Agnosthaetus brouni ( Broun, 1910 ) Clarke, Dave J., 2011

Clarke, Dave J., 2011, A Revision of the New Zealand Endemic Rove Beetle Genus Agnosthaetus Bernhauer (Coleoptera: Staphylinidae), The Coleopterists Bulletin (mo 10) 2011, pp. 1-118 : 33-36

(9) Agnosthaetus brouni ( Broun, 1910) , new combination

( Figs. 12 , 63 , 78–79 , 113 , 121 , 125 , Map 3 )

Dimerus brouni Broun, 1910: 16 (not as new, attributed to Fauvel; Dimerus Broun , not Dimerus Fiori ) (Type locality not originally specified, fixed as Hunua, Drury by lectotype designation herein); Broun 1912: 401.

Lectotype (here designated). ♂, with five labels: “[Broun’ s handwriting] Hunua| Drury./ New Zealand |. Broun Coll.| Brit. Mus.| 1922-482./ [Broun’ s handwriting] Dimerus brouni . / FMNH- INS 0000 048 424 / LECTOTYPE Dimerus brouni Broun, 1910 , ♂, design. D. Clarke 2011”, in BMNH. On card . Paralectotypes (4). ♂, card-mounted: [Broun’ s handwriting] “Hunua, Maketu/ New Zealand. Broun Coll. Brit. Mus. 1922-482/ [ Broun’ s handwriting] Dimerus brouni / [label with unintelligible text]/ FMNH-INS 0000 048 422/ PARALECTOTYPE, Agnosthaetus brouni (Broun) , ♂, desig. D. Clarke 2011” . ♀, clear card-mounted: “ New Zealand. Broun Coll. Brit. Mus. 1922-482/ [Broun’ s handwriting] Pirongia , Dec. 1909 / FMNH-INS 0000 038 242/ PARALECTOTYPE, Agnosthaetus brouni (Broun) , ♀, desig. D. Clarke 2011” . ♂, card-mounted: [Broun’ s handwriting] “D. b./ N. Zealand ( Broun ) 95-216/ FMNH-INS 0000 048 425/ PARALECTOTYPE, Agnosthaetus brouni (Broun) , ♂, desig. D. Clarke 2011” . ♀, card mounted, same labels as previous and [A. Fauvel’ s handwriting] “ Dimerus brouni Fauvel / FMNH-INS 0000 048 426/ PARALECTOTYPE, Agnosthaetus brouni (Broun) , ♀, desig. D. Clarke 2011”. (all in BMNH) .

Additional Material Examined. 81 specimens (42♂, 39♀). NEW ZEALAND: [no locality], 1♂, FMNH-INS 48425, 1♀, FMNH-INS 48426 (in BMNH); Makatote [prob. TO: Makatoti], i.1911, 1♀, FMNH-INS 48427 (in BMNH); North Island: Whangamomona Saddle [=Whangamoa Saddle?], 39°9.336′S, 174°41.802′E, 14.iii.1957, B.M. May, 1♂, FMNH-INS 48248 (in NZAC); AK: Ahuroa Forest Reserve , 36°29.04′S, 174°36.18′E [coord.= Ahuroa Scenic Reserve ], 15.xi.1983, leaf litter, P.M. Hammond, 1♀, FMNH-INS 48160 (in BMNH); Atuanui, Mt. Auckland, i.2002, pitfall A5, A Warren, 1♂, AMNZ63688 (in AMNZ); Bethells, Matuku Res., 36°52.02′S, 174°28.02′E, 26.vii.1983, 83/87, grasses & sedges, P.A. Maddison, 1♀, FMNH-INS 48158 (in NZAC); 23.vi.1987, 87/8, sifted litter, R.C. Craw, 1♂, FMNH-INS 48157 (in NZAC); Duck Creek Scenic Reserve , 36°24′36.63″, 174°41′39.4″, 15.xii.1999, M. Hansen & J. Pedersen, 1♂, 1♀, FMNH-INS 66850–8511 (in ZMUC); Epsom, 36°53.5′S, 174°45.9′E, xii.1911, 1♂, FMNH-INS 48423 (in BMNH); Hunua Gorge, 24.vii.1932, 1♀, AMNZ85004 (in AMNZ); Hunua Range, 150 m, 37°5′S, 175°11′E, 4.i.1973, 73/2, litter, B.M. May, 1♂, FMNH-INS 48161 (in NZAC); Hunua Range, Cosseys Walk Trk., 104 m, 37°3.87′S, 175°5.58′E, Kunzea -Phyllocladus - Agathis forest, 5.vii.2004, FMHD#2004-300, berl., leaf litter, D.J. Clarke, 1♀ (in FMNH); Hunua Range, Maketu, 1♂, FMNH-INS 48422 (in BMNH); Hunua, Drury, 1♂, FMNH-INS 48424 (in BMNH); Little Barrier I., Turners Strm., 21.xii.1983, plant/ soil sample 1576, in small tributary bed, K.A.V. Wise, 1♂, AMNZ85003 (in AMNZ); Manurewa, Murphys Bush, 45 m, 36°58.8′S, 174°55.14′E, 30.viii.1977, litter & wood, G. Kuschel, 1♂, FMNH- INS 48162 (in NZAC); Papakura, 37°4.2′S, 174°57′E, 21.i.1942, A. Richardson, 1♀, FMNH- INS 48167 (in NZAC); Riverhead S.F. Res., 36°43′11″S, 174°34′38″E [coord.= middle of reserve ], 11.xi.1980, P.A. Maddison, 1♂, 2♀, FMNH-INS 48163, 164–165 (in NZAC); Waitakere Ra., Cascade-Kauri Pk., Auckland City Walk Trk. [variant name(?)=Waitakere City Walk], 61 m, 36°53.17′S, 174°31.18′E, podocarp-broadleaf forest, 2.vii.2004, FMHD#2004-297, berl., leaf litter, D.J. Clarke, 1♂, 1♀ (in FMNH); Waitakere Ra., Cascade-Kauri Pk., Upper Kauri Trk., 170 m, ANMT 680, 36°53′S, 174°31′E, kauri-podocarphardwood, 23.xi.1984, FMHD#85-811, litter, forest, Berl., A. Newton & M. Thayer, 1♂, FMNH- INS 48156 (in FMNH); Waitakere Ra., Cascades Kauri Park, 36°53.394′S, 174°31.338′E [coord.= The Cascades], 6.xi.1983, leaf litter, P.M. Hammond, 1♀, FMNH-INS 48159 (in BMNH); CL: Great Barrier I., Kaiarara Valley, 36°11.418′S, 175°23.13′E, 12.iv.1982, 82/52, litter, J.C. Watt, 1♂, FMNH-INS 48108 (in NZAC); Great Barrier I., Little Windy Hill, 160 m, forest, 18.iii.2003, L11961, hand collecting, S.E. Thorpe, 1♂, AMNZ85008 (in AMNZ); Kirikiri Saddle [note: unofficial name; saddle is highest point along SH25 and coord.=trig A6C0], 424 m, 37°8.88′S, 175°40.26′E, 27.x.1981, 81/115, litter, C.F.Butcher, 1♂, FMNH-INS 48104 (in NZAC); Mt. Moehau, Wards Bush, 365 m, 1.xi.1979, 79/136, litter, B.M. May, 1♀, FMNH-INS 48106 (in NZAC); Rakitu I., 36°7.86′S, 175°29.28′E, 7.i.1981, 81/6, litter, L. Roberts, 1♂, FMNH-INS 48107 (in NZAC); Whangapoua, 347 m [elevation does not match anywhere in vicinity of Whangapoua in CL, also locality on Great Barrier I.], 31.viii.1975, 75/161, litter, A.K. Walker, 1♂, FMNH-INS 48105 (in NZAC); ND: Aranga, Trounson Kauri Park, forest, 1.xi.2000, litter in forest remnant, S.E. Thorpe, 1♂, AMNZ35068 (in AMNZ); Bay of Islands, Waikare, 35°21′S, 174°14′E, 24.ix.1960, R.A. Cumber, 1♂, FMNH- INS 48172 (in NZAC); Herekino Forest West, 35°12.594′S, 173°11.447′E, 12.xii.2008, RL 1423, sifting rotten wood and leaf litter, R. Leschen, T. Buckley & D. Seldon, 1♀ (in NZAC); Kamo, 35°40.998′S, 174°17.76′E, 19.i.1966, litter, A.K. Walker, 1♂, FMNH-INS 48155 (in NZAC); Mangamuka Summit, 386 m, 35°12′S, 173°27′E, 13.xii.1976, 76/109, litter, V.A. May, 1♀, FMNH- INS 48154 (in NZAC); Maunganui Bluff Scen. Res. 410–460 m, ANMT 697, 35°46′S, 173°34′E, coastal mixed hardwood forest, 3.xii.1984, FMHD#85-827, litter, forest, Berl., A. Newton & M. Thayer, 1♂, FMNH-INS 48148, 1♀, FMNH- INS 48149 (in FMNH); Mt. Manaia, 150–300 m, 35°49′3″S, 174°31′1″E [coord.=Manaia Hill at Whangarei Heads], 4.xi.1981, 81/123, litter & decayed wood, G. Kuschel, 1♂, FMNH-INS 48152 (in NZAC); Ngaiotonga S.R., Russel Walkway, 35°19′S, 174°15′E, 12.i.0200, RL 507, sifted leaf litter, R. Leschen, 1♀, FMNH-INS 48147 (in NZAC); Omahuta Forest Sanctuary, 230 m, 35°14′S, 173°38′E, kauri forest, 12.xii.1983, litter, P.M. Hammond, 1♂, FMNH-INS 48171 (in BMNH); Omahuta Forest, Omahuta Kauri Sanctuary, 340 m, ANMT 693, 35°14′S, 173°38′E, kauri-podocarphardwood, 29.xi.1984, FMHD#85-824, litter, forest, Berl., A. Newton & M. Thayer, 2♂, FMNH-INS 48169–170 (in FMNH); Omahuta Forest, Omahuta Kauri Sanctuary, 35°14′S, 173°38′E, 10.x.1974, 74/76, litter, J.C. Watt, 1♂, FMNH-INS 48495 (in NZAC); Paihia Opua S.F., 35°16.998′S, 174°4.998′E, 22.i.1981, litter and rotten wood, G. Kuschel, 1♂, FMNH-INS 48153 (in NZAC); Parahaki Park [=William Fraser Memorial Park], 5.xi.1981, 81/127, litter & decayed wood, G. Kuschel, 1♀, FMNH-INS 48166 (in NZAC); Puhipuhi S.F., nr. Quarry, 36°27′S, 174°15′E, 14.vi.2000, RL 591, leaf litter, R. Leschen & G. Hall, 1♂, FMNH-INS 48143, 3♀, FMNH-INS 48144–146 (in NZAC); Puketi Forest HQ, 35°13′S, 173°47′E, 31.iii.1998, RL 378, sifted leaf litter, R. Leschen, R. Hoare & G. Hall, 1♀, KUNHM-ENT SM0650738 (in KSEM); Tangihua Ra., 579 m (as 1800–2000 ft), 27.x.1936, E. Fairburn, 1♀, FMNH-INS 48168 (in NZAC); Waimatenui, 35°37′S, 173°42.6′E, 10.iii.1962, litter, R.A. Cumber, 1♂, FMNH-INS 48150, 1♀, FMNH-INS 48151 (in NZAC); Waipoua S.F., Lookout area, 29.x.1980, 80/93, sifted moss, G. Kuschel, 1♀, FMNH-INS 48141 (in NZAC); Waipoua S.F., Waikohatu Stream bridge (0.8 km S), 270 m, ANMT 692, 35°38′S, 173°34′E, hardwoodpodocarp forest, 28.xi.1984, FMHD#85-823, litter, forest, Berl., A. Newton & M. Thayer, 1♀, FMNH- INS 48139 (in FMNH); Waipoua S.F., Waipoua River bridge, 31.x.1985, 85/50, sifted litter & moss, R.C. Craw, 1♀, FMNH-INS 48142 (in NZAC); Waipoua S.F., Waipoua Strm., 120 m, 10–11.xii.1983, leaf litter, P.M. Hammond, 1♂, FMNH-INS 48140 (in BMNH); Waipoua S.F., Wairau Summit, 400 m, ANMT 687, 35°36′S, 173°32′E, hardwood-podocarp forest, 1.xii.1984, FMHD#85-818, litter, forest, Berl., A. Newton & M. Thayer, 2♂, FMNH-INS 48135– 136, 1♀, FMNH-INS 48137 (in FMNH); Waipoua S.F., Wairau Summit (0.8 km NW), 350 m, ANMT 689, 35°35′S, 173°31′E, hardwood-podocarp forest, 27.xi.1984, FMHD#85-820, litter, forest, Wink., A. Newton & M. Thayer, 2♂, 2♀ (in); 1♂, FMNH- INS 48138 (in FMNH); TK: Egmont N.P., Kapuni Loop Trk., 828 m, 39°19.607′, 174°6.495′, broadleaf-kamahi forest, 16.i.2008, KM176, sifted leaf litter, K. Marske & R. Leschen, 1♂ (in NZAC); Egmont (S), 5.x.1985, F. Chambers, 1♀, FMNH-INS 19801 (in MONZ); Egmont N.P., East Egmont, Plateau, 1172 m, 39°18.66′S, 174°5.82′E [coord.=The Plateau], 24.i.1987, 1♂, FMNH-INS 48133 (in Puthz); Mt. Egmont, Potaema Walk, 540 m, forest, 9.xii.1995, in forest litter, J.T. Nunn, 1♀, FMNH-INS 19684 (in JTNC); Mt. Messenger, 16 m, 38°54.207′S, 174°35.643′E, podocarp/broadleaf forest, 15.i.2008, KM172, sifted leaf litter & dead wood, K. Marske, R. Leschen & T. Buckley, 1♂, 1♀ (in NZAC); Mt. Messenger, 200 m, 20.iii.2005, L14302, litter/ground plants, S.E. Thorpe, 1♂, AMNZ85007 (in AMNZ); Waitewhenua V., Ohura (19 km N), 250 m, Nothofagus truncata forest, 5.xii.1983, litter, P.M. Hammond, 1♀, FMNH- INS 48134 (in BMNH); Whangamomona Saddle, Awahou Ridge Trk., base of ascent to saddle, 188 m, 39°9.266′S, 174°42.807′E, 17.i.2008, KM199, hand collected, ex dead wood/fungus, K. Marske, R. Leschen & T. Buckley, 1♂ (in NZAC); 17.i.2008, KM198, sifted leaf & wood litter, K. Marske, R. Leschen & T. Buckley, 2♂ (in NZAC); TO: Erua, 39°14′S, 175°24′E, 30.xii.1940, 1♂, AMNZ85005 (in AMNZ); WO: Hakarimata, ca. 100–300 m, 37°39.83′S, 175°7.44′E, 20.xi.1983, leaf litter, P.M. Hammond, 1♂, FMNH-INS 48115, 1♀, FMNH-INS 48116 (in BMNH); Hapuakohe Ra., Maungakawa, 500 m, 37°24.96′S, 175°23.76′E, 3.i.1984, nikau leaf bases on ground, P.M. Hammond, 1♂, FMNH-INS 48117, 2♀, FMNH-INS 48118–119 (in BMNH); Mahoenui, Gribbons Rd., 38°32.58′S, 174°49.02′E [coord.= approx. midpoint of rd.], 26.vi.1977, 77/81, litter, G.W. Ramsay, 2♂, FMNH-INS 48110, 112, 1♀, FMNH-INS 48114 (in NZAC); 26.vi.1977, 77/78, sifted litter, N.H. Mancer, 1♂, FMNH-INS 48109 (in NZAC); 26.vi.1977, 77/79, sifted litter (J.C Muricer), 2♀, FMNH-INS 48111, 113 (in NZAC); Pirongia, 38°0′S, 175°11′E, xii.1909, 1♀, FMNH-INS 38242 (in BMNH).

Diagnosis. Agnosthaetus brouni may be reliably distinguished from all other species of Agnosthaetus , except A. tumidus , A. bicolor , and A. zonatus , by the combination of dorsum of head distinctly punctate ( Fig. 12 ) and with distinct subocular band of microsculpture ( Fig. 63 , bm), lack of microsculpture on dorsal sides of head and thorax, smooth mentum (as in Fig. 67 ), complete lateral pronotal sulcus ( Fig. 78 , ls) and carina ( Fig. 79 , pc), metathoracic pleural ridge absent (e.g., Figs. 85–86 ), apex of male labrum with broad, well-defined emargination (without larger medial tooth), and apex of female labrum with distinctly larger medial tooth ( Fig. 113 ). From A. tumidus , A. bicolor , and A. zonatus , the aedeagus of A. brouni may be distinguished by the combination of apical part of median lobe without subparallel ridges near base ( Fig. 125 , arrow, cf. Fig. 134 , arrow), paramere in dorsal view with apical part concave mesially ( Fig. 121 , arrow), and the basal sclerite of internal sac broadly U-shaped and with arms gradually diverging ( Fig. 195 , arrow).

Description. Color: Variable, usually reddish brown, and with segment VI (and frequently the elytra) darker to nearly black. Head: Frontal ridge present (cf. Fig. 12 , fr). Dorsum moderately densely punctate; with punctures distributed evenly over surface, but occasionally absent from middle part. Punctures deep, but indistinctly defined; diameter up to 2X diameter of eye facet; interpuncture dis- tance mostly much less than half puncture diameter ( Fig. 12 ). Dorsal microsculpture absent. Dorsal tentorial sulcus (cf. Figs. 10–11 , dt) distinctly slit-like; width subequal to or less than puncture diameter. Sublongitudinal ridge (cf. Fig. 10 , sr) distinct; not confused by smaller carinae or punctures; crest at antennal tubercle without distinct microsculpture. Area above and behind antennoocular carina ( Figs. 10–11 , arrow) more or less smooth, without subsidiary carinae, or with single subsidiary carina formed by confluent punctures. Antenno-ocular carina joining eye at or behind middle ( Fig. 12 , ao). Temple ( Fig. 11 , tm) short, less than 50% EYL. Subocular surface with narrow well-demarcated band of microsculpture ( Fig. 63 , bm). Labrum distinctly sexually dimorphic ( Fig. 113 ). Apical labral margin in males strongly emarginate medially, evenly dentate, with 18–26 teeth (n =7), all teeth normal, projecting more or less anteriorly. Apical labral margin in females broadly convex, not emarginate medially; with 21–25 teeth (n =13), with medial tooth elongate, twice length of others. Adoral labral surface in males smooth, without subapical transverse ridge. Mandible more or less identical in both sexes; males and females with single, mesially directed tooth, males with distinct preapical spur (cf. Fig. 190 , arrow) or with weakly developed preapical spur (cf. Fig. 190 , arrow); females without spur. Prothorax ( Fig. 78–79 ): Pronotum without microsculpture. Medial pronotal sulci anteriorly separate from and terminating posterior to anterior punctures ( Fig. 78 ). Distance between medial sulci slightly greater posteriorly. Pronotal basolateral carina absent ( Fig. 78 ). Anterior pronotal puncture (cf. Fig. 70 , ap) absent or indistinct; medial puncture (cf. Fig. 70 , mu) distinct; basal puncture ( Fig. 70 , bu) absent. Medial pronotal seta subequidistant from medial and lateral sulci ( Fig. 78 ). Pronotal hypomeron ( Fig. 24 , hy) shiny, without microsculpture. Prosternum without microsculpture. Pterothorax: Elytron ( Fig. 23 , e) without microsculpture; without punctiform, non-setiferous impressions,or with small, punctiform, non-setiferous impressions; with 2 macrosetae, not set in punctures; laterally with single ridge (cf. Fig. 24 , ek). Mesothoracic epimeral region ( Fig. 24 , mer) shiny, without microsculpture. Metathoracic pleural region ( Fig. 24 , m) shiny, without microsculpture. Metathoracic pleural ridge absent; metathoracic pleural groove ( Fig. 24 , gr) complete, continuing to or near to pleurocoxal articulation. Abdomen: Abdominal vestiture short, somewhat appressed, dorsally with radiating fans of setae either side of midline and laterally, or more or less evenly projecting posteriorly but with middle setae directed posteromedially. Abdominal sternite V of male with large diffuse patch of coarser setae medially. Aedeagus ( Fig. 121 ): “ Type A” (see description on p. 8). Apical part of median lobe distinctly trilobate, with rounded lateral lobes, with abruptly distinct, broadly acuminate, and sharply pointed apicomedian lobe. Both apicolateral and apicomedial setae short ( Fig. 125 ). Paramere extending to about level of median lobe apex; lamellate, in lateral view distinctly broad along most of length; with apical part curving ventrally; in dorsal view with outer side gently convex; with 4 large setae near inner edge of apex; internal sac as in Fig. 195 .

Distribution. ( Map 3 ). North Island: AK; CL; ND; TK; WO.

Biology and Ecology. Habitat: Agathis , Nothofagus , and podocarp forest. Specimens have been taken mostly from forest leaf litter. Phenology: year-round. Elevation: 16– 540 m.

Remarks. The name Dimerus brouni Broun , which was overlooked by subsequent authors (e.g., Scheerpeltz 1933; Bernhauer 1939; Herman 2001), is available because Broun (1910) provided a valid description of this species while comparing it to his new species, Dimerus stilbus . The discovery of this name and its transfer here to Agnosthaetus complicates the nomenclature of the type species of Agnosthaetus ( Agnosthaetus brouni Bernhauer ), which, as the junior secondary homonym, is here given a replacement name (for further details, see Etymology under A. brounides , p. 21). Although the Broun name is available, it cannot be firmly established which of the seven BMNH specimens labeled as Dimerus brouni (or by abbreviation D. b.) were studied by Broun when he wrote his description. Designation of a lectotype is particularly important here, as there are several externally indistinguishable New Zealand species resembling the BMNH D. brouni material, and two of the candidate syntype specimens lack specific locality details. The lectotype matches the original description, and is from an area of northern New Zealand where only this species occurs. Thus, selection of this species fixes the type locality (Hunua, Drury) and the application of this otherwise problematic name. The two paralectotype specimens (FMNH-INS 0000 048 425 and 426) appear to belong to a series and have identical data; the female was likely the one Broun described as having sent to A. Fauvel, who allegedly ( Broun 1912) recognized it as belonging to a new genus, Dimerus . Broun attributed Dimerus , as he used it, to Fauvel, and this is not the same as Dimerus Fiori (now a junior generic synonym in Pselaphinae ). Two specimens in BMNH, while conspecific with the lectotype, are excluded from the type series as they have recorded collecting dates later than 1910, and thus Broun could not have seen them when describing D. brouni . Broun describes D. stilbus and D. brouni as differing in mandibular dentition, and since the first paralecto- type specimen has its mandibles closed, that specimen was excluded as a possible lectotype candidate. Nevertheless, although there still remains some doubt surrounding that specimen, the existence of a label with some descriptive text (only “coarse punctation” can be discerned) in Broun’ s handwriting is sufficient to consider it part of the type series.

Agnosthaetus brouni is one of a group of several species (including A. tumidus , A. zonatus , and A. bicolor ) that may be reliably distinguished only by comparing characters of the aedeagus (paramere and median lobe shape, internal sac sclerites), as given in the diagnoses and identification key.

Figures

Map 3. Geographic distribution of all georeferenced specimens of North Island species in the Agnosthaetus brouni species-group. Triangles = A. brouni; stars = A. tumidus; circles = A. akatarawa; squares = A. aequalis.

Figs. 10–13. Head capsules of Agnosthaetus spp. 10) A. stilbus; 11) A. rodmani; 12) A. brouni; 13) A. chiasma. Scale bars = 100 mm.

Figs. 63–68. Heads of Agnosthaetus spp., ventral view. 63) A. brouni; 64) A. stilbus, oblique view; 65) A. chiasma;

Figs. 75–78. Pronota of Agnosthaetus spp. 75) A. vicinus; 76) A. stilbus; 77) A. nitidus; 78) A. brouni. Scale bar =

Figs. 79–82. Prothoraces of Agnosthaetus spp., right lateral view. 79) A. brouni; 80) A. nitidus; 81) A. rodmani;

Figs. 113–120. Male (left) and female (right) labra of species in the Agnosthaetus brouni species-group, dorsal (anterior margin) view. 113) A. brouni; 114) A. tumidus; 115) A. aequalis; 116) A. akatarawa; 117) A. zonatus; 118) A. bicolor; 119) A. impressus; 120) A. nitidus. Scale bar = 100 mm.

Figs. 121–124. Aedeagi of species in the Agnosthaetus brouni species-group, dorsal (left) and right lateral (right) views. 121) A. brouni; 122) A. tumidus; 123) A. aequalis; 124) A. akatarawa. Scale bar = 100 mm.

Figs. 125–128. Apex of median lobe of species in the Agnosthaetus brouni species-group, ventral view.

Figs. 83–88. Pterothoraces of Agnosthaetus spp., right lateral view. 83) A. lanceolatus; 84) A. stilbus; 85) A. chiasma;

Figs. 133–136. Apex of median lobe of species in the Agnosthaetus brouni species-group, ventral view.

Figs. 195–197. Aedeagi of some species in the Agnosthaetus brouni species-group, with details of the internal sac, dorsal view. 195) A. brouni; 196) A. tumidus; 197) A. bicolor. Scale bar = 200 mm.

Figs. 189–194. Mandibles of Agnosthaetus spp., dorsal view (189–193, left; 194, right). 189) A. newtoni; 190) A. falx; 191) A. stenomastax; 192) A. heteromastax (Rakeahua Valley); 193) A. heteromastax (Big South Cape Island); 194) A. heteromastax (Twilight Bay). Scale bar = 100 mm.

Figs. 69–74. Pronota of Agnosthaetus spp. 69) A. chiasma; 70) A. rodmani; 71) A. heteromastax; 72) A. stenomastax;

Figs. 23–24. Agnosthaetus cariniceps, thorax. 23) Dorsal view; 24) Left lateral view. Scale bars = 100 mm.