Ditaeniella grisescens Meigen, 1830

Vikhrev, Nikita E. & Murphy, William L., 2022, Notes on the taxonomy of species of Sciomyzini with a predominantly setulose anepisternum (Diptera, Sciomyzidae), Amurian Zoological Journal XIV (2), pp. 281-298 : 287-289

Ditaeniella grisescens Meigen, 1830

Figs. 7 , 14, 15

Sciomyza grisescens Meigen, 1830

Melina (Ditaenia) grisescens Meigen, 1830 ( Cresson, 1920)

Pherbellia grisescens Meigen, 1830 ( Malloch, 1933)

Ditaeniella grisescens Meigen, 1830 ( Rozkošný, 1987)

Sciomyza patagonensis Macquart, 1851 = Pherbellia grisescens Meigen, 1830 ( Malloch, 1933)

Specimens examined:

BELARUS, Minsk Reg., Borisov , Berezina R., 54.239 ° N, 28.494 ° E, D. Gavryushin, 5.07.2013, 1♂ ( ZMUM) .

CHINA, Liaoning Prov., 50 km north of Mukden (= Shenyang = Fengtian) (42.3 ° N, 123.4 ° E), I. Rubtsov, 20.07.1952, 1♀ ( ZIN) .

HUNGARY, Domsod , Apaj-puszta, (47.1 ° N, 19.1 ° E), K. Gorodkov, 25.06.1970, 2♀ ( ZIN) .

IRAN: Isfahan Prov., Ghomrood R., 1920 m, 33.42 ° N 50.12 ° E, O. Kosterin, 21.05.2017, 2♀ ; Markazi Prov., Arak env., 1700 m, 34.13 ° N, 49.71 ° E, O. Kosterin, 18– 30.05.2017, 1♂ ; Gav Godar env., 1800 m, 34.11 ° N, 49.36 ° E, O. Kosterin, 19.05.2017, 1♂ ; Hendoudar R., 2000 m, 33.76 ° N, 49.22 ° E, O. Kosterin, 20.05.2017, 1♂, 1♀; Shazand env., 2000 m, (33.92 ° N, 49.40 ° E), I. Grichanov, 20.05.2017, 1♂ (all ZMUM).

KAZAKHSTAN: Almaty Reg., Kapchagay Lake env., 43.7 ° N, 77.2 ° E, N. Vikhrev, 22 – 28.05.2016, 2♂; Kyzylorda Reg., Bazykara env., 45.757 ° N, 62.311 ° E, fresh pond, K. Tomkovich, 15 – 19.05.2011, 6♂, 1♀; Perovsk (= Kyzylorda ) (≈ 44.75 ° N, 65.71 ° E), A. Zhelokhovtsev , 14.05.1925, 1♂ (all ZMUM); Turkistan Reg., Saryagash Distr. (≈ 41.6 ° N, 68.7 ° E), P. Ler , 27.04.1957, 1♂ ( ZIN) .

KYRGYZSTAN: Bishkek Reg ., Bishkek Park, 42.90 ° N 74.62 ° E, N. Vikhrev, 17.09.2013, 2♂, 1♀ ( ZMUM); Issyk-Kul Lake, Rybachiy (= Balykchy) (42.46 ° N, 76.2 ° E), 1600 m, E. Nartchuk, K. Gorodkov, 16.08.1969, 4♂, 2♀ ( ZIN) .

MONGOLIA: Dornod aimak, near Choibalsan , 40 km upstream Kerulen R. (48.0 ° N, 113.9 ° E) I. Kerzhner, 27.07.1971, 5♂, 1♀ ; Omnogovi aimak, 20 km WSW Bayan-Ovoo (42.9 ° N, 105.8 ° E), I. Kerzhner, 8 August 1971, 1♂, 1♀ ; Tov aimak, Urga (= Ulaanbaatar), Tola R. (47.88 ° N, 106.90 ° E), P. Kozlov, 29– 30.06.1905, 1♂ ; 5– 7.07.1905, 2♀; Uvs aimak, 50 km ESE of Ulangom , 49.95 ° N, 92.65 ° E, E. Narchuk, 7.08.1975, 1♀ (all ZIN).

RUSSIA: Altai Krai Reg., Zmeinogorsk Distr. , Kolyvanskoe L. (≈ 51.37 ° N, 82.17 ° E), O. Kosterin, 8.09.2007, 1♂, 2♀ ( ZMUM) ;

Amur Reg., Zeya env. (53.7 ° N, 127.2 ° E), A. Shatalkin, 6.06.1978, 1♀ ( ZMUM) ;

Astrakhan Reg., Baskunchak Salt Lake , 48.167 ° N, 46.830 ° E, near lake, K. Tomkovich, 3– 6.05.2010, 1♂ ; 48.193 ° N, 48.813 ° E, fresh pond silt, K. Tomkovich, 1– 2.05.2010, 1♀ (all ZMUM) ;

Buryatia Reg.: Ust-Kiran env., R. Chikoy (50.42 ° N, 106.83 ° E), Khomze, 29.07.1908, 1♀ ( ZIN); Mondy env., 51.62 ° N, 100.91 ° E, 1960 m, 22– 24.06.2021, N. Vikhrev, 1♂ ( ZMUM) .

Crimea Reg., Koktebel env. (44.95 ° N, 35.25 ° E), Dyakonov , 27.08.1923, 1♀ ( ZIN) ;

Irkutsk Reg.: Bratsk (≈ 56.29 ° N, 101.72 ° E), A. Monchadsky, 29.06.1956, 1♀; (Olkhon L.) Peschanka (≈ 53.285 ° N, 107.58 ° E), P. Mikhno, 25.07.1925, 1♀ (all ZIN) ;

Kaliningrad Reg., Khrabrovo env., 54.88 ° N, 20.60 ° E, K. Tomkovich, 23.08.2013, 1♀ ; 54.90 ° N, 20.65 ° E, fresh ponds, K. Tomkovich, 13– 15.08.2013, 1♂, 1♀ ( ZMUM) ;

Karachay-Cherkess Reg., Bol. Zelenchuk R., 44.22 ° N, 41.72 ° E, 580 m, O. Kosterin, 6.07.2019, 1♂, 1♀ ( ZMUM) ;

Khanty-Mansi Reg., Khanty-Mansiysk Airport , 61.04 ° N, 69.11 ° E, fir forest, K. Gorodkov, 28.08.1976, 4♂, 4♀ ( ZIN) .

Krasnodar Reg.: Adler env., Imereti Lowlands (43.41 ° N, 39.95 ° E), N. Vikhrev, 2.10.2008, 2♀ ; Gelendzhik Distr., Krinitsa env., Pshada R., 44.396 ° N, 38.342 ° E, K. Tomkovich, 6– 13.09.2009, 1♂ (all ZMUM) ;

Krasnoyarsk Reg., Ergaki National Park , 1450 m, 52.84 ° N, 93.25 ° E, N. Vikhrev, 27– 29.06.2017, 1♂ ( ZMUM) ;

North Ossetia – Alania Reg., Mozdok Distr., Sukhotskoe (≈ 43.67 ° N, 44.44 ° E), A. Ozerov, 3.08.1988, 1♀ ( ZMUM) ;

Omsk Reg.: Irtysh R. floodplain, oxbow of Zamarayka R. (54.964 ° N, 73.353 ° E), O. Kosterin: 14– 18.07.2007, 1♀; Krugloe L., 54.89 ° N, 73.36 ° E, O. Kosterin, 24.07.2012, 1♂, 1♀; Ptichya Gavan oxbow, 54.97 ° N, 73.35 ° E, O. Kosterin, 23.07.2008, 1♀ (all ZMUM) ;

Primorsky Reg.: Vladivostok, Sedanka (43.2 ° N, 132.0 ° E), A. Stackelberg, 20.06.1927, 1♀ ( ZIN) ; Khanka Lake , 45.06 ° N, 131.99 ° E, N. Vikhrev, 15– 19.07.2014, 2♂, 1♀ ( ZMUM) ;

Rostov Reg., Tsimlyansk env. (47.65 ° N, 42.10 ° E), O. Negrobov, 27.09.1965, 1♂ ( ZIN) ;

Saint Petersburg Reg.: Yukki (60.11 ° N, 30.28 ° E), A. Stackelberg, 27.08.1953, 6♂, 9♀ ; Luga Distr., Yaschera (59.15 ° N, 29.91 ° E), A. Stackelberg, 17.07.1963, 1♀ (all ZIN) ;

Tuva Reg., Dus-Khol Salt Lake , 700m, 51.36 ° N, 94.45 ° E, N. Vikhrev, 2– 5.07.2017, 1♂ ( ZMUM) ;

Tver Reg., Volgo L., 56.877 ° N, 33.244 ° E, A. Ozerov, 18– 19.08.2011, 1♀ ( ZMUM) ;

Ulyanovsk Reg., Radischevo Distr., Vjazovka env., 52.82 ° N, 48.35 ° E, K. Tomkovich, 2– 5.05.2011, 1♂ ( ZMUM) ;

Yamalo-Nenetsk Reg.: Salekhard env., 66.6 ° N, 66.8 ° E, N. Vikhrev, M. Yanbulat, 16– 19.07.2019, 4♂, 4♀ ( ZMUM); Muzhi vill. (65.4 ° N, 64.7 ° E), bank of Ob R., K. Gorodkov, 23.08.1976, 2♂, 2♀ ( ZIN) .

Yaroslavl Reg., Berditsino (57.45 ° N 40.11 ° E), A. Yakovlev, 14.07.1907, 1♂ ( ZIN) .

TAJIKISTAN: Dushanbe env. (38.5 ° N, 68.7 ° E), E. Gurjeva, 2.04.1962, 1♀ ; Gorno-Badakhshan Reg., Shakhdara R. near Roshtqala (≈ 37.2 ° N, 72.0 ° E), E. Nartchuk, 15.07.1965, 1♂ (all ZIN) .

TURKEY: Ankara Prov., Karagol L. env., 1540 m, 40.346 ° N, 31.929 ° E, N. Vikhrev, 2.09.2009, 3♂, 2♀ ; Antalya Prov., Titreyen L., 36.757 ° N, 31.455 ° E, N. Vikhrev, 1– 4.04.2006, 3♂, 2♀ ; 27– 30.03.2007, 4♂, 1♀; 27.09.2007, 2♀ ; 4.10.2007, 1♂; A. Ozerov, 5.10.2007, 1♀ (all ZMUM) ; bank of Manavgat R., 36.871 ° N, 31.527 ° E, N. Vikhrev, 29 March 2007, 1♀; A. Ozerov, 3.10.2007, 1♂; Mersin Prov., near Silifke , 36.31 ° N, 34.01 ° E, N. Vikhrev, 22.04.2010, 2♂, 1♀ (all ZMUM).

TURKMENISTAN, Lebap Prov., Charjou (= Turkmenabat) env. (≈ 38.76 ° N, 64.20 ° E), A. Ozerov, 25.04.1990, 1♀ ( ZMUM); Dasoguz Prov., Tashauz (= Dasoguz, 41.8 ° N, 60.0 ° E), lower Amu-Daria River , Ushinsky, 21– 24.04.1931, 3♂, 3♀ ( ZIN) .

UKRAINE, Kherson Reg, Aleshky, lower Dnepr River (46.64 ° N, 32.72 ° E), L. Zimin, 26.07.1926, 1♀ ; 18.08.1926, 1♀ ( ZIN) .

UZBEKISTAN: Tashkent Reg.: Karjantau Ridge (≈ 41.83 ° N, 69.84 ° E), K. Obukhova, 22.07.1938, 1♀ ; 30.07.1938, 1♂ ( ZMUM); Chinaz , Syr-Darya R. (40.9 ° N, 68.7 ° E), F. Dobrzhansky, 28.04.1925, 1♂ ( ZIN); Xorazm Reg., Khiva (41.4 ° N, 60.4 ° E), L. Zimin, 21– 27.04.1927, 2♀ ( ZIN) .

VIETNAM, Lao Cai Prov., Sin Chai , 1400 m, 22.338 ° N, 103.808 ° E, A. Ozerov, 15.04.2012, 1♀ ( ZMUM) .

Distribution. Remarkably widely distributed in the Palaearctic region from west to east, south to the Near East and Egypt, north to the Polar Circle; also recorded from North Vietnam. Has been recorded in the mountains up to 2000 m asl (Buryatia).

Notes on synonymy of Ditaeniella patagonensis . Malloch (1933) synonymised D. patagonensis Macquart, 1851 with D. grisescens Meigen, 1830 after examination of material from Argentina, Chile and Uruguay. Steyskal (1963) refuted this synonymy and proposed D. patagonensis as a valid species. Steyskal’s (1963: 121, figs. 6–10) drawings of male genitalia of species of Ditaeniella ( Figs. 7–9 ) were provided with no explanation as to which differences were diagnostically important. In the figures, the postabdomens of D. patagonensis and D. grisescens appear quite similar, while that of D. parallela differs slightly, for example, in D. patagonensis and D. grisescens the postgonites bear several spines, rather than a single spine as in D. parallela . Thus, Steyskal’s drawings did not convince us that the differences between D. patagonensis and D. grisescens are significant. We agree with Malloch's synonymy that D. grisescens = D. patagonensis . If this synonymy is confirmed, it will turn out that D. parallela occurs in North America, whereas D. grisescens occurs in Eurasia and South America. The only reasonable explanation for this disjunct distribution would be that D. grisescens was introduced relatively recently from Eurasia to South America.

Figures

Figs. 7–9. Postabdomen ofDitaeniella species in lateral view (modified from Steyskal 1963: figs. 8–10): 7 — D. grisescens; 8 — D. patagonensis; 9 — D. parallela Рис. 7–9. ПостабΔомен таксонов Ditaeniella, виΔ сбоку (по Steyskal 1963: figs. 8–10): 7 — D. grisescens; 8 — D. patagonensis; 9 — D. parallela

Figs. 12–15. Ditaeniella milleri sp. nov., male (12, 13): 12 — ventral spines at apex of anterior femora; 13 — postgonites; Ditaeniella grisescens, male (14, 15): 14 — ventral setae at apex of anterior femora; 15 — postgonites Рис. 12–15. Ditaeniella millerisp. nov., самец (12, 13): 12 — вентраΛьные шипы на переΔнем беΔре; 13 — постгониты; Ditaeniella grisescens, самец (14, 15): 14 — вентраΛьные шипы на переΔнем беΔре; 15 — постгониты