Comibaena birectilinea, Han & Galsworthy & Xue, 2012

Han, Hongxiang, Galsworthy, Anthony C. & Xue, Dayong, 2012, The Comibaenini of China (Geometridae: Geometrinae), with a review of the tribe, Zoological Journal of the Linnean Society 165 (4), pp. 723-772 : 754-756

publication ID

https://doi.org/ 10.1111/j.1096-3642.2012.00826.x

persistent identifier

https://treatment.plazi.org/id/0389DF41-FF89-A922-FC09-AA96FBDBFC7B

treatment provided by

Marcus

scientific name

Comibaena birectilinea
status

sp. nov.

COMIBAENA BIRECTILINEA SP. NOV.

( FIGS 36 View Figures 27–54 , 90 View Figures 79–90 , 140 View Figures 132–155 , 197 View Figures 188–199 )

Description: Antenna in male with basal threequarters bipectinate, outer rami longer than inner rami. Frons white, a small green patch present in the holotype. Labial palpus with upper and lateral surfaces brown, ventral surface white. Vertex green. Dorsal side of thorax and abdomen pale green, intersegment white on abdomen. Hind tibia with two pairs of spurs in both sexes, not dilated, without hair-pencil, terminal extension absent.

Forewing length: ♀ ♂ 16–17 mm. Forewing green, with white antemedial and postmedial lines, both straight, the antemedial inclined outwards, the postmedial inwards. Hindwing pale green, anterior part whitish, with white submarginal line parallel to outer margin; discal spot dark green. Both wings with terminal line and fringes concolorous with wing surface. Underside: forewing same as upperside; hindwing with outwardly curved postmedial line. Frenulum well developed.

Male genitalia. Uncus totally bifid, tapering, bent outwards. Socii longer than uncus, tapering but blunt. Valva simple, apex blunt; costa protruding at middle. Vinculum short, concave at middle, with branches short and broad. Aedeagus slender.

Female genitalia. Sterigma irregularly shaped. Antrum rounded, with posterior margin deeply concave, depth about half length of antrum. Ductus bursae quite short. Corpus bursae weak (incomplete in specimen examined).

Diagnosis: Comibaena birectilinea is similar to Comibaena cenocraspis and Comibaena hypolampes in the male genitalia and wing pattern. However, the antemedial line is distinctly inclined outwards, and close to the postmedial line on the inner margin. In the male genitalia, the costa is protruding and angled, and the lateral protrusions of the vinculum are shorter and broader. The female genitalia are similar to those of Comibaena cenocraspis , but the concavity of the antrum is shallower. The wing pattern of Comibaena birectilinea resembles the members of Thetidia , but the frenulum is present in this species, whereas it is usually absent in Thetidia .

Holotype, ♂, China: Tibet: Jomda , 3400 m, 31.vii.1976, coll. Zhang Xuezhong, IZCAS slide no. Geom-00124 ( IZCAS) . Paratypes ( IZCAS): China: 2♀, same locality as holotype, 26–27.vii.1976, coll. Han Yinheng, Zhang Xuezhong, IZCAS slide no. Geom-00125 ; 1♀, Sichuan: Batang, Yidun , 3350 m, 16.viii.1982, coll. Zhang Xuezhong ; 1♂, Batang (Tibet), Untere Urwaldzone , c. 3800 m, 5.viii.1936, H. Höne ( ZFMK) ; same locality and collector , 3♂, 7.viii.1936, 1♂, 8.viii.1936; 1♂, 14.viii.1936, 1♂, 15.viii.1936 ( ZFMK) ; 1♂, Batang (Tibet), Im Tal des Yangtse , c. 2800 m, 18.ix.1936, H. Höne ( ZFMK) ; 1♂, Tibet, Yargong Zambala, P. Soulié, 1903, ex. coll. Oberthür ( ZFMK) .

Distribution: China (Sichuan, Tibet). Etymology: A combination of the Latin prefix bi - and rectilinea, referring to the linear antemedial and postmedial lines on the forewing.

Species not occurring in China

Square brackets indicate nature of material examined.

Comibaena albimarginata ( Warren, 1893) View in CoL [specimen(s); male and female genitalia photographs, Holloway (1996: figs 248, 265)]

Uliocnemis albimarginata Warren, 1893: 355 View in CoL . ( India: Sikkim)

Comibaena amoenaria ( Oberthür, 1880) View in CoL [specimen(s)]

Phorodesma amoenaria Oberthür, 1880: 48 View in CoL , pl. 9, figure 4. ( Russia: Askold Island)

Comibaena bajularia bajularia ( Denis & Schiffermüller, 1775) View in CoL [male and female genitalia photographs, Hausmann, (2001: fig. 19)]

Geometra bajularia Denis & Schiffermüller, 1775: 97 View in CoL . ( Austria)

Phalaena ditaria Fabricius, 1776: 286 View in CoL . ( Germany)

Phalaena glauca Fourcroy, 1785: 267 View in CoL . ( France)

Phalaena pustulata Hufnagel, 1767: 520 View in CoL . ( Germany) [Junior homonym of pustulata Müller, 1764 View in CoL , Arctiidae .]

Comibaena bajularia tikhonovi Hausmann, 2000

Comibaena bajularia tikhonovi Hausmann, 2000: 302 , fig. d. ( Russia)

Comibaena barnsi barnsi Prout, 1930 View in CoL [specimen(s)]

Comibaena barnsi Prout, 1930: 14 View in CoL . ( Zaïre)

Comibaena barnsi equatorialis Debauche, 1941 View in CoL

Comibaena barnsi equatorialis Debauche, 1941: 302 View in CoL , pl. 10, figure 3; pl. 11, figure 6. ( Zaïre)

Comibaena biviaria Hampson, 1909 View in CoL [specimen(s)]

Comibaena biviaria Hampson, 1909: 126 View in CoL , pl. 4, figure 60. (Ruwenzori [south-west Uganda /north-east Zaïre])

Comibaena castaneata ( Warren, 1906) View in CoL [specimen(s)]

Comostolodes castaneata Warren, 1906: 87 View in CoL . ( Papua New Guinea)

Probolosceles agathia Warren, 1912: 80 View in CoL . ( Papua New Guinea) [specimen(s)]

Comibaena chalybeata Moore, 1868 View in CoL [specimen(s); male genitalia]

Comibaena chalybeata Moore, 1868: 639 View in CoL . ( India)

Comibaena cheramota ( Meyrick, 1886) View in CoL [specimen(s)]

Iodis cheramota Meyrick, 1886: 203 . ( Fiji)

Comibaena connata ( Warren, 1898) View in CoL [specimen(s)]

Probolosceles connata Warren, 1898: 15 View in CoL . ( Australia)

Comibaena coryphata ( Felder & Rogenhofer, 1875) View in CoL [specimen(s)]

Racheospila coryphata Felder & Rogenhofer, 1875 View in CoL : pl. 127, figure 10. ( South Africa: Knysna)

Comibaena diluta ( Warren, 1895) View in CoL [specimen(s)]

Colutoceras diluta Warren, 1895: 88 View in CoL . ( Japan)

Euchloris obsoletaria Leech, 1897: 238 View in CoL . ( Japan) [specimen(s); female genitalia]

Comibaena erythrospila Yazaki, 1998 View in CoL

Comibaena erythrospila Yazaki, 1998: 4 View in CoL , pl. 129, figure 7; text-figure 840. ( Nepal)

Comibaena esmeralda ( Warren, 1898) View in CoL [specimen(s)]

Probolosceles View in CoL ? esmeralda Warren, 1898: 15 View in CoL . ( Nigeria)

Comibaena flavitaenia ( Warren, 1898) View in CoL [specimen(s)]

Oenospila flavitaenia Warren, 1898: 13 View in CoL . ( Nigeria)

Comibaena hemictenes Prout, 1917 View in CoL [specimen(s)]

Comibaena hemictenes Prout, 1917: 114 View in CoL , pl. 7, figure 21. ( Ivory Coast)

Comibaena ingrata ( Wileman, 1911) View in CoL [specimen(s)]

Thalera ingrata Wileman, 1911: 343 View in CoL . ( Japan)

Comibaena insulana Inoue, 1986 View in CoL [specimen(s)]

Comibaena insulana Inoue, 1986: 53 View in CoL , figures 8a, b, 9a, b. ( Japan)

Comibaena integranota Hampson, 1893 View in CoL [specimen(s); male genitalia]

Comiboena integranota Hampson, 1893: 37 , 146, pl. 170, figure 13. ( Sri Lanka)

Comibaena leucochloraria ( Mabille, 1880) View in CoL [specimen(s)]

Phorodesma leucochloraria Mabille, 1880 View in CoL : xxii. ( Madagascar)

Comibaena leucospilata ( Walker, 1863) View in CoL [specimen(s)]

Geometra leucospilata Walker, 1863: 1554 View in CoL . ( South Africa)

Comibaena levequei Leraut, 2009 [specimens(s)]

Chlorochaeta longipennis Leraut, 2009: 46 View in CoL . ( Algeria)

Comibaena longipennis ( Warren, 1904) View in CoL [specimen(s)]

Chlorochaeta longipennis Warren, 1904: 464 View in CoL . ( Niger)

Comibaena mariae mariae ( Lucas, 1888) View in CoL [male genitalia]

Iodis mariae mariae Lucas, 1888: 1266 . ( Australia: Queensland)

Probolosceles albipunctata Warren, 1898: 15 View in CoL . ( Australia: Queensland) [specimen(s)]

Comibaena mariae syndyas Prout, 1916b View in CoL [specimen(s)]

Comibaena mariae syndyas Prout, 1916b: 205 View in CoL . (New Guinea)

Comibaena meyricki Prout, 1933 View in CoL [specimen(s); male genitalia]

Comibaena quadrinotata meyricki Prout, 1933: 92 View in CoL . (Talaut)

Comibaena multigruma Prout, 1920 View in CoL [specimen(s)]

Comibaena multigruma Prout, 1920: 299 View in CoL . ( Papua New Guinea)

Comibaena nepalensis Yazaki, 2000 [specimen and male genitalia photograph, Yazaki, (2000: pl. 162: 1; fig. 1248)]

Comibaena nepalensis Yazaki, 2000: 9 .

Comibaena pseudoneriaria Wehrli, 1926 View in CoL [male and female genitalia photograph, Hausmann (2001: fig. 20)]

Comibaena pseudoneriaria Wehrli, 1926: 163 View in CoL . ( Spain)

Comibaena punctaria ( Swinhoe, 1904) View in CoL [specimen(s)]

Probolosceles punctaria Swinhoe, 1904: 550 View in CoL . ( Madagascar)

Comibaena rhodolopha Prout, 1915 View in CoL [specimen(s)]

Comibaena rhodolopha Prout, 1915: 313 View in CoL . ( Zimbabwe)

Comibaena rhodonia Prout, 1913b View in CoL [specimen(s)]

Comibaena rhodonia Prout, 1913b: 416 View in CoL . (Irian Jaya)

Comibaena rufitornus Prout, 1916c View in CoL [specimen(s)]

Comibaena rufitornus Prout, 1916c: 275 View in CoL . ( Kenya)

Comibaena serrulata Fletcher, 1963 View in CoL [specimen(s); male genitalia]

Comibaena serrulata Fletcher, 1963: 116 View in CoL , pl. 8, figures 1–5 ( Iran)

Comibaena viridifimbria ( Warren, 1906) View in CoL [specimen(s); male genitalia]

Comostolodes viridifimbria Warren, 1906: 87 View in CoL . ( Papua New Guinea)

IZCAS

Institute of Zoology, Chinese Academy of Sciences

ZFMK

Zoologisches Forschungsmuseum Alexander Koenig

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Lepidoptera

Family

Geometridae

Genus

Comibaena

Loc

Comibaena birectilinea

Han, Hongxiang, Galsworthy, Anthony C. & Xue, Dayong 2012
2012
Loc

Chlorochaeta longipennis

Leraut P 2009: 46
2009
Loc

Comibaena bajularia tikhonovi

Hausmann A 2000: 302
2000
Loc

Comibaena nepalensis

Yazaki K 2000: 9
2000
Loc

Comibaena erythrospila

Yazaki K 1998: 4
1998
Loc

Comibaena insulana

Inoue H 1986: 53
1986
Loc

Comibaena serrulata

Fletcher DS 1963: 116
1963
Loc

Comibaena quadrinotata meyricki

Prout LB 1933: 92
1933
Loc

Comibaena barnsi

Prout LB 1930: 14
1930
Loc

Comibaena multigruma

Prout LB 1920: 299
1920
Loc

Comibaena hemictenes

Prout LB 1917: 114
1917
Loc

Comibaena mariae syndyas

Prout LB 1916: 205
1916
Loc

Comibaena rufitornus

Prout LB 1916: 275
1916
Loc

Comibaena rhodolopha

Prout LB 1915: 313
1915
Loc

Comibaena rhodonia

Prout LB 1913: 416
1913
Loc

Probolosceles agathia

Warren W 1912: 80
1912
Loc

Thalera ingrata

Wileman AE 1911: 343
1911
Loc

Comibaena biviaria

Hampson GF 1909: 126
1909
Loc

Comostolodes castaneata

Warren W 1906: 87
1906
Loc

Comostolodes viridifimbria

Warren W 1906: 87
1906
Loc

Chlorochaeta longipennis

Warren W 1904: 464
1904
Loc

Probolosceles punctaria

Swinhoe C 1904: 550
1904
Loc

Probolosceles connata

Warren W 1898: 15
1898
Loc

Probolosceles

Warren W 1898: 15
1898
Loc

Oenospila flavitaenia

Warren W 1898: 13
1898
Loc

Probolosceles albipunctata

Warren W 1898: 15
1898
Loc

Euchloris obsoletaria

Leech JH 1897: 238
1897
Loc

Colutoceras diluta

Warren W 1895: 88
1895
Loc

Uliocnemis albimarginata

Warren W 1893: 355
1893
Loc

Comiboena integranota

Hampson GF 1893: 37
1893
Loc

Iodis mariae

Lucas TP 1888: 1266
1888
Loc

Iodis cheramota

Meyrick E 1886: 203
1886
Loc

Phorodesma amoenaria Oberthür, 1880: 48

Oberthur C 1880: 48
1880
Loc

Comibaena chalybeata

Moore F 1868: 639
1868
Loc

Geometra leucospilata

Walker F 1863: 1554
1863
Loc

Phalaena glauca

Fourcroy AFD 1785: 267
1785
Loc

Phalaena ditaria

Fabricius JC 1776: 286
1776
Loc

Geometra bajularia Denis & Schiffermüller, 1775: 97

Denis M & Schiffermuller I 1775: 97
1775
Loc

Phalaena pustulata

Hufnagel JS 1767: 520
1767
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