Laccophilus javanicus, Regimbart, 1899

Hájek, Jiří & Brancucci, Michel, 2015, A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae), Raffles Bulletin of Zoology 63, pp. 309-326 : 310-311

publication ID

https://doi.org/ 10.5281/zenodo.5385198

publication LSID

lsid:zoobank.org:pub:AC7F1491-CD89-4F10-9D61-55354B55E368

DOI

https://doi.org/10.5281/zenodo.6512177

persistent identifier

https://treatment.plazi.org/id/038A87C4-FFA4-FF9E-F05B-9104FCD0BDBA

treatment provided by

Valdenar

scientific name

Laccophilus javanicus
status

 

The Laccophilus javanicus species group was erected by Guignot (1959) for a single African species L. flavopictus Régimbart, 1889 . Although Guignot was apparently aware of Oriental species, he never listed them. Consequently, the group was adopted and modified by Brancucci (1983), who defined this species group as follows: Surface of pronotum and elytra with simple reticulation, composed entirely of large polygonal meshes. Very large species (body length 3.4–5.1 mm), of very broad and depressed shape. Elytra brown to black, with pale bands and other markings. Another characteristic of the group is: Prosternal apophysis relatively short and lanceolate; posterior margin of apical ventrite rounded in both sexes.

Brancucci (1983) assigned nine Oriental species distributed from India, through continental Southeast Asia, east to the Sunda Islands, to the L. javanicus -group. More recently, Balke et al. (1998) described a tenth species from Kalimantan. In the present work, we adopt the concept of the group from Brancucci (1983) and consider it to be exclusively Oriental; Afrotropical Laccophilus flavopictus is removed from the L. javanicus group, as it does not seems to be closely related to Oriental species (O. Biström, pers. comm.).

At present, the Laccophilus javanicus species group comprises 18 species:

Laccophilus aurofasciatus Vazirani, 1972 – India (Kerala, Tamil Nadu)

Laccophilus auropictus Régimbart, 1899 – India (Karnataka, Tamil Nadu), Myanmar (Tenasserim)

Laccophilus ceylonicus Zimmermann, 1919 – India (Maharashtra, Orissa), Sri Lanka

Laccophilus elegans Sharp, 1882 – India (Andaman and Nicobar Islands, Andhra Pradesh, Karnataka, Kerala, Nagaland, Orissa, West Bengal)

Laccophilus girardi Brancucci, 1983 – Malaysia (Sabah)

Laccophilus hendrichi , new species – Myanmar, Thailand, Laos

Laccophilus jaechi , new species – Cambodia, Thailand

Laccophilus javanicus Régimbart, 1899 – Indonesia (Java, Bali, Lombok)

Laccophilus kalimantanensis , new species – Indonesia (Kalimantan)

Laccophilus komareki , new species – China (Guangdong, Guangxi)

Laccophilus mahakamensis Balke, Mazzoldi & Hendrich, 1998 – Thailand, Malaysia ( Sarawak), Indonesia (Kalimantan)

Laccophilus mazzoldii , new species – Laos, Thailand

Laccophilus nusatenggaraensis , new species – Indonesia ( Flores, Sumba, Sumbawa, Timor)

Laccophilus schillhammeri , new species – Myanmar

Laccophilus stastnyi , new species – Malaysia ( Sarawak)

Laccophilus tobaensis Brancucci, 1983 – Indonesia (Sumatra)

Laccophilus tonkinensis Brancucci, 1983 – China (Hainan), Laos, Vietnam

Laccophilus wittmeri Brancucci, 1983 – China (Yunnan), Myanmar, Thailand, Laos, Vietnam

A key to species of the Laccophilus javanicus View in CoL group (males only)

1. Smaller species (body length 3.5–4.5 mm), elongate oval ( Figs. 3–8 View Figs , 10, 12–16 View Figs ) .......................................................................2

– Large species (body length 4.5–5.0 mm), broadly oval ( Figs. 1, 2, 9 View Figs , 11, 17 View Figs ).......................................................................14

2. Small species (body length 3.4–3.8 mm); Borneo ( L. mahakamensis View in CoL also known from southern Thailand).............3

– Medium sized species (body length 3.8–4.5 mm); so far unknown from Borneo ............................................................................6

3. Subbasal testaceous band very broad, covering at least basal third of elytra; or elytra almost uniformly testaceous ( Fig. 5 View Figs ) .................................................................................................4

– Subbasal testaceous band sinuous, narrower, usually well separated from elytral base ( Figs. 10, 14 View Figs )..............................5

4. Apical part of median lobe in lateral view almost straight, narrowing to apex which is bent dorsally ( Fig. 22 View Figs ); body length 3.5–3.7 mm; Sabah .................................................... L. girardi View in CoL

– Apical part of median lobe in lateral view almost regularly curved to apex ( Fig. 26 View Figs ); body length 3.6–3.8 mm; Kalimantan ........ .............................................. L. kalimantanensis View in CoL , new species

5. Apical part of median lobe in lateral view distinctly sinuous with elongate pointed tip ( Fig. 28 View Figs ); body length 3.4–3.7 mm; Sarawak, Kalimantan; southern Thailand..... L. mahakamensis View in CoL

– Apical part of median lobe in lateral view less sinuous with shortly pointed tip ( Fig. 32 View Figs ); body length 3.5–3.6 mm; Sarawak: Bako NP ............................................. L. stastnyi View in CoL , new species

6. Apical part of median lobe in lateral view with distinct bulge on dorsal side (cf. Figs. 20, 21, 30, 34 View Figs ) .................................7

– Apical part of median lobe in lateral view with dorsal side almost straight or very slightly curved (cf. Figs. 23–25, 30, 33 View Figs ) ..........................................................................................10

7. Subbasal testaceous band sinuous, narrow, well separated from elytral base ( Figs. 3 View Figs , 13 View Figs ) .........................................................8

– Subbasal testaceous band broad, more or less connected with elytral base ( Figs. 4 View Figs , 16 View Figs ) .........................................................9

8. Apical part of median lobe narrow with pointed tip ( Fig. 20 View Figs ); body length 3.8–4.1 mm; Sri Lanka, India........ L. ceylonicus View in CoL

– Apical part of median lobe broad with blunt tip ( Fig. 31 View Figs ); body length 3.8–4.3 mm; Myanmar.................................................. .................................................. L. schillhammeri View in CoL , new species

9. Apex of median lobe short, blunt ( Fig. 21 View Figs ); body length 3.8–4.0 mm; Andaman and Nicobar Islands, India.............. L. elegans View in CoL

– Apex of median lobe elongate with pointed tip ( Fig. 34 View Figs ); body length 3.8–4.1 mm; China (Hainan), Laos, Vietnam............... ............................................................................ L. tonkinensis View in CoL

10. Broadly oval species; elytral subbasal testaceous band divided longitudinally into separate spots ( Fig. 15 View Figs ); apical part of median lobe shorter, almost subparallel ( Fig. 33 View Figs ); body length 4.4–4.5 mm; Sumatra ......................................................... L. tobaensis View in CoL

– Elongate oval species; elytral subbasal testaceous band complete; apical part of median lobe longer.........................................11

11. Anterior margin of elytral subbasal testaceous band almost rectilinear ( Fig. 8 View Figs ); body length 4.0– 4.4 mm; Java, Bali, Lombok.................................................................. L. javanicus View in CoL

– Anterior margin of elytral subbasal testaceous band sinuous ( Fig. 6 View Figs ), or band very broad connected with elytral base ( Figs. 7 View Figs , 12 View Figs )......................................................................................12

12. Ground colour of elytra usually pale brown, thus testaceous pattern not well recognisable ( Fig. 12 View Figs ), if elytra dark, then subbasal testaceous band very broad but well separated from elytral base; body length 4.0– 4.5 mm; Flores, Sumba, Sumbawa, Timor ................................ L. nusatenggaraensis View in CoL , new species

– Elytra dark, subbasal testaceous band differently shaped; continental SE Asia ..............................................................13

13. Subbasal testaceous band sinuous, narrow, well separated from elytral base ( Fig. 6 View Figs ); apex of median lobe nearly straight ( Fig. 23 View Figs ); body length 3.8–4.4 mm; Laos, Myanmar, Thailand....... ......................................................... L. hendrichi View in CoL , new species

– Subbasal testaceous band broad, more or less connected with elytral base ( Fig. 7 View Figs ); apex of median lobe bent ( Fig. 24 View Figs ); body length 4.1–4.4 mm; SW Cambodia, SE Thailand.................... ............................................................... L. jaechi View in CoL , new species

14. Reticulation of elytra composed of well impressed comparatively smaller sized meshes; apical part of median lobe in lateral view slightly curved on dorsal side, apex blunt ( Fig. 29 View Figs ); body length 4.6–4.7 mm; C Laos, E Thailand... L. mazzoldii View in CoL , new species

– Reticulation of elytra composed of shallowly impressed very large meshes; apex of median lobe acute ( Figs. 18, 19, 27, 35 View Figs )..........................................................................................15

15. Elytra without mediolateral spot ( Figs. 1, 2 View Figs ); apical part of median lobe in lateral view with distinct bulge on dorsal side ( Figs. 18, 19 View Figs ); S India...........................................................16

– Elytra either with mediolateral spot ( Fig. 17 View Figs ), or ground colour of elytra pale ( Fig. 9 View Figs ); apical part of median lobe in lateral view with dorsal side almost straight ( Figs. 27, 35 View Figs ); continental SE Asia........................................................................................17

16. Subbasal testaceous band broad, straight, not separated from elytral base ( Fig. 1 View Figs ); body length 4.6–4.9 mm; S India.......... ........................................................................ L. aurofasciatus View in CoL

– Subbasal testaceous band narrow, sinuous, well separated from elytral base ( Fig. 2 View Figs ); body length 4.5–4.7 mm; S India.......... ............................................................................. L. auropictus View in CoL

17. Ground colour of elytra pale, testaceous pattern bordered with narrow black rim ( Fig. 9 View Figs ); apical part of median lobe relatively longer, with dorsal side almost straight, and long pointed apex ( Fig. 27 View Figs ); body length 4.5–4.9 mm; China (Guangdong, Guangxi) ........................................... L. komareki View in CoL , new species

– Ground colour of elytra dark, brown-blackish ( Fig. 17 View Figs ); apical part of median lobe relatively shorter, with dorsal side lightly curved, and short and hooked apex ( Fig. 35 View Figs ); body length 4.5–5.1 mm China (Yunnan), Laos, Myanmar, Thailand, Vietnam..... ................................................................................. L. wittmeri View in CoL

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Dytiscidae

Genus

Laccophilus

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