Agnosthaetus ecarinatus Clarke, 2011

Clarke, Dave J., 2011, A Revision of the New Zealand Endemic Rove Beetle Genus Agnosthaetus Bernhauer (Coleoptera: Staphylinidae), The Coleopterists Bulletin (mo 10) 2011, pp. 1-118 : 48-49

publication ID	https://doi.org/ 10.1649/0010-065X-65.mo4.1
publication LSID	lsid:zoobank.org:pub:0818A3A2-AB42-43D8-8F76-4F65F367C584
persistent identifier	https://treatment.plazi.org/id/3CDE351F-6066-41CA-9B85-B30850A9EA5F
taxon LSID	lsid:zoobank.org:act:3CDE351F-6066-41CA-9B85-B30850A9EA5F
treatment provided by	Carolina
scientific name	Agnosthaetus ecarinatus Clarke
status	sp. nov.

Treatment

(18) Agnosthaetus ecarinatus Clarke View in CoL , new species

( Figs. 140–142 View Figs , Map 4 View Map 4 )

Type Material. Holotype. ♂, “ New Zealand : South Island: CO: Mt. Benger, 1140 m, 45°58′S, 169°26′E, tussock grassland, 7.i.2000, plot 9, turf 11, tussock litter (B.I.P. Barratt), FMNH-INS 0000 048 483 / HOLOTYPE Agnosthaetus ecarinatus Clarke , ♂, design. D. Clarke 2011”, in AGRL. The aedeagus is everted from the abdomen GoogleMaps . Paratypes. 47 specimens (27♂ 20♀; all except two ♀ s in AGRL). NEW ZEALAND: South Island: CO: Mt. Benger [see Barratt et al., 2009 for locality and collection details], 1140 m, 45°58′S, 169°26′E, tussock grassland, 7.i.2000, plot 9, turf 7, turf, B.I.P. Barratt GoogleMaps , 4♂, FMNH-INS 48478–480, 486; 7.i.2000, plot 9, turf 11, turf, B.I.P. Barratt, 1♂, 1♀ [in glyc., no #], 3♂, 1♀, FMNH-INS 48481–482, 484–485; Mt. Benger , 1100 m, 8.i.2003, plot 2, turf 2, turf, B.I.P. Barratt , 1♂, FMNH-INS 66698; 8.i.2003, plot 7, turf 9, turf, B.I.P. Barratt, 1♀, FMNH-INS 66702; 8.i.2003, plot 2, turf 8, turf, B.I.P. Barratt, 1♂, FMNH-INS 66703; 7.i.2000, plot 9, turf 14, turf, B.I.P. Barratt, 1♂, FMNH-INS 66707; 8.i.2003, plot 2, turf 18, turf, B.I.P. Barratt, 1♀, FMNH-INS 66715; 7.i.2000, plot 9, turf 5, turf, B.I.P. Barratt, 1♂, 1♀, FMNH-INS 66716–717; 8.i.2003, plot 2, turf 11, turf, B.I.P. Barratt, 1♀, FMNH-INS 66718; 7.i.2000, plot 9, tuss. 9, tussock litter, B.I.P. Barratt, 3♂, 1♀, FMNH-INS 66683–685, 686; 7.i.2000, plot 9, tuss. 7, tussock litter, B.I.P. Barratt, 1♂, FMNH-INS 66688; 8.i.2003, plot 7, tuss. 2, tussock litter, B.I.P. Barratt, 1♀, FMNH- INS 66689; 8.i.2003, plot 2, tuss. 9, tussock litter, B.I.P. Barratt , 1♀, FMNH-INS 66690; 8.i.2003, plot 2, tuss. 1, tussock litter, B.I.P. Barratt, 2♀, FMNH- INS 66691–692; 8.i.2003, plot 7, tuss. 1, tussock litter, B.I.P. Barratt , 3♂, 1♀, FMNH-INS 66693– 695, 696; 8.i.2003, plot 2, tuss. 3, tussock litter, B.I.P. Barratt, 1♂, FMNH-INS 66697; 8.i.2003, plot 2, tuss. 6, tussock litter, B.I.P. Barratt, 3♀, FMNH-INS 66699–701; 16.i.2004, plot 9, tuss. 1, tussock litter, B.I.P. Barratt, 3♂, FMNH-INS 66704–706; 7.i.2000, plot 9, tuss. 4, tussock litter, B.I.P. Barratt, 1♂, FMNH-INS 66708; 7.i.2000, plot 9, tuss. 1[a], tussock litter, B.I.P. Barratt, 2♂, 2♀, FMNH-INS 66709–710, 711–712; 8.i.2003, plot 2, tuss. 4, tussock litter, B.I.P. Barratt, 1♂, FMNH-INS 66713; 8.i.2003, plot 2, tuss. 7, tussock litter, B.I.P. Barratt, 1♀, FMNH-INS 66714; Old Man Ra., Shingle Rd. , 15.iii.1975, 75/27, litter, J.C. Watt , 2♀, FMNH-INS 48258–259 (in NZAC) .

Diagnosis. As for the species-group. In addition to the combination of those characters, A. ecarinatus may be positively separated from all others by the

single elytral ridge ( Fig. 24 View Figs ek) and metathoracic pleural ridge. The parameres of the aedeagus are distinctly slender in both dorsal and lateral view, and have a distinct point of inflection at the middle ( Fig. 141 View Figs , arrow).

Description. Color: More or less uniformly reddish brown. Head: Frontal ridge absent. Dorsum sparsely punctate; with punctures distributed anteriorly, laterally, and posteriorly on disc, middle part impunctate. Punctures shallow, well-defined; diameter subequal to or slightly greater than diameter of eye facet; interpuncture distance approximately 0.5–1.0X puncture diameter. Dorsal microsculpture present on entire or most of surface; distinctly reticulate (cf. Fig. 11 View Figs ). Dorsal tentorial sulcus (cf. Figs. 10–11 View Figs , dt) ovate; small, width slightly greater than puncture diameter. Sublongitudinal ridge (cf. Fig. 10 View Figs , sr) indistinct; confused by dorsolateral secondary carinae or punctures; crest at antennal tubercle with distinct microsculpture. Area above and behind antenno-ocular carina ( Figs. 10–11 View Figs , arrow) with distinct subsidiary carina formed by confluent punctures. Antenno-ocular carina (cf. Fig. 10 View Figs , ao) more or less absent (cf. Figs. 10–11 View Figs ). Temple ( Fig. 11 View Figs , tm) short, much less than 50% EYL. Subocular surface more or less evenly microsculptured (cf. Fig. 65 View Figs ). Labrum not distinctly sexually dimorphic ( Fig. 140 View Figs ). Apical labral margin in males moderately broadly and shallowly emarginate medially, evenly dentate, with 14–17 teeth (n =7), all teeth normal, projecting more or less anteriorly. Apical labral margin in females broadly convex, not emarginate medially; with 14 teeth (n =1), all teeth subequal in length. Adoral labral surface in males smooth, without subapical transverse ridge. Mandible more or less identical in both sexes; males and females with single, mesially directed tooth, males and females without preapical spur (cf. Fig. 189 View Figs ). Prothorax: Pronotum with distinctly reticulate microsculpture. Medial pronotal sulci anteriorly subcontinuous with anterior punctures. Distance between medial sulci very slightly greater posteriorly. Pronotal basolateral carina absent (cf. Figs. 23 View Figs , 69 View Figs , 77 View Figs ), or present, distinct (cf. Figs. 73 View Figs , 75 View Figs ). Anterior pronotal puncture (cf. Fig. 70 View Figs , ap) indistinct; medial puncture (cf. Fig. 70 View Figs , mu) absent or indistinct; basal puncture ( Fig. 70 View Figs , bu) absent. Medial pronotal seta adjacent to lateral sulcus (cf. Fig. 70 View Figs ). Pronotal hypomeron ( Fig. 24 View Figs , hy) with distinct reticulate microsculpture. Prosternum with distinctly reticulate microsculpture. Pterothorax: Elytron ( Fig. 23 View Figs , e) with distinct microsculpture; with 3 macrosetae, not set in punctures; laterally with single ridge (cf. Fig. 24 View Figs , ek). Mesothoracic epimeral region ( Fig. 24 View Figs , mer) with distinct microsculpture. Metathoracic pleural region ( Fig. 24 View Figs , m) with distinct reticulate microsculpture. Metathoracic pleural ridge present (cf. Fig. 24 View Figs , mp), fully developed; metathoracic pleural groove ( Fig. 24 View Figs , gr) incomplete posteriorly, forming elongate oval punctiform impression. Abdomen: Abdominal vestiture long, dorsally more or less evenly projecting posteriorly. Aedeagus ( Fig. 141 View Figs ): “ Type A” (see description on p. 8). Apical part of median lobe slightly narrower at base, forming weak apicolateral lobes; gradually converging to rounded point. Apicolateral setae up to 4X longer than apicomedial setae; apicomedial setae small ( Fig. 142 View Figs ). Paramere not exceeding apex of median lobe; slen- der in both dorsal and lateral view; in dorsal view with distinct inflection point at middle ( Fig. 141 View Figs , arrow); with 4 large setae at apex.

Etymology. The specific epithet ecarinatus is an adjective from the Latin ex- (out or not) and carinatus (-s, -um), with a carina, and refers to the lack of the antenno-ocular carina which is characteristically present in other species.

Distribution. ( Map 4 View Map 4 ). South Island: CO.

Biology and Ecology. Habitat: tussock grassland. Specimens have been taken from tussock litter. Phenology: January; March. Elevation: 1,140 m.

Remarks. The lack of a discernable antennoocular carina is unique within Agnosthaetus (all other species have a distinct carina, though this may be difficult to discern because of other sculpturing). This species may also be readily distinguished from species with distinct microsculpture on the head and thorax in the next large species grouping by having the medial pronotal seta positioned closer to the lateral than the medial pronotal sulcus (as in Figs. 69–72 View Figs ).