Dasysyrphus venustus (Meigen)

Locke, Michelle M. & Skevington, Jeffrey H., 2013, Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae), Zootaxa 3660 (1), pp. 1-80 : 66-69

Dasysyrphus venustus (Meigen)

Figures: 17A–D, 22

Synonyms and References:

Scaeva arcuata Fallén : 1817: 42 (original description)

Dusek & Laska 1969: 171 (name suppression proposal)

International Commission on Zoological Nomenclature 1972 1972: 9 (name suppression)

Syrphus venustus Meigen 1822: 299 (original description)

Syrphus lunulatus Meigen 1822: 299 (original description)

Syrphus reflectipennis Curran 1921: 157 (original description)

Metasyrphus reflectipennis (Curran) :

Fluke 1933: 109 (key, redescription and transfer to Metasyrphus )

Metasyrphus venustus (Meigen) :

Fluke 1933: 112 (key, redescription and transfer to Metasyrphus )

Dasysyrphus lunulatus (Meigen) :

Enderlein 1938: 208 (transfer to Dasysyrphus )

Stone et al. 1965: 564 (catalogue and transfer back to Dasysyrphus )

Goeldlin de Tiefnau 1974: 208 (lectotype designation)

Syrphella lunulatus (Meigen) :

Goffe 1944: 129 (transfer to Syrphella )

Syrphella venustus (Meigen) :

Goffe 1944: 129 (transfer to Syrphella )

Dasysyrphus arcuatus (Fallén) :

Stone et al. 1965: 563 (catalogue and transfer to Dasysyrphus )

Peck 1974: 128 (key)

Dasysyrphus reflectipennis (Curran) :

Stone et al. 1965: 564 (catalogue and transfer to Dasysyrphus )

Dasysyrphus venustus (Meigen) :

Dusek & Laska 1967: 354 (transfer to Dasysyrphus ?)

Goeldlin de Tiefnau 1974: 208 (lectotype designation)

Vockeroth 1986: 203 (discussion)

Peck 1988: 16 (catalogue)

Vockeroth 1992: 70 (key and redescription)

Mutin & Barkalov 1999: 397 (key)

Stubbs & Falk 2002: 88 (key and discussion)

van Veen 2004: 87 (key)

Barkalov 2007: 294 (key and redescription)

Bartsch et al. 2009: 202 (key and redescription)

Speight 2011: 72 (discussion)

Diagnosis: Abdominal maculae cross abdominal margin; maculae on tergite 2 large (>⅓ width of tergite) ovalshaped; maculae on tergites 3&4 are transverse, constricted slightly medially, with medial edge swollen slightly ( Figs 17A & B ). Similar to D. limatus however maculae on tergite 2 are always small in D. limatus ( Fig. 9A ), less than ¼ the width of the tergite, and are much larger in D. venustus , greater than ⅓ the width of the tergite. Also often confused with species in D. intrudens complex, however, species in this complex have maculae that are arcuate, lunulate and/or greatly constricted medially.

Redescription:

Body Length: 7.9–11.2mm; Wing Length: 7.4–10.4mm

Head: Frons dark with light pollinose fascia (in males it runs along ventral edge of where eyes meet, in females it is ⅓–½ the length between the antennae and ocelli and widely separated medially) and dark pile; face light with dark vitta ¼–⅓ the width of the face, usually reaching antennal socket, face with light to dark pile or combination of light and dark pile; gena dark with dark to light pile or a combination of light and dark pile; occiput dark, covered in light pollen, pile light; scape and pedicel light, flagellomere usually light basally and darker apically, sometimes mostly dark.

Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light with darker anterolateral edges, light pile some with few dark pile along posterior edge; wings can range from being entirely densely microtrichose to having wing cells r 1, br and bm bare base (in br above spurious vein only) and cell cup with small bare areas below vein CuP ( Fig. 17C ); haltere light; pro- and mesofemora ¼–½ dark basially, ½–¾ light apically, metafemur ⅔ dark basially, ⅓ light apically, tibiae light, metatibia with dark band ¼ of the way from the apex, tarsi light, metatarsi dark anteriorly.

Abdomen: Maculae on tergite 2 large and oval with anterolateral edge extending to edge of abdomen, maculae on tergites 3&4 are transverse, constricted slightly medially, with medial edge swollen slightly, almost never meeting medially, maculae cross abdominal margin ( Figs 17A & B ); sternite 2 either all light or with dark fascia (usually distinct, sometimes faint), sternites 3&4 light with dark fasciae.

Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened anteroposteriorly with long pile on posterior side and spines on anteroventral side, similar to D. intrudens complex ( Fig. 7F ) and D. limatus ( Fig. 9D ); cercus oval with long pile over entire surface; gonostylus is pointed ventrally, dorsally flattened at base then it expands dorsally at apex, oblique ridge near apex, with ends of ridge projecting into points; pile on dorsal surface; basiphallus is basally bent at about a 90 degree angle towards dorsum, apical end enlarged and curved ventrally, fully sclerotized all the way around with no pile or spines; distiphallus long with no enlarged area basally, apical end flared out into horn with the dorsal side flattened slightly, which is mostly sclerotized but membranous at apex with setulae dorsally ( Fig. 17D ).

Intraspecific variation: The abdominal maculae of the male are sometimes more arcuate and with a more swollen medial edge of maculae. Wings cells r 1, br, bm, and cup range from being entirely microtrichose to having bare areas.

Etymology: From the Latin venustus , meaning charming, lovely, beautiful, graceful.

Distribution: Canada (AB, BC, MB, NT, NS, ON, QC, YT) and US (AK, ID, ME, MI, NH, NM, NC, PA, UT, VT, VA, WV), also widespread over Europe and Asia ( Fig. 22).

Ecology: Collected from May–August, but more frequently in May and June. Collected at elevations from 260–2957m. Habitats listed on labels include “bog”, “fell-field tundra”, “Mh/Be forest” (mixed hardwood/beech), “pine forest”, “boreal mixed wood forest”, “black spruce forest (25%) with moss”, “Picea-Betula forest”. Specimens have been collected from hilltops.

Discussion: The species concept for D. venustus has been uncertain for quite some time now, especially in the Nearctic. Fluke’s (1933) concept was much like that of this one, distinguishing venustus as having markings that are more straight than specimens in the intrudens complex, which is treated by Fluke as laticaudatus , amalopis (in part) and osburni . Vockeroth (1986) synonymized the intrudens complex under venustus as he believed it was simply a variable species. While abdominal maculae vary greatly, many of the other characters are similar, including the male genitalia.

Dasysyrphus osburni , intrudens , disgregus and laticaudatus were all considered synonyms of venustus by Vockeroth (1992), however these four names fall within what we recognize as the intrudens complex.

The only specimen of D. reflectipennis is that of the holotype (located at the CNC). It is a female with its abdomen broken and glued to a point below the specimen. It was treated as a unique species by Fluke (1933) and by Vockeroth (1986). However, at the time, Vockeroth believed that reflectipennis was a specimen of venustus with aberrant markings. He stated that he was not ready to treat it as a synonym until other specimens were found. Other specimens were not found, but in 1992 he synonymized reflectipennis with venustus . The abdominal maculae of tergite 3 are like that of venustus , however they narrowly meet medially. We have seen no other specimens where the maculae meet, but since the specimen of reflectipennis looks like that of venustus in every other way, it is logical to conclude that Vockeroth was correct in saying that the specimen of reflectipennis is actually a specimen of venustus with aberrant markings.

There are five specimens that appear to be morphologically identical and sympatric with venustus but differ from it in their COI sequences (CNCD 5458, 30889, 30890, 30902, JSS 20982; Fig. 4). It is possible that this is a cryptic species but until more specimens are available to provide more precise geographical and ecological evidence and/or morphological and DNA characters they are retained under the name venustus .

Type Material: From Germany, deposited in the Muséum National d'Histoire Naturelle (The National Museum of Natural History), Paris, France ( Pape & Thompson 2010b). (not examined; requested but not obtained) Material Examined: Canada: Alberta: Itaska Audubon Reserve, Pigeon Lake, [53, -114], 18.v.2008, ♂, JSS 21073, ♀, JSS 21074, F.A.H. Sperling (FAHS). Drayton, [53.220865, -114.978236], 29.vi.1988, ♀, UASM# 149016, E.H. Strickland (UASM). Edmonton, Division No.11, [53.540941, -113.493698], 5.vi.1948, ♀, CNCD 14537, J.R. Vockeroth (CNC); 20.v.1923, ♀, CNCD 14538, E.H. Strickland (CNC); 12.v.1924, ♀, JSS 19540–19545, 19551–19554, 19561–19564, 13.v.1924, ♀, JSS 19559, 19560, 19565, 16.v.1924, ♀, JSS 19558, O. Bryant (CNC). Elk Island National Park, [53.604578, -112.859291], 19.v.1981, ♀, CNCD 14540, J.E. O’Hara (CNC). Bilby, Division No. 13, [53.698357, -114.10198], 1.vi.1924, ♀, JSS 19546, O. Bryant (CNC). British Columbia: CENTRAL KOOTENAY L.D.: Robson, [49.333015, -117.692871], 14.v.1954, ♂, JSS 20034, 17.v.1954, ♂, JSS 20025, 19.v.1954, JSS 20127, 26.v.1954, ♂, JSS 19999, 20.v.1955, ♂, JSS 19998, 7.v.1956, ♀, JSS 20015, 8.v.1956, ♂, JSS 20019, 14.v.1956, 3♂, JSS 20023, 20164, 15.v.1956, ♂, JSS 20132, ♀, JSS 20130, 6.v.1957, ♂, JSS 20012, 11.v.1957, ♀, JSS 20002, 21.v.1957, ♀, JSS 20095, 7.vi.1965, ♀, JSS 20122, 25.v.1965, ♀, JSS 20092, H.R. Foxlee (UBCZ); 14.v.1947, ♂, CNCD 14510, 11.v.1948, ♀, CNCD 14242, 2.vi.1948, ♀, CNCD 14303, H.R. Foxlee (CNC). Miracle Beach, Near Oyster Bay, [49.854574, -125.098409], 13.vi.1955, ♀, CNCD 14314, J.R. McGillis (CNC). Salmon Arm, [50.703923, -119.273924], 2.v.1930, ♀, JSS 20159, Hugh B. Leech (UBCZ). 32 miles South West of Terrace, [54.194076, -129.140296], 30.5m, 8.vi.1960, ♂, CNCD 14509, G.E. Shewell (CNC); 11.vi.1960, 2♀, CNCD 14217, 14341, B. Heming (CNC). Spring Creek, Terrace, [54.539511, -128.618192], 1.vi.1960, ♀, CNCD 14231, C.H. Mann (CNC). Kitsumkalum Lake, 20 Miles North of Terrace, [54.762473, -128.770968], 16.vi.1960, ♀, CNCD 14227, J.G. Chillcott (CNC). Kispiox, [55.348319, -127.687346], 24.v.1958, ♂, JSS 20152, J. Foster (UBCZ). Atlin, [59.5775, -133.69236], 670.56m, 9.vi.1955, 2♀, CNCD 14251, 14252, 20.vi.1955, ♀, CNCD 14223, 6.vii.1955, ♀, CNCD 14284, B.A. Gibbard (CNC); 20.vi.1955, ♂, CNCD 14360, 7.vii.1955, ♀, CNCD 14289, H.J. Huckel (CNC). Blanchard R. Haines Rd., [11.vi.1979, ♀, JSS 20146, G.J. Spencer (UBCZ). Kiskatinaw Public Campground, Alaska Highway DC-20 (31.4km), [26.vi.1978, ♀, JSS 23719, Paul H. Arnaud, Jr. (CAS). Manitoba: Aweme, [49.708529, -99.602758], 19.v.1926, ♀, CNCD 14550, 27.v.1925, ♀, CNCD 14554, 1.vii.1966, ♀, CNCD 14555, R.D. Bird (CNC); 20.vi.1925, ♀, CNCD 14551, R.M. White (CNC). 5 miles South West of Shilo, [49.763964, -99.717403], 28.v.1958, ♀, CNCD 14552, J.F. McAlpine (CNC); 5.vi.1958, ♀, CNCD 14553, C.D.F. Miller (CNC). Erickson, [50.498283, -99.9078], 17.vi.1983, ♀, JSS 20195, D.H. Pengelly (JBWM). CHURCHILL: Burn site N[orth] of Twin lakes, [58.61861111, -93.82888889], 4.vii.2007, ♀, CNCD 9495, 5.vii.2007, ♀, CNCD 9582, J. Skevington (CNC); Bus Stop, [7.viii.2007, ♀, JSS 24644, P. Kevan (CNC). New Brunswick: CHARLOTTE CO.: Chamcook, [45.126407, -67.071693], 28.vi.1965, 3♀, CNCD 14817-14819, G.E. Shewell (CNC). NORTHUMBERLAND CO.: Doaktown, [46.556585, -66.126203], 6.vii.1971, ♀, CNCD 14820, B.V. Peterson (CNC). KENT CO.: Kouchibouguac National Park, [46.819201, -64.96788], 19.vi.1978, 2♀, CNCD 14815, 14816, S.J. Miller (CNC). Snow Cap Park, Hwy 17, [5.vii.1971, ♀, CNCD 14821, B.V. Peterson (CNC). Northwest Territories: Jean-Marie Creek, 42 Miles South West of Fort Simpson, [61.372555, -122.224182], 13.vi.1972, ♀, CNCD 14047, B.V. Peterson (CNC). Coppermine, [62.562732, -115.092764], 30.vi.1951, ♀, CNCD 14058, S.D. Hicks (CNC). Wrigley, Mackenzie, [63.209367, -123.34594], 15.vi.1969, ♀, CNCD 14056, G.E. Shewell (CNC). Nova Scotia: KINGS CO.: [45.017428, -64.610757], 22.vi.1983, ♀, CNCD 14779, 23.vi.1983, ♀, CNCD 14778, J.R. Vockeroth (CNC). COLCHESTER CO., Truro, [45.364959, -63.279846], 7.vii.1919, ♂, CNCD 14769, (CNC). VICTORIA CO., Baddeck, Cape Breton Island, [46.10001, -60.753507], 15.vi.1931, ♀, CNCD 14780, G. Fairchild (CNC); Dingwall, [46.896638, -60.469402], 23.vi.1983, ♀, CNCD 14777, J.R. Vockeroth (CNC); INVERNESS CO., Cape Breton Highlands National Park: French Lake, [46.728452, -60.86483], 24– 30.vi.1984, 2♀, CNCD 14774, 14775, H.J. Teskey (CNC); North Mtn. Bog, [46.805111, -60.688224], 400m, 7.vi.1984, 2♂, CNCD 14759, 14760, B.E. Cooper (CNC); Mackenzie Mountain, [46.773977, -60.818772], 400m, 9.vi.1984, ♂, CNCD 14762, B.E. Cooper (CNC); Pleasant Bay, [46.822961, -60.799065], 6.vi.1984, ♂, CNCD 14765, 11.vi.1984, ♂, CNCD 14763, 12.vi.1984, 3♂, CNCD 14757, 14758, 14766, 2♀, CNCD 14770, 14771, B.E. Cooper (CNC); 25–29.vi.1984, ♀, CNCD 14776, H.J. Teskey (CNC). Ontario: Crawford, [44.253882, - 80.929867], 7–16.vi.1998, 4♀, ROMEnt Spec. No. 57872, 57887–57889, 8–17.vii.1998, ROMEnt Spec. No. 74031, A. Sugar (ROME). Cranberry Lake, Runtz property, [44.44111111, -76.29055556], 18–24.v.2007, 2♀, CNCD 30889, 30890, J. Skevington, H. Douglas, D. Runtz (CNC). LEEDS CO.: Chaffeys Lock, [44.579098, - 76.319816], 28.v.1972, ♀, CNCD 14593, P. Ward (CNC); SIMCOE CO.: Orillia, [44.609505, -79.42068], 26.v.1913, ♀, CNCD 15386, 30.v.1921, ♀, CNCD 14595, 7.vi.1925, ♀, CNCD 14594, C.H. Curran (CNC); Midland, [44.752113, -79.887253], 26.v.1959, ♀, CNCD 14600, J.G. Chillcott (CNC). Griffith, 7 miles East, [45.243279, - 77.031193], 22.v.1982, ♂, CNCD 14567, B.E. Cooper (CNC). Stittsville, [45.263479, -75.925163], 24.vi.1963, ♀, CNCD 14598, W.R.M. Mason (CNC). Nepean, 24 Gervin Street, [45.31694444, -75.72], 28.v.2006 4.vi.2006, ♀, CNCD 30888, J.E. O’Hara (CNC). Mer Bleu, 5 Miles East of Ottawa, [45.40331, -75.5083], 31.v.1923, ♀, CNCD 14579, C.H. Curran (CNC); 5.vi.1966, 2♀, CNCD 14585, 14586, 13.vi.1966, ♀, CNCD 14587, 15.vi.1966, ♀, CNCD 14588, 17.vi.1966, ♀, CNCD 14589, 19.vi.1966, 2♀, CNCD 14590, 1.vii.1966, ♀, CNCD 14592, D.D. Munroe (CNC). Ottawa, [45.411604, -75.688669], 29.v.1925, ♀, CNCD 14571, C.H. Curran (CNC). Waubamick, Parry Sound District, [45.446658, -80.017278], 14.vi.1915, 2♀, JSS 19533, 19534, E.S. Parish (CNC). Dirleton, [45.494944, -76.142874], 29.iv.1968, ♀, CNCD 14599, J.E.H. Martin (CNC). ALGONQUIN PROVINCIAL PARK: Florence Lake [45.44371, -78.49012], 13.v.2007 29.v.2007, ♂, JSS 21385, E. Proctor (CNC); Cecil Lake, [45.4511, -78.4956], 17.v.2007 29.v.2007, ♂, JSS 21371, E. Proctor (CNC); 2 km E of Pondweed Lake, [45.4654, - 78.4298], 16–28.v.2007, 2♂, JSS 21394, 21397, E. Proctor (CNC); Algonquin Park, [46, -78.08333333], 12– 19.vi.2003, ♀, ROMEnt Spec. No. 98532, 13.–20.vi.2003, 7♀, ROMEnt Spec. No. 98522–98525, 98535, 98536, 98540, 17–24.vi.2003, ♀, ROMEnt Spec. No. 98541, 18–25.vi.2003, 3♀, ROMEnt Spec. No. 98526, 98527, 98528, 98533, 20–27.vi.2003, 2♀, ROMEnt Spec. No. 98530, 98531, 10.vii.2003 17.vii.2003, ♀, ROMEnt Spec. No. 98534, Mark Vanderwel (ROME). Atikokan, 14 miles East on Hwy 11, [48.723518, -91.313476], 3.vii.1978 4.vii.1948, ♀, CNCD 14582, H.J. Teskey (CNC). Iroquois Falls, [48.761811, -80.685262], 20.vi.1987, ♀, JSS 20773, J.R. Vockeroth (CNC). LowBush, Lake Abitibi, [48.91608, -80.139852], 6.vi.1925, ♂, BOC 34653, 7.vi.1925, 2♀, ROMEnt Spec. No. 115126, 115127, 13.vi.1925, 2♀, ROMEnt Spec. No. 115128, 115130, 16.vi.1925, ♀, ROMEnt Spec. No. 115131, 17.vi.1925, ♀, ROMEnt Spec. No. 115132, 23.vi.1925, ♀, ROMEnt Spec. No. 115133, 1.vii.1925, 4♀, BOC 34649–34652, 14.vii.1925, ♀, BOC 34654, 18.vii.1925, 4♀, BOC 34655– 34657, ROMEnt Spec. No. 115129, N.K. Bigelow (ROME); 9.vi.1925, ♀, CNCD 14572, 15.vi.1925, ♀, CNCD 14573, 16.vi.1925, 2♀, CNCD 14574–14576, 14.vii.1925, ♀, CNCD 14578, N.K. Bigelow (CNC); 1.vii.1925, ♀, UASM# 149015, N.K. Bigelow (UASM); 1.vii.1925, ♀, JSS 21349, 14.vii.1925, ♀, JSS 21350, N.K. Bigelow (MCZ). Macdiarnid, Lake Nipigon, [49.438216, -88.12695], 7.vi.1921, ♀, BOC 34659, 24.vi.1923, ♀, BOC 34660, N.K. Bigelow (ROME). Quebec: Covey Hill, Le Haut-Saint-Laurent, [45.019681, -73.745746], 20.vi.1924, ♀, CNCD 14755, G.S. Walley (CNC). Aylmer, [45.400224, -75.817137], 28.v.1923, ♂, CNCD 14653, C.H. Curran (CNC). Ste. Anne de Bellevue, [45.406417, -73.950944], 12.v.1965, ♀, LEM 8806, 1.vi.1966, ♀, LEM 8804, W. Boyle (LEMQ); 13.v.1965, ♀, LEM 8807, G. Jamieson (LEMQ); 15.v.1967, ♂, LEM 8805, A. Campbell (LEMQ). Abbotsford, [45.437637, -72.887923], vii.1935, ♀, CNCD 14711, G.E. Shewell (CNC); 25.vi.1937, ♀, CNCD 14725, G. Shewell (CNC); 14.vi.1972, ♀, CNCD 14689, D.M. Wood (CNC). VAUDREUIL CO.: Mount Rigaud, [45.46638889, -74.32638889], 19.v.2004, ♂, CNCD 68, J. Skevington, L. Bartels, E. St. Louis, J. King (CNC); 24.v.2007, CNCD 8656, 8658, 8675, J. Skevington, A. Bailie, R. Farmahin, A. Kealey, S. Kelso, J. King (CNC). LES COLLINES- DES- L'OUTAOUAIS: Summit of King Mountain, Old Chelsea, [45.498058, -75.864116], 351m, 14.vi.1963, ♂, CNCD 14680, 2♀, CNCD 14737, 14738, 21.vi.1966, ♂, CNCD 14651, ♀, CNCD 14686, 13.vi.1970, 6♂, CNCD 14606, 14608–14612, 11.vi.1971, 2♂, CNCD 14654, 14679, ♀, CNCD 14736, 16.vi.1971, ♂, CNCD 14671, 12.vi.1980, ♀, CNCD 14693, J.R. Vockeroth (CNC); 9.vi.1971, ♂, CNCD 14681, D.M. Wood (CNC); Champlain Lookout, Gatineau Park, [45.508314, -75.913273], 335m, 1.vi.1965, ♂, CNCD 14672, J.R. Vockeroth (CNC). Montreal Area, Communaute-Urbaine-de-Montreal, [45.526155, - 73.702807], 6.vi.1964, 2♀, CNCD 14692, 14694, J.W. Boyes (CNC). Mont Saint-Hilaire, La Vallee-du-Richelieu, [45.552948, -73.155276], 411m, 11.vi.1967, ♂, CNCD 14675, J.W. Boyes (CNC). Lac-Megantic, Le Granit, [45.57781, -70.884105], 18.vi.1923, ♂, CNCD 14646, 20.vi.1923, ♀, CNCD 14753, C.H. Curran (CNC); 30.v.1971, ♂, CNCD 14667, D.M. Wood (CNC). Duncan Lake, near Rupert, [45.68138889, -76.05027778], 21.vi.1969, ♀, CNCD 14745, J.F. McAlpine (CNC); 19.v.1987, ♂, JSS 20767, D.M. Wood (CNC). Lac Cloutier, [46.184459, -73.64769], 30.v.1987 7.vii.1987, ♀, LEM 13620, (LEMQ). Schefferville, Iron Arm Fern, [54.805276, -66.805274], 28.vi.1990, ♀, LEM 13928, 3–4.vii.1990, ♀, LEM 13459, P. McElligott (LEMQ). Great Whale River, [55.116816, -76.405554], 7.vii.1949, ♀, CNCD 14718, 12.vii.1949, 3♀, CNCD 14715–14717, 24.vii.1949, ♀, CNCD 14724, 25.vii.1949, ♀, CNCD 14714, 31.vii.1949, ♀, CNCD 14719, J.R. Vockeroth (CNC). Indian House Lake, [56.328482, -64.720845], 8.vii.1954, ♀, CNCD 14721, 10.vii.1954, ♂, CNCD 14683, 17.vii.1954, ♀, CNCD 14722, W.R. Richards (CNC). Kuujjuaq (Fort Chimo), [58.100076, -68.406179], 29.v.1948, ♂, CNCD 14684, H.N. Smith (CNC); 14.vii.1948, ♀, CNCD 14712, 28.vii.1948, ♀, CNCD 14713, R.H. MacLeod (CNC). Saskatchewan: Great Deer, [52.580321, -107.056629], 21.v.1949, ♀, CNCD 14547, J.R. Vockeroth (CNC). Nipawin, Division No.14, [53.365608, -104.011519], 2.vi.1948, ♀, CNCD 14548, J.R. Vockeroth (CNC). Yukon Territory: Otter Lake, [62.486817, -130.4166667], 1219m, 17.vii.1960, ♀, CNCD 13966, J.E.H. Martin (CNC). Sheldon Lake, [62.616763, -131.266603], 1067m, 3.vii.1960, ♀, CNCD 13967, 7.vii.1960, ♀, CNCD 13968, J.E.H. Martin (CNC); 3.vii.1960, ♀, CNCD 13976, 7.vii.1960, 2♀, CNCD 13974, 13975, E.W. Rockburne (CNC). La Force Lake, [62.683333, -132.333333], 1006m, 28.vi.1960, ♀, CNCD 13955, E.W. Rockburne (CNC); 7.vii.1960, ♀, CNCD 13949, J.E.H. Martin (CNC). DEMPSTER HIGHWAY: Mile 40, [64.448494, -138.223783], 1– 6.vii.1973, ♂, CNCD 14029, G. & D.M. Wood (CNC); Engineer Cr., km 152 [65.070828, -138.2399], 30.vi.1982, ♀, JSS 20140, S.G. Cannings, L. Vasington, R.A. Moore (UBCZ); km 416 [66.644581, -136.324471], 750m, 22– 28.vi.1980, ♀, CNCD 14005, Wood & Lafontaine (CNC). Rampart House, [67.421455, -140.983336], 18.vii.1951, ♀, CNCD 13981, C.C. Loan (CNC). Old Crow, [67.571219, -139.833877], 2.vii.1983, ♀, JSS 20138, R.A. Cannings (UBCZ). British Mountains, Firth River, [68.848549, -140.48328], 24.vii.1956, ♀, CNCD 14053, R.E. Leech (CNC); 24.vi.1984, ♀, JSS 18367, S.G. Cannings (CNC). Kluane; Ruby Ranges; Pika Camp, [16.vi.2004 23.vi.2004, ♀, JSS 25586, E.S. Bakker (WVSC). United States of America: Alaska: ANCHORAGE CO.: Anchorage, [61.175333, -149.861715], 13.vi.1921, ♀, JSS 19547, J.M. Aldrich (CNC). MATANUSKA- SUSITNA CO.: Matanuska, [61.541945, -149.229674], 4.vi.1944, ♀, JSS 19557, 10.vi.1944, ♀, JSS 19537, 11.vi.1944, ♀, JSS 19536, 17.vi.1944, ♀, JSS 19538, 1.vi.1945, 2♀, JSS 19535, 19539, J.C. Chamberlin (CNC). Trapper Creek, 36 miles North of Willow, [62.31666667, -150.2166667], 19.vii.1984, CNCD 60064, S. & J. Peck (CNC). Unalakleet, [63.86666667, -160.7833333], 17.vi.1961, ♀, CNCD 26575, B.S. Heming (CNC); 16.viii.1961, 2♀, CNCD 26577, 26578, R. Madge (CNC). SOUTHEAST FAIRBANKS CO.: Big Delta, Lake Boleo, [64.154432, - 145.836524], 19.v.1951, ♀, CNCD 26553, W.R.M. Mason (CNC). NOME AREA: Mile 55 of Kougarok Road, [65.08972222, -164.6722222], 20–26.vi.2005, ♀, CNCD 30893, J.&R. Skevington (CNC); Nome Area, Mile 17 of Kougarok Road, [64.70611111, -165.2972222], 21–23.vi.2005, 2♀, CNCD 30881, 30902, 26–28.vi.2005, 2♀, CNCD 30895, 30897, 30898, J.&R. Skevington (CNC). Colorado: Chambers Lake, [40.603663, -105.853645], 3.viii.1932, ♀, JSS 20714, Chas L. Fluke (CSUC). Idaho: Keene Valley, [44.18333333, -73.78333333], 25.v.1910, JSS 21351, H. Notman (MCZ). Maine: Bar Harbor, [44.38333333, -68.2], ♀, JSS 21348, (MCZ). FRANKLIN CO.: Dryden, [44.586729, -70.209783], 20.vii.1959, ♂, CNCD 26734, G.H. Heinrich (CNC). Massachusetts: HAMPSHIRE CO.: [42.335786, -72.644117],. vi.1922, ♂, CNCD 26739, P.A. Readio (CNC). MIDDLESEX CO.: Melrose Highlands, [42.470929, -71.061441], 16.vi.1911, ♂, CNCD 26740, J.D. Tothill (CNC). Michigan: Manistee Lake, [44.24749, -86.30471], 22.v.1955, ♂, JSS 21138, R.W. Hodges (MSUC). Sagola, [46.09028, - 88.0775], 6.vi.1983, ♀, JSS 21136, M. Arduser (MSUC). Isle Royale, [48, -88.83333333], 3.vii.1957, ♀, JSS 21142, 15.vii.1957, 2♀, JSS 21144, JSS 21147, 18.vii.1957, ♀, JSS 21143, 11.viii.1957, ♀, JSS 21141, R.W. Hodges (MSUC). KALKASKA CO.: 14.vi.1949, ♀, JSS 21139, R.R. Dreisbach (MSUC); 12.vi., ♀, JSS 21140, R.R. Dreisbach (MSUC). CHIPPEWA CO.: 3.vi.1957, 2♀, JSS 21145, 21146, R. & K. Dreisbach (MSUC). CLARE CO.: 16.v.1951, ♀, JSS 21119, 23.v.1959, 2♀, JSS 21148, 21149, R. Dreisbach (MSUC). New Hampshire: Franconia, Rt.3 Fanconia North, [44.209935, -71.685178], 20–21.vi.1972, ♀, USNM ENT 258066, B.J. & F.C. Thompson (USNM). Pittsburg, Rt.3 Connecticut Lakes, [45.104264, -71.259207], 22–24.vi.1972, 4♀, USNM ENT 258067–258070, B.J. & F.C. Thompson (USNM). CHESHIRE CO.: Jaffrey, [42.814957, -72.022399], 1941, ♂, CNCD 26741, C.W. Johnson (CNC). Bretton Woods, [44.25, -71.43333333], 27.vi., ♂, CNCD 26742, C.W. Johnson (CNC). COOS CO.: Mount Washington, [44.270481, -71.30255], 6.vii., ♂, CNCD 26735, (CNC). Lancaster Mt. Prospect, [44.449317, -71.570888], 625m, 19.vi.1982, ♀, JSS 20780, J.R. Vockeroth (CNC). New Mexico: LINCOLN CO.: Sierra Blanca, [33.478065, -105.552907], 2957m, 10–26.vi.1979, ♀, JSS 18336, S. & J. Peck (CNC). New York: ULSTER CO.: Oliverea, [42.065369, -74.460152], 18.vi.1918, ♂, CNCD 26745, (CNC). Maplecrest, Catskill Mts., [42.275919, -74.186809], 18.vi.1927, JSS 19255, F.M. Schott (FMNH). ESSEX CO.: Heart Lake, Mount Jo, [44.182434, -73.966816], 13.vi.1935, ♂, CNCD 26743, C.W. Johnson (CNC). North Carolina: Shining Rock Wild. Area, [35.355, -82.84388889], 23.v.1993, ♀, JSS 20807, Kondratieff & Kirchner (CSUC). Pisgah National Forest, 10km SE Linville on Road, [36, -81.8], 760m, 25.v.1999, ♀, JSS 20988, M.I. Erwin (MHPC). 13km W Boone on Blue Ridge Pkwy, [36.23333333, -81.51666667], 1050m, 24.v.1999, ♀, JSS 20987, 25.v.1999, ♂, JSS 20983, M.I. Erwin (MHPC). Pennsylvania: Black Moshannon State Park, [40.896041, - 78.044398], 13.v.1990, ♂, USNM ENT 258081, FC Thompspn (USNM). CENTRE CO., Bear Meadows Natural Area, 11.v.1983, ♂, CNCD 5458, F.C. Thompson (USNM). Utah: CACHE CO., Tony Grove Crk, [41.892197, - 111.634869], 8–15.vii.1994, ♀, JSS 24642, W.J. Hanson (EMUS). Vermont: ESSEX CO.: 5 Miles West of Bloomfield, [44.753513, -71.727243], 28.vi.1972, 2♀, JSS 18338, 18339, H.J. Teskey (CNC). Virginia: SMYTH CO., Hurricane Creek, 22.v.1981, ♂, CNCD 5456, F.C. & B.J. Thompson (USNM). West Virginia: Standard, off I- 77, [38.136111, -81.400556], 6.v.2005, ♀, JSS 20692, B. Kondratieff, R. Zuelig, R.F. Kirchner (CSUC).

Figures

FIGURE 7. Dasysyrphus intrudens complex: A: female (CNCD 14312) dorsal habitus; B: female (CNCD 14312) lateral habitus; C: female (CNCD 30911) dorsal habitus; D: female (CNCD 30911) lateral habitus; E: male (JSS 23704) dorsal habitus; F: lateral view of male (CNCD 30894) genitalia. Dorsal and lateral habitus scale bar = 1mm, genitalia scale bar = 500µm.

FIGURE 9. Dasysyrphus limatus: A: male (CNCD 13659) dorsal habitus; B: male (CNCD 13659) lateral habitus; C: wing base: (CNCD 13686); D: lateral view of male (JSS 20796) genitalia. Dorsal and lateral habitus scale bar = 1mm, genitalia and wing scale bar = 500µm.

FIGURE 17. Dasysyrphus venustus: A: female (CNCD 30888) dorsal habitus; B: female lateral (CNCD 30888) habitus; C: wing base (CNCD 5458); D: lateral view of male (CNCD 14567) genitalia. Dorsal and lateral habitus scale bar = 1mm, genitalia and wing scale bar = 500µm.