Glyphiulus species

Likhitrakarn, Natdanai, Jeratthitikul, Ekgachai, Jirapatrasilp, Parin & Wesener, Thomas, 2024, Integrated taxonomy of three new species of Glyphiulus Gervais, 1847 (Diplopoda, Spirostreptida, Cambalopsidae) from Laos, Raffles Bulletin of Zoology 72, pp. 42-61 : 57-59

publication ID

https://doi.org/ 10.26107/RBZ-2024-0003

publication LSID

lsid:zoobank.org:pub:DC2582A7-C5DD-4726-9B59-3F2C00E9485E

persistent identifier

https://treatment.plazi.org/id/038A87F0-FFCA-E711-2ED3-F885FCBFFA37

treatment provided by

Felipe

scientific name

Glyphiulus species
status

 

Key to Glyphiulus species presently known to occur in Laos ( Fig. 7 View Fig ), chiefly based on male characters, modified after Likhitrakarn et al. (2017)

1. Male leg 1 very strongly reduced, completely lacking any median structures.....................................................................2

– Male leg 1 either normal or reduced in size, but with a pair of paramedian coxal processes ( Figs. 2B, C View Fig , 4C, D View Fig , 6C, D View Fig ).....3

2. Paraprocts with a row of several strong setae near median marginal ridge; posterior gonopods broadly subquadrate, each half with a plumose apical flagellum (f) .......... G. subbedosae View in CoL

– Paraprocts with a bare marginal ridge devoid of setae; posterior gonopods narrowly subrectangular ........................ G. bedosae View in CoL

3. Carinotaxy formula of midbody rings 2+I/i+3/3+I/i+2..........4

– Carinotaxy formula of midbody rings 2/2+I/i+3/3+I/i+2/2 ( Figs. 1D, E View Fig , 3D, E View Fig , 5D, E View Fig ) ..............................................................5

4. Carinotaxy formula of collum, I–VI+7a+pc+ma+pc+7a+VI–I, texture of both lateral-most crests micropunctate; male leg 1 with two low, paramedian, contiguous cones; each posterior gonopod with a long and bare flagellum............. G. costulifer View in CoL

– Carinotaxy formula of collum, I+2c+III–VI+5c+6a+pc+ma+pc+ 6a+5c+VI–III+2c+I, both lateral-most crests smooth; male leg 1 with medially contiguous, apically diverging cones; each posterior gonopod with a long, distally plumose flagellum .... ....................................................................... G. semicostulifer View in CoL

5. Ommatidia invisible or absent ................................................6

– Ommatidia present ..................................................................7

6. Colouration entirely pallid; adult body length 14–17 mm; carinotaxy formula of collum 1–4+5a+pc+ma; apicomesal processes (d) on the anterior gonopods broad and stout, apicolateral process (ap) being small and rounded; a distally plumose flagellum process (f) present on the posterior gonopods.......................................................... G. percostulifer View in CoL

– Colouration usually brownish; adult body length 18.5–22.5 mm; carinotaxy formula of collum 1+2c+3–4+5c+6a+pc+ma ( Fig. 5A–C View Fig ); apicomesal processes (d) on the anterior gonopods long and slender ( Fig. 6J, K View Fig ), apicolateral process (ap) large, curved, and subdigitiform ( Fig. 6H, I View Fig ); a slim and smooth flagellum process (f) present on the posterior gonopods ( Fig. 6J, K View Fig ) .... .............................................. G. houaphanhensis , new species

7. Carinotaxic formula of collum:1–2+3c+4a+5c+6+7c+8a+pc+ma; male leg 1 telopodites strongly reduced in size, 4-segmented; anterior gonopod with a subsecuriform coxosternum.............. ......................................................................... G. subcostulifer View in CoL

– Carinotaxic formula of collum: 1–5+6a+pc+ma ( Figs. 1A, B View Fig ; 3A, B View Fig ); male leg 1 telopodites nearly normal, 5-segmented ( Figs. 2B, C View Fig , 4C, D View Fig ); anterior gonopod coxosternum with particularly high median outgrowths ( Figs. 2H, I View Fig , 4H, I View Fig )......8

8. Carinotaxic formula of midbody rings: 2+I/i+3/3+I/i+2 ( Fig. 1A, B, D, E, G, H View Fig ); apicomesal processes (d) on the anterior gonopods longer and slender ( Fig. 2H, I View Fig ); without apicolateral process (ap) ( Fig. 2H, I View Fig ); tip of flagellum process (f) on the posterior gonopods pointed ( Fig. 2J, K View Fig )................................... ............................................. G. pseudocostulifer , new species

– Carinotaxic formula of midbody rings: 2/2+I/i+3/3/3+I/i+2/2 ( Fig. 3A, B, D, E, G, H View Fig ); apicomesal processes (d) on the anterior gonopods broader and stout ( Fig. 4H, I View Fig ); with an evident, swollen, rounded apicolateral process (ap) ( Fig. 4H, I View Fig ); tip of flagellum process (f) on the posterior gonopods axe-shaped ( Fig. 4J, K View Fig ) ........................... G. steineri , new species

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