Eurygnathodus costatus Staesche, 1964

Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane & Goudemand, Nicolas, 2022, A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China, Swiss Journal of Palaeontology (19) 141 (1), pp. 1-61 : 52

Eurygnathodus costatus Staesche, 1964

Fig. 14A

*1964 Eurygnathodus costatus n. sp.; Staesche, p. 269, pl. 28, figs. 1–6.

1977 Platyvillosus costatus (Staesche) ; Goel, p. 1098, pl. 2, figs. 15–21.

1981 Platyvillosus costatus (Staesche) ; Wang & Cao, p. 371, pl. 2, figs. 1–4, 28–30, 33.

1981 Platyvillosus paracostatus n. sp.; Wang & Cao, p. 371, pl. 2, figs. 9,10.

1984 Platyvillosus costatus (Staesche) ; Matsuda, p. 128, pl. 6, figs. 6–10.

1988 Platyvillosus costatus (Staesche) ; Koike, p. 65, pl. 1, figs. 1–57, pl. 2, figs. 1–37.

1991 Platyvillosus costatus (Staesche) ; Beyers & Orchard, pl. 5, fig. 10.

2009 Eurygnathodus costatus Staesche ; Igo in Shigeta et al., p. 183, figs. 152.23–152.24.

2010 Eurygnathodus costatus Staesche ; Orchard, p. 145, fig. 5.9–5.10.

2013 Platyvillosus hamadai Koike ; Zhao et al., p.535, figs. 10K, L.

2013 Platyvillosus costatus (Staesche) ; Zhao et al., p.535, figs. 10M, 10N, 10O.

2014 Eurygnathodus costatus Staesche ; Maekawa & Igo in Shigeta et al., p. 220, figs. 161.4–161.6

2015 Eurygnathodus hamadai Staesche ; Maekawa in Maekawa et al., p. 316, fig. 5.1.

2016 Eurygnathodus costatus Staesche ; Chen et al., fig. 11.3, 11.6–11.7.

2018 Eurygnathodus costatus Staesche ; Maekawa in Maekawa et al., pp. 45–50, figs. 25–27.

2019 Eurygnathodus costatus Staesche ; Li et al., p. 6, fig. 4.22–4.36.

2019 Eurygnathodus costatus Staesche ; Lyu et al., fig. 7.12.

2019a Eurygnathodus costatus Staesche ; Wu et al., fig. 4.20.

Material.> 30 specimens.

Remarks. Goel (1977), Matsuda (1984), Koike (1988) and Maekawa et al. (2018) documented a large intraspecific variation of the platform morphology and its oral ornamentation. It follows that Platyvillosus paracostatus Wang and Cao (1981) likely corresponds to a variant of Eu. costatus (Form A in Koike 1988). Maekawa et al., (2018, figs. 25.1d) documented also specimens covered with microgranules.

Occurrence. Tis species has been reported worldwide from the early Smithian Flemingites zone. Japan: Tahogawa member within the Novispathodus ex gr. waageni Zone ( Koike, 1988; Maekawa et al., 2018), Europe: South Tyrol, Slovenia, Croatia and Bosnia and Herzegovina (Aljinović et al., 2006; Chen et al., 2016; Kolar-Jurkovšek et al., 2021; Staesche, 1964), India: Spiti and Kashmir ( Goel, 1977; Matsuda, 1984; Orchard, 2010; Orchard & Krystyn, 2007), Russia; South Primorye ( Shigeta, 2009), Canada: British Columbia (Beyer & Orchard, 1991), South China and North-eastern Vietnam ( Chen et al., 2015; Shigeta et al., 2014).

Figures

Fig. 14 Neogondolellinae, Novispathodinae and uncertain from Shanggang. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, Eurygnathodus costatus (Staesche); SHA325, PIMUZ 39130. B Eurygnathodus hamadai (Koike); SHA326, PIMUZ 39131. C, D, F, G Novispathodus ex gr. waageni (Sweet); C SHA328,PIMUZ 39238; D SHA328, PIMUZ 39239; F SHA325, PIMUZ 39240;G SHA326, PIMUZ 39241.E Neospathodus ex gr. cristagalli (Huckriede); SHA330, PIMUZ 39182.H Neospathodus dieneri (Sweet); SHA338, PIMUZ 39181