Notoxus impavidus, Kejval, 2011

Kejval, Zbyněk, 2011, Taxonomic revision of the Oriental species of Notoxus (Coleoptera: Anthicidae), Acta Entomologica Musei Nationalis Pragae 51 (2), pp. 627-673 : 649-650

Notoxus impavidus sp. nov.

( Figs. 22 , 53 , 73 )

Type locality. Laos, Vientiane Province, Phou Khao Khouay National Park, Tad Leuk, 18°23′N, 103°04′E, 150–200 m a.s.l.

Type material. HOLOTYPE: J, ‘ LAOS central, Saisombun zone PHOU KHAO KHOUAY Nat. Park TAD LEUK 18°23′N, 103°04′E 15.–21.V.2001, alt. 150–200 m E. Jendek & O. Šauša leg.’ ( NMPC) . PARATYPES: 16 JJ 3 ♀♀, same data as holotype ( ZKDC, 1 spec. each in BMNH, DCDC, MHNG, MNHN, NHMB, ZMHB) ; 1 J, ‘ LAOS centr., Khammouan pr., BAN KHOUN NGEUN vill. env., 17.v.–6.vi.2007, ~ 300 m, M. Štrba leg.’ ( ZKDC) ; 8 JJ, ‘C-LAOS: Prov. Viangchan Phou Khao Khouay NP Nam Leuk env. Tad Leuk Waterf. // 1.- 8.6.1996, 200m at light leg. Schillhammer (15a) // Notoxus inbasalis Pic det. G. Uhmann 1998’ ( NHMW, ZSMC) .

Description. Male (holotype). Body length 3.5 mm. Head and pronotum reddish brown. Elytra brown, with reddish tinge basally, with indication of two vaguely outlined, transverse, pale reddish bands. Legs and antennae reddish.

Body only moderately glossy, rather finely, densely and evenly punctured; pronotum with some scattered coarser punctures; punctures of elytra rather fine but distinct ( Fig. 73 ), slightly coarser than those on pronotum, simple, with smooth margins. Body setation mostly subdecumbent; pronotum with numerous conspicuously long, erect setae, especially anterolaterally; elytra with numerous, moderately longer suberect to erect setae. Antebasal setose band of pronotum conspicuous, widely interrupted medially.

Head with rather large eyes. Antennae long, only moderately enlarged in terminal third. Pronotum 1.1 times as wide as head across eyes, globose in dorsal view. Pronotal horn moderately long, slender, largely subparallel, with continuously fringed apical margins and 3 lateral lobules on each side, becoming distinctly smaller towards horn base; horn crest only moderately raised, evenly lowering towards horn apex, its lateral margins feebly marked by series of somewhat coarser, distinctly separate lobules / rugules; dorsal surface rather evenly and densely covered with distinct rounded rugules. Elytra 1.9 times as long as wide; humeri distinctly protruding; omoplates and postbasal moderately indicated; elytral apices distinctly modified, subtruncate, with rounded gland opening near distinctly produced lateral angle. Legs simple. Abdominal sternum VII moderately impressed postero-medially, its posterior margin somewhat unevenly rounded. Tergum VIII forming simple, evenly arched sclerite. Aedeagus as figured ( Fig. 22 ).

Female. Externally differing from male by simply rounded elytral apices.

Variation. Body length (J ♀) 3.2–3.6 mm. Moderately variable in width of pronotal horn; its lateral margins with 2–4 distinct lobules on each side.

Differential diagnosis. Notoxus impavidus sp. nov. shares long bifurcate dorsal plate of the tegmen of the aedeagus with N. ales and N. garuda sp. nov., but differs by somewhat paler body colouration (dark markings of elytra less distinct, vaguely outlined), rather fine and dense punctation of the elytra ( Fig. 73 ), and by the morphology of the aedeagus.

Etymology. From the Latin, impavidus (fearless, intrepid); named in reference to the location of the type locality in the former Saisomboun special zone, administered by military forces.

Distribution. Laos.

Figures

Figs. 22–26. 22 – Notoxus impavidus sp. nov., aedeagus. 23–26 – N. katthapa sp. nov.: 23 – sternum VII; 24 – tergum VIII; 25 – aedeagus in dorsal view; 26 – the same, lateral view. Scale (0.5 mm): A – Figs. 22, 23; B – Figs. 25, 26; C – Fig. 24.

Figs. 52–60. 52–58 – pronotal horn in dorsal view: 52 – Notoxus brunneorufus Bonadona, 1989 (Pachmarhi); 53 – N. impavidus sp. nov.; 54 – N. variabilis Krekich-Strassoldo, 1913 (Mulshi); 55 – N. noctivagus Krekich- Strassoldo, 1913 (Punjab); 56 – N. sinensis Pic, 1907 (Xishan); 57 – N. pachodemba sp. nov.; 58 – N. safraneki sp. nov.; 59 – N. inbasaliformis sp. nov., head in dorsal view; 60 – N. psammophilus Telnov, 2007 (Cam Ranh Bay), the same.

Figs. 61–75. 61–64 – elytra: 61–62 – Notoxus noctivagus Krekich-Strassoldo, 1916 (61 – Karadj, 62 – Punjab); 63–64 – N. variabilis Krekich-Strassoldo, 1913 (63 – syntype, 64 – Mulshi). 65–67 – elytral apices: 65 – N. peguensis Pic, 1914 (Myitkyina); 66 – N. psammophilus Telnov, 2007 (Cam Ranh Bay); 67 – N. variabilis (Mulshi). 68 – N. ales Telnov, 2007 (Ban Nape), laterobasal fovea of sternum III; 69–70 – N. ravana sp. nov.: 69 – laterobasal fovea of sternum III; 70 – apex of elytron. 71–72 – N. ales (Ban Nape): 71 – male sternum VII; 72 – punctation of elytra (basal half). 73 – N. impavidus sp. nov., the same. 74 – N. donckieri Pic, 1908 (Phuyen Binh), male protibia. 75 – N. hilaris sp. nov., the same.