Agnosthaetus affinis Clarke

Clarke, Dave J., 2011, A Revision of the New Zealand Endemic Rove Beetle Genus Agnosthaetus Bernhauer (Coleoptera: Staphylinidae), The Coleopterists Bulletin (mo 10) 2011, pp. 1-118 : 70-71

publication ID	https://doi.org/ 10.1649/0010-065X-65.mo4.1
publication LSID	lsid:zoobank.org:pub:0818A3A2-AB42-43D8-8F76-4F65F367C584
persistent identifier	https://treatment.plazi.org/id/DA354009-37D1-46A7-9792-616E8E97323F
taxon LSID	lsid:zoobank.org:act:DA354009-37D1-46A7-9792-616E8E97323F
treatment provided by	Carolina
scientific name	Agnosthaetus affinis Clarke
status	sp. nov.

Treatment

(34) Agnosthaetus affinis Clarke View in CoL , new species

( Figs. 179 View Figs , 183 View Figs , 187 View Figs , Map 7 View Map 7 )

Type Material. Holotype. ♂, with three labels: “ NEW ZEALAND: CO| Piano Flat [45°32′S, 169°1′E], Titan Rocks| track 9-XII-1998 | C. Hall ex. litter/ SM0673434 | KUNHM-ENT / HOLOTYPE Agnosthaetus affinis Clarke , ♂, design. D. Clarke 2011”, in KSEM GoogleMaps . Paratypes. 6 specimens (5♂ 1♀). Same data as holotype GoogleMaps : 5♂, KUNHM-ENT SM0673340, KUNHM-ENT SM0673398, KUNHM- ENT SM0673414, KUNHM-ENT SM0673428, KUNHM-ENT SM0673430, 1♀, KUNHM-ENT SM0673427 (in KSEM) .

Diagnosis. The distinctive and even microsculpture on the head and thorax of A. affinis distinguishes this species from others in the nunni species-group, but makes it closely resemble species in the thayerae and truncatus speciesgroups, as well as A. lanceolatus (North Island) and A. ecarinatus . From these species, A. affinis may be distinguished by the combination of distinct antenno-ocular carina ( Fig. 10 View Figs , ao; which alone distinguishes it from A. ecarinatus ), more or less distinct sublongitudinal ridge ( Fig. 10 View Figs , sr; which distinguishes it from the truncatus speciesgroup), shallow and sparse dorsal head punctuation, elytra with single lateral ridge ( Fig. 24 View Figs , ek) (these last two easily distinguish it from the thayerae species-group), and males with unmodified labral teeth ( Fig. 179 View Figs ; distinguishes males from all aforementioned species except A. ecarinatus ).

Description. Color: More or less uniformly yellowish brown, with abdominal segment VI distinctly darker, nearly black ( Fig. 5 View Figs ). Head: Frontal ridge absent. Dorsum sparsely punctate; with punctures distributed anteriorly, laterally, and posteriorly on disc, middle part impunctate. Punctures shallow, rather indistinct; diameter greater than diameter of eye facet; interpuncture distance approximately 0.5–1.0X puncture diameter. Dorsal microsculpture present on entire or most of surface; distinctly reticulate, fainter posteriorly. Dorsal tentorial sulcus (cf. Figs. 10–11 View Figs , dt) narrowly ovate; width subequal to or slightly greater than puncture diameter. Sublongitudinal ridge (cf. Fig. 10 View Figs , sr) distinct; not confused by smaller carinae or punctures (cf. Fig. 10 View Figs , sr); crest at antennal tubercle with distinct microsculpture. Area above and behind antenno-ocular carina ( Figs. 10–11 View Figs , arrow) more or less smooth, without subsidiary carinae. Antenno-ocular carina joining eye at or behind middle (cf. Fig. 10 View Figs , ao). Temple ( Fig. 11 View Figs , tm) short, less than 50% EYL. Subocular surface more or less evenly microsculptured (cf. Fig. 65 View Figs ). Labrum not distinctly sexually dimorphic ( Fig. 179 View Figs ). Apical labral margin in males very shallowly emarginate medially, evenly dentate, with 16–21 teeth (n =4), all teeth normal, projecting more or less anteriorly. Apical labral margin in females broadly convex, not emarginate medially; with 17 teeth (n =1), all teeth subequal in length. Adoral labral surface in males smooth, without subapical transverse ridge. Mandible sexually dimorphic; males with single, dorsally directed tooth, without preapical spur (cf. Fig. 189 View Figs ); females with single, mesially projecting tooth, without spur. Prothorax: Pronotum with distinctly reticulate microsculpture. Medial pronotal sulci anteriorly continuous with anterior punctures (cf. Fig. 76 View Figs ). Distance between medial sulci very slightly greater posteriorly. Pronotal basolateral carina distinct (cf. Fig. 76 View Figs , bp). Pronotal macrosetal punctures distinct (cf. Fig. 73 View Figs ). Medial pronotal seta subequidistant from medial and lateral sulci (cf. Fig. 73 View Figs , mu). Pronotal hypomeron ( Fig. 24 View Figs , hy) with distinct reticulate microsculpture. Prosternum with distinctly reticulate microsculpture. Pterothorax: Elytron ( Fig. 23 View Figs , e) with distinct microsculpture; with 2 macrosetae or with 3 macrosetae, set in distinct punctures; laterally with single ridge (cf. Fig. 84 View Figs , ek). Mesothoracic epimeral region ( Fig. 24 View Figs , mer) with distinct microsculpture. Metathoracic pleural region ( Fig. 24 View Figs , m) with distinct reticulate microsculpture. Metathoracic pleural ridge fully developed (cf. Fig. 24 View Figs , mp); metathoracic pleural groove ( Fig. 24 View Figs , gr) incomplete posteriorly, forming elongate oval punctiform impression. Abdomen: Abdominal vestiture short, somewhat appressed, dorsally more or less evenly projecting posteriorly but with middle setae directed posteromedially. Abdominal sternite V of male with surface impressed apico- medially, flanked by coarse acuminate setae forming small tufts apically, apex of sternite not distinctly sinuous; VI with subapicomedial patch of denser setae. Aedeagus ( Fig. 183 View Figs ): “ Type B” (see description on p. 8). Apical part of median lobe slightly narrower basally, not forming distinct lateral lobes; produced concavely into narrow point. Apicolateral setae small; apicomedial setae up to 10X longer than apicolateral setae ( Fig. 187 View Figs ). Paramere not exceeding apex of median lobe; in lateral view produced apically into lobe; with apical part perpendicular to median lobe; in dorsal view with outer side more or less straight; with 2 small setae at apex and 2 large ones on ventral edge.

Etymology. The specific epithet affinis is an adjective from the Latin affinis (-e), near or related to, for its general similarity to other species in both the nunni species-group, and those other species with distinctive microsculpture.

Distribution. ( Map 7 View Map 7 ). South Island: CO.