Sedum marmorense Otting & R. E. Brainerd, 2018
publication ID |
https://doi.org/ 10.11646/phytotaxa.368.1.1 |
persistent identifier |
https://treatment.plazi.org/id/03888A7D-5332-1F3F-FF2D-DC36FCE4FDCE |
treatment provided by |
Felipe |
scientific name |
Sedum marmorense Otting & R. E. Brainerd |
status |
sp. nov. |
7. Sedum marmorense Otting & R. E. Brainerd View in CoL , sp. nov. Figs. 32 G–L View FIGURE 32 , 33 View FIGURE 33
Species Sedum oblanceolatum proxima sed differt foliis basalibus rosulatis obovatis, a Sedum oregonensis rosulis densissimis et inflorescentiis granularibus ceraceisque.
Type: — UNITED STATES. California: Siskiyou Co., Klamath Mountains, Marble Mountain Wilderness Area , Upper Wright Lake , 2295 m, 12 July 2013, R. E . Brainerd 2180 & N . Otting (holotype, OSC; isotypes, GH, NY, RSA, UC, WTU) .
Rhizomes and stolons 1–6.5 cm long, 1.2–3.5 mm in diameter. Rosettes not numerous, 17–36 mm diameter, densely arranged, generally without visible internodes in exposed situations. Rosette leaves green or glaucous, but not granular waxy, often with red margins, aging red then orange, strongly flattened dorsiventrally, obovate, oblanceolate, or lanceolate, 6–24 × 3–11 mm, length to width ratio 1.3–2.8; base cuneate; tip retuse to rounded. Stem leaves ascending to slightly spreading, slightly to strongly glaucous, but not granular waxy, green, pink, or red; ovate, obovate, oblong, or occasionally lanceolate, 7–10 × 2–6 mm, length to width ratio 1.3–3.6, flattened, base truncate; tip acute to rounded or shallowly retuse. Fertile stems green, pink, or red, 4.4–15 cm, nodding in bud, erect in flower and fruit. Inflorescences (0.5–)1.4–6.0 × (0.5–) 1.8–4.5 cm, length to width ratio 0.7–1.8, often globose to subglobose, occasionally more elongated, usually densely flowered, paniculate cymes. Proximal branches 3–7, ascending, (0.3–) 1–2.8 cm, solitary at nodes; all branches with granular waxy flakes early in season. Inflorescence bracts resembling stem leaves but smaller, 4–9 × 1–3 mm, length to width ratio 2–4; bases truncate, tips (acute to) obtuse or rounded. Flowers 6–42 per inflorescence. Calyx green, with granular waxy flakes early in season, 3.5–5.2 × 2.7–4.3 mm; 32–69% as long as perianth; sepals fused basally for 0.5–1.5 mm, lobes 2.4–4.7 mm, acute, less often obtuse, the very tip rounded. Petals 5, 5.3–7.5 mm, 1.4–2.8 mm wide at midlength, fused at base for 0.9–2.5 mm, ascending at an angle of 5–20° from the floral axis at anthesis, white, cream, or light yellow, aging pale yellow or pink; midvein green, cream, or pale yellow, often aging pink; apiculate, the mucro 0.1–0.3 mm. Stamens 10, shorter than the petals; filaments cream or yellow, often almost transparent, aging red; anthers 0.8–1 × 0.3–0.6 mm (when dried), yellow, aging red-orange or reddishbrown, papillose at 20×. Nectaries shallowly crescent-shaped, sunken in middle, yellow, 0.5–0.7 mm, 0.1 mm high. Ovaries 3.8–8.5 mm, erect, fused for 0.7–1.5 mm, maturing into 5 dark red-brown, erect follicles. Follicles 6.6–7.2 mm, with erect to slightly curved style remnant forming a narrow beak approximately 2 mm; bases fused for 0.8–1.5 mm. Seeds 12–31 per follicle, brown, narrowly elliptic, shiny, longitudinally striate, 0.7–1 × 0.15–0.3 mm (including stipe), stipe 0.05–0.1 mm.
Paratypes: — UNITED STATES. CALIFORNIA. Siskiyou County: Marble Mountains , 8 August 1939, Howell 15141 ( CAS, POM) ; Marble Mountains , NE of Red Mountain, 2345 m, 12 July 2013, Brainerd 2185 & Otting ( OSC) ; Tom Martin Peak , 2023 m, 11 July 2014, Otting 3837 ( CAS, MO) ; Lake Mountain , 2080 m, 6 July 2014 m, Otting 3831 & Brainerd ( OSC) ; Lake Mountain , 2034 m, 6 July 2014, Otting 3832 & Brainerd ( CAS, JEPS, WTU) .
Distribution and ecology: —Endemic to a narrow zone 19 km long in the Marble Mountains, in the Klamath Ranges of Siskiyou Co., California ( Fig. 22 View FIGURE 22 ). We uncovered three extant populations, at elevations of 2023–2345 m, on broad flat stony crests to moderate slopes. Aspects varied from south to west to north. Flowering was observed in July, but this was following winters with very low snow cover. The sites were dry sunny cracks and crevices on rocky ridgelines, in low ledges or massive outcrops, as well as in soil pockets on a talus slope among boulders. The underlying bedrock was metasedimentary, metavolcanic, and ultramafic, according to geologists on the Klamath National Forest (Juan de la Fuente and Derek Beal, pers. comm.). Some common associates included: Achillea millefolium L., Arctostaphylos sp. , Calamagrostis sp. , Eriogonum umbellatum Torr. , Festuca roemeri (Pavlick) E.B. Alexeev , Heuchera sp. , Holodiscus discolor (Pursh) Maxim. , Lomatium macrocarpum (Hook. & Arn.) J.M. Coult. & Rose , Myriopteris gracillima (D.C. Eaton) J. Sm. , Penstemon deustus Douglas ex Lindl. , Polystichum sp. , and Pseudoroegneria spicata (Pursh) Á. Löve.
Etymology: — Sedum marmorense , or Marble Mountains stonecrop, was named for the Marble Mountains and for the marble substrate on which it sometimes grows.
Notes: — Sedum marmorense ( Fig. 32G–L View FIGURE 32 ) resembled S. oregonense , but the former has waxy granules on the calyces and inflorescence branches, and was a more delicate plant with looser rosettes. Sedum marmorense differed from S. albomarginatum and S. oblanceolatum in its relatively wide rosette and stem leaves ( Fig. 33E–F View FIGURE 33 ), and in the relative difference in leaf shape and size between the rosettes and stems ( Fig. 33H View FIGURE 33 ). Sedum oblanceolatum and S. albomarginatum had oblanceolate foliage that was quite similar on the stems and the rosettes (e.g., Fig. 21B View FIGURE 21 ). As with some farinose species of North American primrose ( Primula Linnaeus 1753: 142 ), the waxy particles can melt away from specimens dried with heat, complicating identification.
R |
Departamento de Geologia, Universidad de Chile |
E |
Royal Botanic Garden Edinburgh |
N |
Nanjing University |
OSC |
Oregon State University |
GH |
Harvard University - Gray Herbarium |
NY |
William and Lynda Steere Herbarium of the New York Botanical Garden |
UC |
Upjohn Culture Collection |
WTU |
University of Washington |
CAS |
California Academy of Sciences |
POM |
Pomona College |
NE |
University of New England |
MO |
Missouri Botanical Garden |
JEPS |
University of California |
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