Agnosthaetus lanceolatus Clarke, 2011

Clarke, Dave J., 2011, A Revision of the New Zealand Endemic Rove Beetle Genus Agnosthaetus Bernhauer (Coleoptera: Staphylinidae), The Coleopterists Bulletin (mo 10) 2011, pp. 1-118 : 56-57

publication ID	https://doi.org/ 10.1649/0010-065X-65.mo4.1
publication LSID	lsid:zoobank.org:pub:0818A3A2-AB42-43D8-8F76-4F65F367C584
persistent identifier	https://treatment.plazi.org/id/038787B5-FFA8-537F-44EA-BF58803DFD00
treatment provided by	Carolina
scientific name	Agnosthaetus lanceolatus Clarke
status	sp. nov.

Treatment

(24) Agnosthaetus lanceolatus Clarke View in CoL , new species

( Figs. 60 View Figs , 68 View Figs , 74 View Figs , 83 View Figs , 147 View Figs , 157 View Figs , 160 View Figs , Map 5 View Map 5 )

Type Material. Holotype. ♂, with four labels: “New Zealand WN| Tararua Ra | Logan East Basin [40°44′S, 175°28′E] 12 Feb 1985 | C.F.Butcher / Moss and| liverworts| 85/28/ FMNH-INS 0000 038 336 / HOLOTYPE Agnosthaetus lanceolatus Clarke , ♂, design. D. Clarke 2011”, in NZAC GoogleMaps . Paratypes. 26 specimens (16♂ 10♀). NEW ZEALAND: North Island: WN: Tararua Ra., Dundas Hut, 1250 m, 40°43′S, 175°28′E, 13.ii.1985, 85/25, litter & tussock, B.A. Holloway GoogleMaps , 3♂, FMNH-INS 38326–328 (in NZAC) ; Tararua Ra., Dundas Hut Ridge , 950 m, 40°43.08′S, 175°27.9′E, 3.xii.1984, 84/87, sifted litter, R.C. Craw GoogleMaps , 1♂, 1♀, FMNH-INS 38322–323 (in NZAC) ; Tararua Ra., Dundas Hut Ridge , 990 m, 40°43.08′S, 175°27.9′E, 3.xii.1984, 84/83, sifted litter, B.G. Bennett & T.K. Crosby GoogleMaps , 1♂, FMNH-INS 38319 (in NZAC) ; 3.xii.1984, 84/85, sifted litter, B.G. Bennett & T.K. Crosby, 2♂, 1♀, FMNH-INS 38316–317, 318 (in NZAC) ; 13.ii.1985, 85/20, sifted litter, C.F. Butcher, 1♂, 1♀, FMNH-INS 38320, 321 (in NZAC) ; Tararua Ra., Dundas Hut Ridge, River Forks , 700 m, 40°43.086′S, 175°27.954′E [coord.=Dundas Hut], 11.ii.1985, 85/13, litter, G.W. Ramsay GoogleMaps , 1♂, 1♀, FMNH-INS 38330, 331 (in NZAC) ; Tararua Ra. , Dundas Ridge, E side, 1350 m, 40°43.578′S, 175°27.414′E [coord.=Dundas Ridge], 6.xii.1984, 84/92, turf, plants, moss, J.S. Dugdale GoogleMaps , 2♀, FMNH-INS 38324–325 (in NZAC) ; Tararua Ra., Dundas Ridge , west slope, 1430 m, 40°43.578′S, 175°27.414′E [coord.=Dundas Ridge], 13.ii.1985, 85/24, mixed swards, C.F. Butcher GoogleMaps , 1♂, FMNH-INS 38329 (in NZAC) ; Tararua Ra., Logan , 1490 m, 40°43.332′S, 175°27.408′E [coord.=Logan Hill], 2.xii.1984, 84/91, Swards, moss, plants, J.S. Dugdale GoogleMaps , 2♂, FMNH-INS 38332–333 (in NZAC) ; Tararua Ra., Logan Basin , 1300 m, 40°44′S, 175°28′E [coord.=“ Logan E Basin”, taken from McLellan 1996], 6.xii.1984, 84/96, turf. Plants, R.C. Craw GoogleMaps , 1♂, FMNH-INS 38334 (in NZAC) ; Tararua Ra., Logan E. Basin , 40°44′S, 175°28′E, 12.ii.1985, 85/28, moss & liverworts, C.F. Butcher GoogleMaps , 1♂, FMNH-INS 38335 (in NZAC) ; Tararua Ra., Mt. Dome , 40°46.002′S, 175°22.998′E, 10.ii.1985, 85/10, moss, liverworts and mixed plants, G.W. Ramsay GoogleMaps , 1♂, FMNH-INS 38337 (in NZAC) ; 10.ii.1985, 85/9, moss, mixed swards & tussock, G.W. Ramsay, 1♂, FMNH-INS 38338 (in NZAC) ; Tararua Ra., Mt. Pukemoremore , 1474 m, 40°42.72′S, 175°28.02′E, 12.ii.1985, 85/30, mixed swards, C.F. Butcher GoogleMaps , 1♀, FMNH-INS 38339 (in NZAC) ; Tararua Ra., Pukemoremore summit, 1474 m, 40°43.002′S, 175°28.002′E, 12.ii.1985, 85/33, moss and lichen, B.A. Holloway GoogleMaps , 1♀, FMNH- INS 38340 (in NZAC) ; Tararua Ra., Ridge to Dundas Hut , 1275 m, 40°43.998′S, 175°27′E, 6.xii.1984, 84/97, moss, R.C. Craw GoogleMaps , 1♀, FMNH- INS 38314 (in NZAC) ; Tararua Ra., Ridge to Dundas Hut , 1100 m, 40°43′45″S, 175°28′25″E, 6.xii.1984, 84/95, moss, R.C. Craw GoogleMaps , 1♀, FMNH- INS 38315 (in NZAC) .

Diagnosis. Agnosthaetus lanceolatus is the most distinctive species on North Island in being the only one with microsculpture on the head and thorax. It can be easily separated from all other known species by the combination of that feature and the distinct basal impression of the mentum ( Fig. 68 View Figs ).

Description. Color: Variably reddish brown, occasionally with darker pigmented abdominal segments. Head ( Figs. 60 View Figs , 68 View Figs ): Frontal ridge absent. Dorsum sparsely punctate; with punctures distributed anteriorly, laterally, and posteriorly on disc, middle part impunctate. Punctures shallow, rather indistinct; diameter subequal to or slightly greater than diameter of eye facet; interpuncture distance approximately 0.5–1.0X puncture diameter. Dorsal microsculpture present on entire or most of surface; distinctly reticulate. Dorsal tentorial sulcus (cf. Figs. 10–11 View Figs , dt) narrowly ovate; width subequal to or slightly greater than puncture diameter. Sublongitudinal ridge (cf. Fig. 10 View Figs , sr) distinct; not confused by smaller carinae or punctures ( Fig. 60 View Figs , sr); crest at antennal tubercle with distinct microsculpture. Area above and behind antennoocular carina ( Figs. 10–11 View Figs , arrow) usually without, or else only indistinct, subsidiary carinae.Antenno-ocular carina joining eye at about middle ( Fig. 60 View Figs ). Temple ( Fig. 11 View Figs , tm) short, less than 50% EYL. Subocular surface more or less evenly microsculptured (cf. Fig. 65 View Figs ). Labrum distinctly sexually dimorphic ( Fig. 147 View Figs ). Apical labral margin in males strongly emarginate medially, with 12–17 teeth (n =6), with 2 triangular, dorsally projecting, and transversely oriented medial teeth, without other teeth between them. Apical labral margin in females broadly convex, not emarginate medially; with 15–21 teeth (n =9), all teeth subequal in length. Adoral labral surface in males with subapical transverse ridge ( Fig. 147 View Figs , arrow). Mandible sexually dimorphic; males with single, dorsally directed tooth, without preapical spur (cf. Fig. 189 View Figs ); females with single, mesially projecting tooth, without spur. Mentum with distinct basomedian groove ( Fig. 68 View Figs ). Prothorax: ( Figs. 74 View Figs , 83 View Figs ). Pronotum with distinctly reticulate microsculpture. Medial pronotal sulci anteriorly separate from and terminating posterior to anterior punctures ( Fig. 74 View Figs ) or continuous with anterior punctures (cf. Fig. 76 View Figs ). Distance between medial sulci slightly greater posteriorly. Pronotal basolateral carina present, but only weakly developed ( Fig. 74 View Figs , bp). Pronotal macrosetal punctures distinct (cf. Fig. 73 View Figs ). Medial pronotal seta subequidistant from medial and lateral sulci ( Fig. 74 View Figs ). Pronotal hypomeron ( Fig. 24 View Figs , hy) shiny, without microsculpture, or with distinct reticulate microsculpture. Prosternum with faintly reticulate microsculpture. Pterothorax ( Fig. 83 View Figs ): Elytron ( Fig. 23 View Figs , e) with distinct microsculpture; with 2 macrosetae, not set in punctures; laterally with single ridge ( Fig. 83 View Figs ). Mesothoracic epimeral region ( Fig. 24 View Figs , mer) with at most faint microsculpture. Metathoracic pleural region ( Fig. 24 View Figs , m) with faint reticulate microsculpture. Metathoracic pleural ridge present, poorly developed and often partially effaced; metathoracic pleural groove ( Fig. 24 View Figs , gr) incomplete posteriorly, forming elongate oval punctiform impression. Abdomen: Abdominal vestiture short, somewhat appressed, dorsally more or less evenly projecting posteriorly, but with middle setae directed posteromedially. Abdominal sternite III of male glabrous apicomedially; IV with surface very slightly impressed apicomedially, flanked by small diffuse patches of coarse acuminate setae; V with surface impressed apicomedially, flanked by coarse acuminate setae forming small tufts apically, apex of sternite slightly sinuous; VI with surface glabrous apicomedially; VII with surface glabrous apicomedially, apex slightly sinuous medially. Aedeagus ( Fig. 157 View Figs ): “ Type B” (see description on p. 8). Median lobe with apical part delimited ventrally from basal part ( Fig. 157 View Figs , arrow). Apical part of median lobe lanceolate, produced into long process. Apicolateral setae small ( Fig. 160 View Figs , arrow); apicomedial setae up to 10X longer than apicolateral setae (and forming distinct fan-like arrangement; Fig. 160 View Figs ). Paramere not exceeding apex of median lobe; slender in both dorsal and lateral view; in dorsal view with outer side outwardly curving from about middle; with 4 setae at apex.

Etymology. The name lanceolatus is an adjective from Latin (a diminutive of lanceatus), lancelike, referring to the unique shape of the median lobe of the aedeagus.

Distribution. ( Map 5 View Map 5 ). North Island: WN.

Biology and Ecology. Apparently a subalpine to alpine species. Habitat: presumably alpine scrub vegetation and tussock grasslands. Specimens have been taken from moss and litter mostly from tussock and other plants. Phenology: December– February. Elevation: 700–1,490 m.