like, straight, distinctly impressed between sublongitudinal ridge and medial part of vertex ( Fig. 11 View Figs , dt). Sublongitudinal ridge (cf. Fig. 10 View Figs , sr) indistinct; confused by dorsolateral secondary carinae or punctures; crest at antennal tubercle with distinct microsculpture. Area above and behind antenno-ocular carina ( Figs. 10–11 View Figs , arrow) with several secondary carinae formed by subconfluent to confluent punctures. Antennoocular carina (cf. Fig. 10 View Figs , ao) indistinct; confused by secondary carinae and sculpture; joining eye at or in front of middle ( Fig. 11 View Figs , ao). Eye distinctly bulged anteriorly ( Fig. 11 View Figs ). Temple ( Fig. 11 View Figs , tm) very short, much less than 50% EYL. Suboccular surface more or less evenly microsculptured (cf. Fig. 65 View Figs ). Labrum not distinctly sexually dimorphic ( Fig. 92 View Figs ). Apical labral margin in males moderately broadly and shallowly emarginate medially, evenly dentate, with 20–21 teeth (n =7), and with medial tooth equal in length to, but approximately 3X width of, paramedial teeth. Apical labral margin in females very slightly concave medially; with 21–25 teeth (n =9), all teeth subequal in length. Medial oblique labral setae (cf. Fig. 20 View Figs ) absent or present. Adoral labral surface in males smooth, without subapical transverse ridge. Mandible sexually dimorphic; males with single, dorsally directed tooth, with distinct preapical spur (cf. Fig. 190 View Figs , arrow); females with single, mesially projecting tooth, without spur. Mentum with distinct basomedian tooth ( Fig. 66 View Figs , arrow). Prothorax: Pronotum without microsculpture. Medial pronotal sulci (cf. Fig. 23 View Figs ) distinctly chromosome-shaped, abruptly deflected obliquely in apical third; anteriorly separate from, and deflected obliquely to anterior punctures ( Fig. 70 View Figs , ap). Distance between medial sulci in front of middle about half that at base. Pronotal basolateral carina absent (cf. Figs. 23 View Figs , 69 View Figs , 77 View Figs ). Anterior pronotal puncture (cf. Fig. 70 View Figs , ap) distinct; medial puncture (cf. Fig. 70 View Figs , mu) distinct ( Fig. 70 View Figs , mu); basal puncture ( Fig. 70 View Figs , bu) indistinct. Medial pronotal seta adjacent to lateral sulcus ( Fig. 70 View Figs , mu). Lateral pronotal carina, anteriorly complete, continuous with anterior pronotal margin but often less distinct anteriorly than posteriorly ( Fig. 81 View Figs , pc). Pronotal hypomeron ( Fig. 24 View Figs , hy) shiny, without microsculpture. Prosternum with distinctly reticulate microsculpture. Pterothorax: Elytron ( Fig. 23 View Figs , e) without microsculpture; with large, scattered, punctiform, non-setiferous impressions, or with small, punctiform, non-setiferous impressions; with 2 or 3 three macrosetae set in distinct punctures; laterally with single ridge (cf. Fig. 85 View Figs , ek). Elytral epipleuron ( Fig. 24 View Figs , ef) very narrow, more or less evenly wide for most of length. Mesothoracic epimeral region ( Fig. 24 View Figs , mer) with at most faint microsculpture. Metathoracic pleural region ( Fig. 24 View Figs , m) with faint reticulate microsculpture. Metathoracic pleural ridge absent; metathoracic pleural groove ( Fig. 24 View Figs , gr) complete, continuing to or near to pleurocoxal articulation. Abdomen: Abdominal vestiture short, somewhat appressed; vestiture dorsally with radiating fans of setae either side of midline and laterally. Abdominal sternite IV of male with surface glabrous and slightly impressed apicomedially; V with surface glabrous and impressed apicomedially, flanked by diffuse coarser acuminate setae, apex slightly sinuate; VI with dense patch of coarse setae apicomedially, apex slightly convex; VII with surface nearly glabrous apicomedially. Aedeagus ( Fig. 97 View Figs ): “ Type A” (see description on p. 8). Apical part of median lobe with small lateral denticles, abruptly produced into broadly triangular, sharply pointed apicomedian lobe. Both apicolateral and apicomedial setae short ( Fig. 101 View Figs ). Paramere exceeding apex of median lobe; in lateral view produced apically into lobe; with apical part perpendicular to median lobe; in dorsal view with outer side irregularly curving; and with ventral excavation at about middle of mesal face ( Fig. 97 View Figs , arrow); with single seta at apex, 1 on mesial face, and 2 on ventral edge.

Etymology. The epithet rodmani   is a noun in the genitive case, honoring Dr. James Rodman, formerly of the U.S. National Science Foundation, and principal architect and supporter of the National Science Foundation’ s Partnerships for Enhancing Expertise in Taxonomy (PEET) program. Support from that program made the present monograph possible.

Distribution. ( Map 1 View Map 1 ). South Island: WD.

Biology and Ecology. Habitat: diverse forested vegetation types, including sand dune forest. Specimens have been collected from forest leaf and log litter, by sweeping ferns and ground plants, and from mossy litter. Phenology: Year-round; most specimens taken in April (but from a single collecting event). Elevation: 10–315 m.

Remarks. Agnosthaetus rodmani   is morphologically closest to A. bisulciceps   . Both species have distinct tooth-like apicolateral lobes of the median lobe ( Figs. 96–97 View Figs , 100–101 View Figs ), but these are much smaller in A. rodmani   , and the parameres extend much further beyond the median lobe apex than in A. bisulciceps   . See also Remarks under A. bisulciceps   .

NZAC

New Zealand Arthropod Collection

LUNZ

Lincoln University Entomology Research Museum

FMNH

Field Museum of Natural History