Ephippiochthonius cabreriensis ( Mahnert, 1993 ) Mahnert, 1993

Zaragoza, Juan A., 2017, Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae), Zootaxa 4246 (1), pp. 1-221 : 58

Ephippiochthonius cabreriensis ( Mahnert, 1993) , n. comb.

Chthonius (Ephippiochthonius) cabreriensis Mahnert 1993: 972 ‒973, figs 1‒4. Chthonius (Ephippiochthonius) cabreriensis: Zaragoza 2007: 15 .

Type locality. Spain, Balearic Islands province, Cabrera archipelago, Conillera Island (39°11′06″N, 02°57′48″E; 72 m a.s.l.) .

Distribution. Spain: Balearic Islands.

Diagnosis (♂). A small, epigean Ephippiochthonius species tentatively assigned to the tetrachelatus -group. Movable cheliceral finger without isolated subapical tooth (di) and with flattened spinneret in male (unknown for female); two pairs of eyes with lenses, anterior margin of carapace with 1 preocular microseta on each side, posterior margin with 2 medial macrosetae and 2 lateral microsetae; chelal hand depressed at level of ib / isb, with a long and rounded hump distad of ib / isb and an abrupt slope between trichobothria ib / isb and eb; fixed chelal finger with 16 teeth; distal half of movable chelal finger with 7 pointed teeth, proximal half without marginal lamina, with 5 vestigial teeth; pedipalpal femur (♂) 5.0 times longer than broad, length (♂) 0.34 mm; chela (♂) 4.6 times longer than deep, length (♂) 0.51 mm; ratio movable chelal finger/chelal hand (♂) 1.4.

Material examined. None. Holotype lost (see Remarks).

Description (♂; ♀ unknown). Small, epigean species. Integument pigmented.

Carapace slightly longer than broad and weakly constricted posteriorly; anterior eyes with convex lens, distance from anterior margin of carapace equal to their diameter, posterior eyes with more weakly convex lens than anterior pair. Chaetotaxy: 18 macrosetae, 1 preocular microseta on each side and 2 lateral microsetae in posterior row, formula m 4m:6:4:2:m 2m.

Cheliceral hand with 6 setae and 1 lateral microseta. Fixed finger with 7 teeth. Movable finger without an isolated subapical tooth (di), with 5 teeth; spinneret flattened in male. Rallum with 11 blades.

Chaetotaxy of sternites II–V: 9:(3)10(3):(1)9(1):8; in addition, genital notch of male flanked by 6 setae on each side and 4+4 internal glandular setae. Anal cone probably with 0+2 setae [ 2+2 according to Mahnert (1993)].

Pedipalpal coxa with 5 setae (including 2 on manducatory process); coxa I 3 + 3 marginal microsetae; II 4 + 5 bipinnate coxal spines, III 5 + 4 bipinnate coxal spines and IV 6; intercoxal tubercle bisetose.

Pedipalp with femoral chaetotaxy 3:6:2:5:1. Chelal hand depressed at level of ib / isb, with a long hump distad of ib / isb and an abrupt slope between trichobothria ib / isb and eb. Fixed finger with 16 teeth, dental row reaching up to level of sensilla pc; one pair of short, close together, antiaxial sensory setae (as) at the base, distance between them markedly shorter than the fixed finger depth at the base. Distal half of movable finger with 7 pointed teeth; proximal half without marginal lamina, with 5 vestigial teeth, dental row reaching up to trichobothrium sb; basal apodeme long; coupled sensilla pc between sb and b, slightly closer to s b. Trichobothrium ist level with or slightly distad of esb; distance between ib / isb and base of hand equal to that between ib / isb and esb; distance between st and sb 2.8 times longer than that between sb and b.

Measurements and ratios. Male holotype. Carapace 0.31/0.29 (1.1). Pedipalp: femur 0.34/0.07 (5.0), patella 0.14/0.08 (1.8), chela 0.51/0.11 (4.6), hand 0.21 (1.9), movable finger 0.30; ratio movable finger/hand 1.4, femur/ movable finger 1.1, femur/carapace 1.1, chela/carapace 1.6, chela/femur 1.5.

Remarks. The only other species within the tetrachelatus -group that, like E. cabreriensis , has 4 setae in the posterior row of the carapace and lacks an isolated subapical tooth on the movable cheliceral finger is E. serengei n. sp. However, the differences between them are very evident: E. cabreriensis is an epigean species with eyes, 1 preocular microseta on each side of carapace, a pedipalpal femur 5.0 (♂) times longer than broad, length (♂) 0.34 mm, chela 4.6 (♂) times longer than broad, length (♂) 0.51 mm, whereas E. serengei n. sp. is a hypogean species, without eyes or preocular microsetae on the carapace, a pedipalpal femur 6.7 (♂) times longer than broad, length (♂) 0.60 mm, and a chela 6.2 (♂) times longer than broad, length (♂) 0.80 mm. The above description is based on that of Mahnert (1993). The only known specimen of the species, the male holotype, is not present in MHNG (Peter Schwendinger, in litt.), the depositary indicated in the original description, or in MNCM (Guillem Pons, in litt.), and can therefore be assumed lost.