Koponenius unicornis, Golovatch, Sergei I. & Vandenspiegel, Didier, 2014

Golovatch, Sergei I. & Vandenspiegel, Didier, 2014, Koponenius gen. nov., a new genus of the millipede family Haplodesmidae from the Himalayas of India and Nepal (Diplopoda: Polydesmida), Zootaxa 3894 (1), pp. 141-151 : 144-147

publication ID	https://doi.org/ 10.11646/zootaxa.3894.1.11
publication LSID	lsid:zoobank.org:pub:78A995DA-7D11-48CE-A547-C7653E505720
DOI	https://doi.org/10.5281/zenodo.6126354
persistent identifier	https://treatment.plazi.org/id/4C790A59-BF16-4562-BA43-8AEF3EE2430D
taxon LSID	lsid:zoobank.org:act:4C790A59-BF16-4562-BA43-8AEF3EE2430D
treatment provided by	Plazi
scientific name	Koponenius unicornis
status	sp. nov.

Treatment

Koponenius unicornis View in CoL sp. nov.

Figs 1–4 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4

Material examined: Holotype male ( ZMUM), India, West Bengal, Darjeeling District, Kalimpong (Gandhi Ashram), 27,06ºN, 88,45ºE, ca 880 m a.s.l., 5–15 November 2013, leg. K. Tomkovich. Paratypes: 7 males, 8 females ( ZMUM), 1 male, 1 female ( SMF), 2 males (SEM), same data, together with holotype.

Name: To emphasize the conspicuous central projection at the fore margin of the collum in both sexes; adjective.

Diagnosis: Differs clearly from the only other known congener, K. biramus sp. nov., in the presence of a highly conspicuous central process at the fore margin of the collum in both sexes, coupled with a uniramous gonopod telopodite.

Description: Length in both sexes ca 12–14 mm, width ca 1.3–1.5 and 1.8–2.0 mm on midbody pro- and metazonae, respectively. Holotype ca 13 mm long, 1.3 and 1.8 mm wide on midbody pro- and metazonae, respectively. Coloration in alcohol mostly brown to dark grey-brown, including vertigial region of head, as well as most of pleurosternal and midsternal regions, but pattern evident due to contrasting pale, whitish to light greyish clypeolabral and occipital regions of head, antennae, prozonae, porosteles, legs, epi-, hypo- and paraprocts ( Fig. 1 View FIGURE 1 ).

Body with 19 segments in both sexes, subcylindrical, not capable of volvation. Collum and following metaterga clothed with a dense, dull, microvillose cerategument crust ( Fig. 2 View FIGURE 2 M–P); vertigial region down to a well-expressed border with clypeolabral region clearly granular ( Fig. 2 View FIGURE 2 D). Clypeolabral region densely setose ( Fig. 2 View FIGURE 2 D, Q). Antennae short and clavate, in situ each placed inside a deep, transverse, nearly C-shaped groove ( Fig. 2 View FIGURE 2 D); in length, antennomere 6> 2> 1> 3=4=5; both antennomeres 5 and, especially, 6 with a tight dorso-apical group of bacilliform sensilla ( Fig. 2 View FIGURE 2 A, D, L). Ventral surface of cardo and gnathochilarium largely finely squamate ( Fig. 2 View FIGURE 2 L).

In width, head <collum = segment 3 = 4 <2 = 5=16, thereafter body gradually tapering towards telson ( Fig. 2 View FIGURE 2 H–J). Collum at fore margin dentate, with a highly conspicuous, central, tuberculate protuberance overhanging the head and concealing it from above ( Fig. 2 View FIGURE 2 A, D, H). Paraterga rather well developed, strongly declined, mostly wing-shaped, set low (at about lower ¼ of midbody height), starting from collum, laterally vaguely lobulate ( Figs 1 View FIGURE 1 & 2 View FIGURE 2 A–J, M, P). Paraterga 2 clearly enlarged, subtending the head on both sides. Dorsum very convex ( Fig. 2 View FIGURE 2 M). Postcollum metaterga usually with 4, rarely 5, regular, transverse rows of rather flat, sometimes rather clearly obliterated, setigerous, isostictic (= regular in axial direction), mostly roundish bosses or tubercles ( Figs 1 View FIGURE 1 & 2 View FIGURE 2 A–C, H–J); caudal row of particularly elongate bosses forming a faint waist ( Fig. 2 View FIGURE 2 I), a series of lobulations and ending up in a distinct crenulate limbus ( Fig. 2 View FIGURE 2 I, N); fore row likewise forming a row of similar lobulations ( Figs 1 View FIGURE 1 & 2 View FIGURE 2 A–C, H–J, N, P). Caudolateral corner of postcollum paraterga mostly well rounded, nearly pointed only in segments 17 and 18 ( Fig. 2 View FIGURE 2 J, K). Ozopores borne on conspicuous dorsolateral porosteles, these lying close to caudolateral corners of paraterga; pore formula 5, 7–18 ( Figs 1 View FIGURE 1 & 2 View FIGURE 2 A–C, I, M, O, P). Tergal setae filiform, rather long ( Fig. 2 View FIGURE 2 O, P). Epiproct strongly flattened dorsoventrally, lobulated laterally, subtruncate caudally, but tip ventral, invisible from above ( Fig. 2 View FIGURE 2 C, G, J, K). Hypoproct subtrapeziform ( Fig. 2 View FIGURE 2 G).

Sterna narrow, but evident, mostly slightly elevated due to small subtriangular lobules observed between both coxae ( Fig. 2 View FIGURE 2 D–F). Spiracles seem to be absent. Legs mostly tightly appressed to venter, densely setose, short and stout, about as long as body height and subequally incrassate in both sexes ( Figs 1 View FIGURE 1 & 2 View FIGURE 2 B–F, M, Q, R), but only male coxae and, to a lesser degree, prefemora micropapillate ( Fig. 4 View FIGURE 4 A).

Gonopods ( Figs 3 View FIGURE 3 & 4 View FIGURE 4 B) with large, subcylindrical, mediobasally fused coxae carrying a few long setae ventrally and a small, but evident, rounded, lateral lobe apically; cannulae long and slender. Telopodite strongly elongated, considerably longer than coxae, slender; prefemoral (= densely setose) portion suberect, about half as long as entire telopodite, set off from a subfalcate, rather gradually attenuating and narrowly truncate acropodite (= solenomere, sl) by a strong sulcus (s) on mesal face and by a ventral projection (p); seminal groove rapidly shifted laterad due to a twisted prefemoral portion, thereafter running only on lateral side all over sl extent. Neither an accessory seminal chamber nor a hairy pulvillus.