Odontocroton Clarke, 2018

Clarke, Robin O. S., 2018, Provisional revision of the genus Odontocera Audinet-Serville, 1834 (Coleoptera: Cerambycidae). I: exclusions, new rank, synonymies and the description of two new genera, Insecta Mundi 637, pp. 1-27 : 3-8

Odontocroton Clarke , n. gen.

Fig. 1–34

Type species. Odontocera flavicauda Bates, 1873 , here designated.

Description. Slender, lengths forebody/abdomen in male 0.73–0.78, in female 0.82; length body/width metasternum in male 6.45–8.30, in female 6.05 (Group A); or moderately slender, lengths forebody/ abdomen in male 0.87–0.92, in female 0.93–1.00; length body/width metasternum in male 5.42–5.85, in female 5.00–5.36 (Group B).

Structure. Smaller species 9.0– 12.7 mm (Group A), larger species 13.5–17.1 mm, but see O. monnei 11.0 mm (Group B species).

Head. Head (h) narrower or wider than prothorax (p), p/h 0.94–1.06 (Group A), 1.08–1.21 (Group B). Rostrum parallel-sided; comparatively short, width (w)/length (l) 2.30–2.50 (Group A), longer w/l 1.93–2.40 (Group B); usually shorter in males w/l 2.33–2.50 (but 2.08 in O. septemtuberculatus ), usually longer in females, w/l 1.93–2.12 (but 2.30 in O. flavicauda ). Labrum small, apex declivous, almost rectangular, approximately 3× wider than long (in O. flavicauda ), or 2× wider than long (in most species). Surface of clypeus separated from frons by strong transverse ridge and adjacent declivity (Group A); surface of clypeus convex (lacking ridge), frons depressed (Group B); clypeus almost impunctate, that of frons strongly ornamented (in most species), or both similarly ornamented (in O. quinquecallosus ). Coronal suture only traceable on interocular (in O. flavicauda , O. melzeri ), or extended as narrow stria and impunctate line beyond antennal tubercles (in O, soror , O. apicalis , and males of O. quinquecallosus , O. rufifrons , and both sexes of O. septemtuberculatus ), or frontal suture complete, from near apex of frons to beyond antennal tubercles (in females of O. quinquecallosus , O. rufifrons ). Area of submentum transverse, flattened, not well delimited at sides; separated from gula by distinct declivity.

Eyes. Inferior lobes of eyes large (in males of Group A, most males of Group B, and female O. septemtu- berculatus), smaller (in male O. rufifrons ), or much smaller (in most females); contiguous to subcontiguos in male, width of one lobe/interocular 4.83–11.15, in female 0.83–1.40; strongly convex laterally (Group A, in male O. rufifrons and both sexes of O. quinquecallosus , O. septemtuberculatus ), or moderately so (in male O. apicalis and female O. rufifrons ); mesally slightly more prominent than interocular in male, in female about coplanar with interocular; distal margins placed on frons (in most males), or adjacent to genae (in most females, and male of O. septemtuberculatus ); proximal margins weakly oblique. Superior lobes rather short and lobate, not widely separated, interocular distance/width of one lobe 1.80–3.33 in male, 2.20–2.75 in female; mesally with 10–12 rows of ommatidia (in both sexes of O. flavicauda and males of O. melzeri , O. soror ), or 13–14 rows (in male O. apicalis , and both sexes of O. quinquecallosus , O. septemtuberculatus , O. rufifrons ); laterally reduced to 4–5 rows (in O. soror ), 6–7 rows (in males of O. flavicauda , O. melzeri and both sexes of O. quinquecallosus ), or 8–9 rows (in female O. flavicauda , male O. apicalis and both sexes of O. rufifrons , O. septemtuberculatus ).

Antennal tubercles. Moderately prominent, but rounded, not widely separated; distance between tubercles/width of scape 1.67–2.25 (narrowest in O. soror , widest in female O. septemtuberculatus ).

Antennae. Filiform (Group A), subfiliform to subcrassate (Group B), moderately short, reaching api- cal third of urosternite II to base of III (Group A and male O. septemtuberculatus ), reaching middle of urosternite I to middle of II (Group B most species). Antennomeres VI to X the widest (in most species), V–X (in O. monnei ), VII–X (in O. melzeri ); with acute, clipped serrations. Scape variable, pyriform to subpyriform (in most species), short and cylindrical (in O. monnei ); from 2.3× longer than pedicel (in O. apicalis ) to 3.3× longer (in O. septemtuberculatus ); approximately 1/3 shorter than antennomere III (in most species), approximately 1/5 shorter (in O. rufifrons ). Antennomere III: approximately 1/3 longer than any other segment (in females of O. rufifrons , O. septemtuberculatus ), 1.1–1.2× longer than any other segment (in most species), equal in length to V, or V and VI (in males of O. rufifrons , O. flavicauda ); almost cylindrical, narrower than pedicel at base; slightly wider than pedicel at apex (in O. flavicauda , O. apicalis , O. rufifrons ), not wider than apex of pedicel (in O. melzeri , O. soror , O. quinquecallosus , O. septemtuberculatus ). Male. IV and V gradually widened to apex (Group A species and O. apicalis , O. sanguinolentus ), hardly widened to apex (in remaining Group B species); V not subserrate, (in Group A species, and O. quinquecallosus , O. septemtuberculatus ), V subserrate (in O. apicalis , O. rufifrons ). VI–X narrower at base, moderately widened to apex; incrementally shorter (in most species), or nearly so (in male O. flavicauda ,), but not incrementally shorter, lengths VI 1.5, VII 1.6, VIII 1.4 (in O. melzeri ). XI subcylindrical, short (but longer than X); with longer narrow cone (in Group A species and O. septem- tuberculatus); with shorter, broad apical cone (in most Group B species); cone centered (in most Group A species and O. apicalis ), cone slightly displaced to one side, leaving lateral margin somewhat dentate (in O. melzeri and most Group B species). Female. basal segments tend to be longer than in male, and apical ones shorter; IV–X similar structure to males (in Group A species and O. septemtuberculatus ), but somewhat tumid compared with males (in O. quinquecallosus , O. rufifrons ); XI not dentate as apical cone not displaced (Group A), or also dentate, if only weakly so (Group B).

Prothorax. Cylindrical; but sides wider at middle, before and after middle, and thereby multi-sinuate (in Group A, and O. quinquecallosus , O. septemtuberculatus ), or with sides straight for apical half, sinuate for basal half (in O. apicalis ), or sides of prothorax regularly rounded from apex to basal constriction (in O. monnei , O. sanguinolentus , O. rufifrons ); in male: length (l)/width (w) 1.23–1.27 (Group A, and O. apicalis ), l/w 1.09–1.14 (Group B); in female: l/w 1.00-1.14; in male: widest from middle to before middle, prothoracic quotient 2.00–2.20; in female: widest at middle to well in front of middle, quotient approximately 2.00–2.46 (Group A), or near middle, quotient 1.92–2.19 (in Group B species). Surface of pronotum not callose (in Group B (i) species), or callose (in Group A and Group B (ii) species, weakly so in O. melzeri , O. quinquecallosus ), with narrow callus at midline, to either side with paired, somewhat fused, protuberant calli, and lateral profile with four indentations produced by three carina overhang- ing sides (in Group A species), or callosities similar to above, but lateral carina absent and profile bisinuate (in Group B (ii) species). Apical constriction evident (Group A, and in O. septemtuberculatus ), evanescent or absent (in most Group B species). Basal constriction declivous laterally, medially sloping to pronotal disc (in Group A and O. septemtuberculatus ), or hardly declivous (in O. quinquecallosus ), or basal constriction semi-evanescent (in O. apicalis , O. rufifrons ); towards sides with deep, rounded fossa (Group B), or not fossate (Group A). Apical border narrow and prominent; front margin slightly narrower than basal margin (in most species), or slightly wider than basal margin (in female O. quinquecallosus ).

Prosternum. Declivous across apical third, weakly inclined to base of prosternal process (in O. flavi- cauda, O. melzeri , O. septemtuberculatus ), or moderately strongly inclined (in O. soror , O. apicalis , O. quinquecallosus , O. rufifrons ). Base of prosternal process: short, arced (in nearly all species), or flat (in female O. flavicauda ); moderately broad, approximately 5–7× narrower than width of procoxal cavity (in both sexes of O. quinquecallosus , in females of O. rufifrons and O. septemtuberculatus ); moderately narrow, approximately 9–12× narrower than procoxal cavity (in both sexes of O. flavicauda , and males of O. soror , O. apicalis , O. rufifrons , and O. septemtuberculatus ); narrow, approximately 16× narrower than procoxal cavity (in male O. melzeri ). Apex of prosternal process a large equilateral triangle, with or without slightly rounded sides; postcoxal process narrow (in female O. flavicauda , O. melzeri ), or moderately narrow (in male O. flavicauda , O. soror , and all Group B species). Procoxal cavities plugged at sides, firmly closed behind.

Mesothorax. Mesosternum deep and abruptly declivous. Base of mesosternal process with weakly raised sides; comparatively narrow, 3–6× narrower than width of coxal cavity (Group A), or broader, approximately 2× narrower than width of coxal cavity (Group B); apex of process (best illustrated in Figure 28 of male O. quinquecallosus ) short, broad and Y-shaped, moderately and regularly widening from apex of base into two short, truncate lobes (these separated by a short V-shaped excavation), the apical margin of each lobe oblique. Mesocoxal cavities narrowly open to mesoepimeron, the latter narrow and irregularly widened at apex. Mesothorax shorter than metathorax, lengths meso-/metathorax 0.67–0.84 in male, 0.63–0.73 in female. Scutellum . Small; oval or scutate.

Elytra. Strongly subulate, narrow, and nearly flat; humeri weakly prominent, not projecting, broad (hiding mesepimeron when viewed from directly overhead), with narrow, slightly raised, smooth line running towards apex of elytra (which might represent the remains of a humero-apical costa); elongate, apex reaching from apex of urosternite III to apex of IV (Group A); apex reaching from middle of IV to middle of V (Group B); in male, 3.76–4.12× longer than width of humeri, in female: 3.69× longer (Group A); in male, 3.56–3.84× longer than width of humeri, in female, 3.45–3.84× longer (Group B); variably dehiscent or narrowly fissate (in most species) or not dehiscent nor fissate (in O. flavirostris ). Each elytron variable, but narrowed for, at least basal half (usually more) and almost parallel-sided towards apex; sides arced from behind humerus to apex (in O. flavicauda , male O. rufifrons ), or weakly recurved for pre apex (in female O. rufifrons , and all other species); apical margin obliquely truncate (in most species), weakly armed at sutural apex (in male O. flavicauda , O. soror ,), or weakly armed at both sutural and lateral apex (in most Group B species), or unarmed (in female O. flavicauda , O. melzeri , O. rufifrons ).

Metathorax. Sides moderately rounded and weakly oblique to middle of apex (in males of O. flavicauda , O. soror , O. rufifrons ), or sides rounded and sinuate to middle of apex (in female O. flavicauda and both sexes of O. quinquecallosus and O. septemtuberculatus ), or sides subparallel and rounded behind to middle of apex (in O. melzeri , O. apicalis ), or sides subparallel, but oblique to apex (in female O. rufifrons ). Metasternum in male: tumid (in most males), mod tumid (in O. soror , O. apicalis ); in female: tumid, not flattened (Group A and O. septemtuberculatus ), tumid, moderately flattened (in most Group B species); more or less coplanar with mesocoxae; metepisternum cuneate, not broad at base, acuminate at apex in both sexes of most species, more parallel-sided in O. soror .

Abdomen. Group A male ( Fig. 4 ). Cylindrical, hardly annulate (in O. melzeri ), or subcylindrical, distinctly annulate (in O. flavicauda , O. soror ); narrow, at its widest distinctly narrower than width across mesocoxal cavities, widest at apex of urosternite II (in O. flavicauda , O. melzeri ), or III (in O. soror ); urosternite I longest, II–IV subequal, lengths urosternites I/V 1.89–1.94. Urosternite V: trapezoidal, transverse (in O. flavicauda ), quadrate (in O. melzeri , O. soror ); soleate depression represented by wide, flattened area, with strongly raised sides (in O. flavicauda , O. melzeri ), or weakly raised sides (in O. soror ). Abdominal process triangular, with 45–60° slope (in O. flavicauda , O. melzeri ), or vertical (in O. soror ).

Group B male ( Fig. 18 , 22 , 28 , 32 ). Cylindrical, distinctly annulate; broad, at its widest about as wide as width across mesocoxal cavities, widest at apex of urosternite III; urosternite I longest, II–IV sequentially shorter, lengths urosternite I/V 2.11–2.44. Urosternite V: trapezoidal, transverse; soleate depression represented by wide, flattened area, with strongly raised sides towards apex (in O. apicalis , O. quinquecallosus ), or moderately raised, rounded sides (in O. rufifrons ), or weakly raised sides (in O. septemtuberculatus ). Abdominal process triangular, with 45–60° slope (in most species), or almost vertical (in O. rufifrons ).

Group A female ( Fig. 6 , 8 ). Fusiform; hardly annulate; at its widest (middle of urosternite II) about as wide as width across mesocoxal cavities (in O. flavicauda ), or at its widest (apex of urosternite II) wider than width across mesocoxal cavities (in O. flavirostris ); uniformly tapering from near apex of II to apex of V; V conical and subacuminate at apex (in O. flavicauda ), or trapezoidal and truncate at apex (in O. flavirostris ). Abdominal process triangular with 30° slope (in O. flavicauda ).

Group B female ( Fig 24 , 30 , 34 ). Fusiform; distinctly annulate; at its widest (middle, or apex of urosternite III) distinctly wider than width across mesocoxal cavities, irregularly and abruptly narrowed to apex of V; V trapezoidal, broadly rounded at apex. Abdominal process triangular with 20–30° slope (in O. quinquecallosus , O. septemtuberculatus ), or approximately 45° slope (in O. rufifrons ).

Legs. Front leg. Femoral peduncles short, approximately 1/10 length femora; clave fusiform, tumid mesally (hardly so laterally) and abrupt; tibia narrow at base, regularly, but weakly widening to apex; apex hardly excavate laterally, mesally without tooth. Middle leg. Femur relatively long, 1.20–1.40× longer than tibia; peduncle flattened; clave fusiform, distinctly longer than peduncle, lengths clave/ peduncle in male 1.56–1.59, in female 1.35 (Group A), or clave/peduncle in male 1.52–1.78, in female 1.50–1.72 (Group B); clave tumid mesally (when viewed from above), moderately wide dorso-ventrally, length of femur/width of clave 3.24–3.95. Tibia cylindrical, straight, narrow, gradually widening from base to apex. Hind leg. Apex of femur reaching from near apex of urosternite III to basal quarter of IV; femoral peduncle cylindrical, narrow. Tibia cylindrical; straight when viewed from directly behind (in O. flavicauda , O. flavirostris , O. melzeri , O. quinquecallosus ), or sinuate (in O. soror , O. apicalis , O. rufifrons , O. septemtuberculatus ); straight when viewed from the side (in O. flavirostris , O. soror , O. apicalis , O. septemtuberculatus ), or sinuate (in O. flavicauda , O. melzeri , O. quinquecallosus , O. rufifrons ); shorter than femora, lengths femora/tibia 1.00–1.08 (in most species), 1.14 –1.16 (in O. melzeri , O. soror ); usually weakly, or not at all thickened from base to apex; moderately densely setose for apical half (the setae too short to be called a brush). Tarsi. Protarsus shorter than mesotarsus, but similar in structure; moderately slender (Group A), or more robust (Group B), widening to apex; tarsomere I subcylindrical, II trapezoidal, transverse, III wide and strongly bilobed (and larger than those of metatarsus). Metatarsus differs from pro- and mesotarsi by much longer first tarsomere, and slightly longer second tarsomere; metatarsomere I cylindrical, long, 1.40–1.55× longer than II+III; II trapezoidal, or cylindrical (in O, septemtuberculatus ), weakly pediculate; III smaller than those of front and middle legs; onychium long and slender (Group A and in O. septemtuberculatus ), or more robust (in rest of Group B).

Group A. Slender and, given the smaller size of the species (approximately 12 mm), relatively long (especially hind leg), ratio of length from front to hind leg 1.0:1.3:2.5–2.6. Metafemoral clave subabrupt, fusiform, shorter than peduncle, lengths clave/peduncle 0.75–0.91.

Group B. Less slender and, given the larger size of the species (13–17 mm), relatively short (especially hind leg), ratio of length from front to hind leg 1.0:1.3–1.4:2.1–2.3. Metafemoral clave not abrupt, cylindrical, longer than peduncle, lengths clave/peduncle 1.20–1.40.

Male genitalia ( Fig. 44–50 ). Median lobe (length approximately 1.7 mm) moderately broad and arced, acuminate to apex (when viewed laterally); internal sack seems to lack dark bodies. Tegmen. Group A. Less chitinized; with paired, parallel, finger-like lobes. Each lobe moderately long (approximately 1.3 mm) and narrow, 3–4× longer than basal piece; parallel-sided, apex blunt; apex (and mesal side in O. melzeri ) clothed with sparse, straight setae. Basal piece short, transverse. Y-piece moderately long, not much shorter than lateral lobes; fork not much longer than stem. Group B. More chitinized, with paired, divergent, thumb-shaped lobes. Each lobe long (approximately 1.7 mm) and broad, 2–3× longer than basal piece; widest at middle, mesal side arced from base to apex, lateral side nearly straight; apex sub-acuminate (or blunt in O. septemtuberculatus ); apex and sides liberally clothed with long setae (those at apex slightly longer and curved). Basal piece moderately long, weakly transverse; Y-piece moderately short, fork distinctly longer than stem.

General pubescence. Reduced in males (of O. flavicauda , O. quinquecallosus , O. septemtuberculatus ), or less so (in O. melzeri , O. soror , O. apicalis , O. rufifrons ); in female almost glabrous (in O. flavicauda ), or as in male (in O. quinquecallosus , O. rufifrons , O. septemtuberculatus ). Dorsad: fimbriate below inferior lobes of eyes. Sides of pronotum with isolated groups of setae laterally. Scutellum clothed with short, recumbent, white or silvery hairs; elytra usually with few short setae basally, or absent (in O. apicalis , O. rufifrons ). Underside: prosternum moderately densely clothed with short, untidy, semierect setae; sides of metasternum, and metepisternum with similar setae, but less dense and not untidy (in most species), or hardly setose (in O. rufifrons , O. septemtuberculatus ). Short, moderately dense, recumbent, white pubescence, or ochreous (in O. soror ) covering metepisternum and sides of metasternum; in female recumbent pubescence much reduced (in O. flavicauda , O. quinquecallosus ). Abdomen in male glabrous at base, towards apex incrementally setose (Group A males); or almost entirely glabrous (in Group B species and female O. flavicauda ). Antennae (Group A) micropubescent towards apex; basal segments fimbriate below, from scape to antennomere VI (in O. flavicauda ), or from scape to antennomere V (in O. melzeri , O. soror ). Antennae (Group B) entirely micropubescent (in O. apicalis , O. rufifrons ), or only towards apex (in O. quinquecallosus , O. septemtuberculatus ); densely clothed with thick setae from scape to antennomere IV (in female O. rufifrons ), or scape to V (in male O. rufifrons and female O. quinquecal- losus), or scape to VI (in males of O. apicalis , O. quinquecallosus ); or scape to VI densely clothed with narrower setae (in both sexes of O. septemtuberculatus ). Legs generally clothed with semierect setae, densest on femoral clave and tibia, or rather weakly setose (in O. apicalis , O. septemtuberculatus ).

Puncturation. Generally dense, small and alveolate, on forebody rarely beveled. Dorsal head almost impunctate on labrum and clypeus (in both sexes of Group A, O. apicalis , O. rufifrons ), or sparsely punctured (in O. quinquecallosus , O. septemtuberculatus ); adjacent area of frons and vertex, densely punctured, these deep, small and simple or alveolate on frons, on vertex contiguous and alveolate (in Group A males, O. apicalis , O. quinquecallosus , O. sanguinolentus and both sexes of O. rufifrons , O. septemtuberculatus ); in other females, frons and vertex with sparser, shallower, larger, alveolate punctures (in O. flavicauda , O. quinquecallosus ); mentum-submentum with simple punctures scattered between arced carinae (in nearly all species), but lacking carinae (in female O. flavicauda ). Pronotum with punctures of mixed size, shallow, alveolate or subalveolate (in most species); usually denser in males, in females usually sparser. In species with well-developed tubercles, punctures generally scattered, others in clusters (in O. flavicauda , O. soror , O. septemtuberculatus ), in species with more even surface, punctures much denser on disc, towards sides scattered (in O. melzeri ), or almost uniformly and densely punctured (in males of O. apicalis , O. quinquecallosus , O. sanguinolentus and both sexes of O. rufifrons ). Mesosternum entirely micropunctate (in O. melzeri ), or at sides impunctate or nearly so (in all other species); mesepimera micropunctate. Punctures at base of elytra alveolate, confluent and concentrated on humeri. Lateral and sutural elytral margins densely punctured, sometimes as far as apex, puncturations small, alveolate, contiguous and moderately deep. In Group A, basal half of elytra with double row of larger punctures along lateral margins, apical half with single row of very small punctures. In Group B, basal half of elytra with various rows of punctures along lateral margins, apical half with single row of very small punctures (in O. apicalis ), or double or triple row of larger punctures (in the remaining species); translucent panels with weak, sparse, shallow, simple punctures (in most species of this group), or densely punctured (in O. sanguinolentus ). Abdomen incrementally punctate from base to apex, punctures small, shallow, simple or beveled. In male, urosternites I–III almost impunctate towards midline, sparsely punctured towards sides, IV entirely sparsely punctate, V densely punctate in some areas (in most species), or generally more punctate with V entirely densely punctate (in O. melzeri , O. quinquecallosus ). In female, abdomen entirely sparsely punctate.

Color. Generally black, or black and rufous/chestnut, or mostly rufous; lacking bright colors, but characteristic yellow tips of elytra serve as a primary diagnostic for most species. Males are dichromatic in O. flavicauda .

Host plant. Some species of Odontocrotron have been observed as adults on an unidentified species of Croton Linnaeus (Euphorbiaceae) . This plant is referred to elsewhere (e.g., Clarke 2011) as Croton sp. A ; its common name is “Tinajero”. Known host records are provided in Table 1 .

Diagnosis. Separation of Odontocroton from Rhinobatesia is provided under the description of the latter.

Odontocroton may be separated from most species of Odontocera by the following character combination: Group A and B species usually with contrasting yellow spot on apex of elytra. Group A species with lateral profile of prothorax interrupted by overhanging carinae. Group B species with characteristic tegmen of aedeagus, possessing divergent, broader lobes in contrast to the parallel, narrower lobes seen in most Odontocera .

Species of Odontocroton may be separated from Odontocera fasciata by the following character combination: in Odontocroton antennae longer, in male apex reaching base of urosternite II–III, in female reaching apical third of I to apex of III (in Odontocera fasciata antennae shorter, in male apex reaching middle of urosternite I, in female not passing metacoxae); in Odontocroton antennomere III 1.2–1.6× longer than scape, and not longer, or not much longer than V and VI (in Odontocera fasciata antennomere III 0.95–1.1× length of scape, and distinctly longer than V and VI); in Odontocroton prothorax not globular, surface most often with callosities and/or sulcation (in Odontocera fasciata prothorax globular, surface almost without callosities or sulcation); in Odontocroton prosternal process from approximately 5–16× narrower than width of coxal cavity (in Odontocera fasciata process at least 30× narrower than width of coxal cavites); in Odontocroton elytra narrow for apical half, except in O. flavirostris (in Odontocera fasciata elytra not much narrower for apical half); in Odontocroton abdomen cylindrical in males, fusiform in females, abdominal process with marked sexual differences of structure and slope (in males approximately 45–80°, to vertical in O. soror ), in females wider, blunter and about half as steep (in Odontocera fasciata abdomen vespiform in both sexes, structure and slope (approximately 80°) of abdominal process the same in both sexes); in male Odontocroton mesofemur 1.3–1.4× longer than tibia (in male O. fasciata mesofemur 1.1× longer than tibia); in Odontocroton metafemur moderately slender, clave not more than 1.4× longer than peduncle (in Odontocera fasciata metafemur robust, clave more than 2× length of peduncle); in Odontocroton metatarsomere I not less than 1.4× longer than length of II+III (in O. fasciata metatarsomere I not more than 1.2× longer than length of II+III); in Odontocroton general color orange/rufous and black, pronotum without transverse bands of different color (in Odontocera fasciata general color yellow, pronotum with transverse bands of different color); in Odontocroton lateral lobes of tegmen either divergent and narrowed to apex, or parallel, straight and hardly longer than Y-piece (in Odontocera fasciata lateral lobes of tegmen parallel, straight and much longer than Y-piece).

Species included in this genus are: Group A: Odontocroton flavicauda ( Bates, 1873) , O. flavirostris (Melzer, 1930) , O. melzeri ( Fisher, 1952) and O. soror ( Gounelle, 1911) . Group B (i): Odontocroton apicalis ( Klug, 1825) , O. rufifrons ( Fisher, 1937) , O. sanguinolenta ( Bates, 1873) and provisionally O. monnei ( Zajciw, 1968) . Group B (ii): Odontocroton quinquecallosa ( Zajciw, 1963) and O. septemtuberculata ( Zajciw, 1963) .

Etymology. The name of this genus is derived from Odontocera and Croton , the genus of the known host flower for some Group A and Group B species included in the new genus. The genus is masculine.

Figures

Figures 1–6. Odontocroton flavicauda (Bates, 1873). 1) Holotype, male. 2) Male, pale form. 3) Male, dark form, dorsal aspect. 4) Male, dark form, ventral aspect. 5) Female, dorsal aspect. 6) Female, ventral aspect.

Figures 7–12. Species of Odontocroton (Group A). 7-8. Odontocroton flavirostris (Melzer, 1930), holotype female. 7) Dorsal aspect. 8) Ventral aspect. 9-10. Odontocroton melzeri (Fisher, 1952). 9) Holotype male. 10) Topotype, male. 11-12. Odontocera soror (Gounelle, 1911), syntype male. 11) Dorsal aspect. 12) Ventral aspect.

Figures 13–18. Species of Odontocroton (Group A and Group B (i)). 13) Odontocroton soror (Fisher, 1952), syntype female. 14) Odontocroton monnei (Zajciw, 1968), holotype male. 15–18. Odontocroton apicalis (Klug, 1825). 15) Holotype female. 16) Detail, holotype. 17) Dorsal aspect, male. 18) Ventral aspect, male.

Figures 19–24. Species of Odontocroton (Group B (i)). 19) Odontocroton sanguinolentus (Bates, 1873), syntype female. 20–24. Odontocroton rufifrons (Fisher, 1937). 20) Holotype female. 21) Male, dorsal aspect. 22) Male, ventral aspect. 23) Female, dorsal aspect. 24) Female, ventral aspect.

Figures 25–30. Species of Odontocroton (Group B (ii)). 25) Odontocroton quinquecallosus (Zajciw, 1963), holotype, male. 26) Odontocroton septemtuberculatus (Zajciw, 1963), holotype male. 27–30. Odontocroton quinquecallosus. 27) Male, dorsal aspect. 28) Male, ventral aspect. 29) Female, dorsal aspect. 30) Female, ventral aspect.

Figures 31–36. Species of Odontocroton (Group B (ii)) and Rhinobatesia n. gen. 31–34. Odontocroton septemtuberculatus (Zajciw, 1963). 31) Male, dorsal aspect. 32) Male, ventral aspect. 33) Female, dorsal aspect. 34) Female, ventral aspect. 35–36. Rhinobatesia rugicollis (Bates, 1880). 35) Holotype male. 36) Female, holotype of junior synonym Odontocera nevermanni Fisher, 1930.

Figures 43–51. Genitalia (tegmen) of the genera Odontocera, Odontocroton and Rhinobatesia. 43) Odontocera fasciata. 44–46. Odontocroton (Group A). 44) Odontocroton flavicauda. 45) Odontocroton melzeri. 46) Odontocroton soror. 47–50. Odontocroton (Group B). 47) Odontocroton apicalis. 48) Odontocroton rufifrons. 49) Odontocroton quinquecallosus. 50) Odontocroton septemtuberculatus. 51) Rhinobatesia rugicollis.