Elmomorphus sulcatus, Selnekovič & Jäch & Kodada, 2024

Selnekovič, Dávid, Jäch, Manfred A. & Kodada, Ján, 2024, Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae), European Journal of Taxonomy 957, pp. 1-229 : 66-69

Elmomorphus sulcatus sp. nov.

urn:lsid:zoobank.org:act:968EDB5A-A7FD-47A9-98E0-E02FDB215071

Figs 29–30 , 107B–F , 109F

Differential diagnosis

Elmomorphus sulcatus sp. nov. ( Fig. 29 ) is characterised by having a plastron on the cranial surface (except for a semicircular area on the vertex), on the anterior angles of the pronotum, and on the lateral sides of the elytra. The body is small and broadly oval ( Fig. 29 ). The elytron has nine longitudinal rows of large punctures and numerous small punctures scattered between the rows. Similar characters are also present in E. hongkong sp. nov. and E. depressus sp. nov. The elytral striae are well impressed. The strial punctures are close to each other, not well separated, often confluent, while in E. depressus , they are well separated by ca 0.5–2.0 × puncture diameters. Moreover, the first two ventrites lack plastron on the median portions, while in E. hongkong and E. depressus , the plastron is missing only on the median portions of the first ventrite. The phallobase is longer (PhL/PrL: 1.40–2.00 (1.80 ± 0.17, n =10)) and the parameres differ from those of E. hongkong (PhL/PrL: 1.93–2.02 (n =3); Fig. 30 ).

Etymology

The epithet is a Latin adjective in the nominative singular, referring to the presence of nine well-impressed longitudinal striae on each elytron.

Type material

Holotype

CHINA – Guangxi Autonomous Region • ♂; “CHINA: SE-Guangxi Distr. Yulin Liuwan Mts. SW Yulin | 16.11.1993 350-400m leg. Schönmann (20)”; IAECAS.

Paratypes

CHINA – Guangxi Autonomous Region • 2 ♂♂, 1 ♀; same collection data as for holotype; CKB, NMW • 1 ♀; “ CHINA: SE-Guangxi, Distr. Yulin, Liuwan Mts. , SW Yulin | 17.11.1993, Kuishan , 300m leg. Schillhammer (22) [CWBS 44]”; NMW • 1 ♀; “ CHINA: SE-Guangxi, Distr. Yulin, Liuwan Mts., SW Yulin | 20.11.1993 120 m, (26) [CWBS 49] leg. Schönmann ”; NMW. – Fujian Province • 1 ♀; “ CHINA: Fujian, Longyan 2 km S Dachi, 750m Ceyan Shan , 29.1.1997 leg. Ji & Wang (CWBS 262)”; NMW • 1 ♀; same label data, but “leg. H.Schönmann ”; NMW. – Guangdong Province • 3 ♂♂, 1 ♀; “ CHINA: Guangdong Prov. 37 km E Zengcheng 23°16′33″N 114°03′27″E 10.11.2001, ca. 200 m Komarek & Wang (CWBS 488)”; NMW • 4 ♂♂, 5 ♀♀; “ CHINA: Guangdong Prov. 38 km ENE Zengcheng 23°16′37″N 114°03′19″E 11.11.2001, ca. 200 m, Komarek & Wang (CWBS 489)”; CKB, NMW .

Type locality

China, Guangxi Autonomous Region, Yulin Prefecture; Liuwan Da Shan (= Sixty-thousand Mountains), 30 km southwest of Yulin City; several streams in the vicinity of Liuwan Forest Station, 0.5–2.0 m wide, rather sandy, original riparian vegetation, slopes covered with a planted forest of mainly Cunninghamia lanceolata , crystalline rock, 350–400 m a.s.l. (CWBS 42; Jäch & Ji 1995).

Description

Measurements (mm): TL: ♂♂ 2.36–2.61 (2.44 ± 0.08, n =10), ♀♀ 2.53–2.76 (2.63 ± 0.07, n=10); PL: ♂♂ 0.61–0.67 (0.64± 0.02, n=10), ♀♀ 0.65–0.72 (0.69 ± 0.02, n =10); PW: ♂♂ 1.11–1.22 (1.16 ± 0.03, n =10), ♀♀ 1.16–1.30 (1.23 ±0.03, n =10); EL: ♂♂ 1.73–1.96 (1.80± 0.07, n =10), ♀♀ 1.87–2.04 (1.94 ±0.05, n=10); EW: ♂♂ 1.31–1.42 (1.36± 0.04, n= 10), ♀♀ 1.35–1.57 (1.46± 0.06, n= 10); PhL: 0.36–0.52 (0.48 ± 0.04); PrL: 0.25–0.29 (0.27± 0.01).

Body ovate, moderately convex dorsally ( Fig. 29 ). Integument black; mouthparts, antennae, and legs reddish brown. Pubescence consists of very short and sparse yellowish setae. Plastron present on head (except for small semicircular area on vertex), on anterior angles of pronotum, lateral portions of elytra, and ventral surface, except for prosternal process, middle of metaventrite, and middle of ventrites 1–2.

Dorsal surface of head with round punctures slightly smaller than diameter of an eye facet, separated by 0.5–1.0 × puncture diameter; surface covered with plastron, except for semicircular area on vertex. Labrum transverse with anterior margin broadly emarginate; exposed portion microreticulate, with minute setiferous punctures; setae equal in length in both sexes. Anterior margin of clypeus straight. Eyes oval, interfacetal setae short; ID: ♂♂ 0.41–0.46 mm (0.44 ± 0.02, n =10), ♀♀ 0.42–0.49 mm (0.47 ± 0.02, n=10); APD/ID: ♂♂ 1.44–1.73 (1.63± 0.08, n= 10), ♀♀ 1.56–1.66 (1.61 ± 0.04, n =10). Antennae 10-segmented, densely setose.

Pronotum transverse, moderately convex, widest at base, PW/PL: ♂♂ 1.77–1.87 (1.81 ± 0.04, n= 10), ♀♀ 1.74–1.83 (1.78 ±0.03, n=10); anterior angles strongly deflexed, prominent; lateral sides convergent, rounded; surface smooth, with round punctures; plastron confined to anterolateral portions. Prosternal process wider than long, lateral edges divergent, straight, posterior edge rounded; lateral portions moderately raised, without groups of long setae; median keel rather flat. Scutellum wider than long, smooth, with round punctures. Metaventral disc slightly convex, glabrous area without plastron widening posteriorly. Lateral margins of metaventral process moderately raised. Elytra ovate, dorsally convex, widest around middle, EL/EW: ♂♂ 1.29–1.38 (1.32± 0.03, n= 10), ♀♀ 1.25–1.38 (1.33± 0.04, n= 10); surface distinctly microreticulate, with small, scattered punctures and large deep punctures arranged in nine longitudinal rows, punctures often confluent; plastron confined to lateral band, anteriorly almost reaching elytral base, extending from elytral lateral margin to sixth elytral row. Tibiae slightly curved; protibia ca 1.3× as long as protarsus; PrTL/PL: ♂♂ 0.99–1.10 (1.03± 0.03, n=10), ♀♀ 0.93–1.02 (0.98± 0.03, n=10). Terminal protarsomere as long as three preceding tarsomeres combined.

Ventrites covered with plastron, except for middle of ventrites 1–2. Ventrite 5 in females with short longitudinal keel before apex. Aedeagus ( Fig. 30 ): parameres shorter than phallobase, widest at basal third, strongly narrowed to narrow subacute apices, dorsal outline strongly arcuate, ventral one almost straight (all in lateral aspect); phallobase narrow, PhL/PrL: 1.40–2.00 (1.80± 0.17, n =10); penis rounded apically; sclerotised fibula distinct.

Secondary sexual dimorphism

Females with short longitudinal keel at the apex of ventrite 5.

Distribution

China (Fujian, Guangdong, Guangxi) ( Fig. 109F ).

Figures

Fig. 29. Elmomorphus sulcatus sp. nov., paratype, male from type locality (CKB), TL: 2.44 mm.

Fig. 30. Elmomorphus sulcatus sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.

Fig. 107. Habitats of selected species of Elmomorphus Sharp, 1888. A. Vietnam, Phong Mỹ, Sông Ô Lâu River (16°31′15.3″ N, 107°15′00.3″ E); E. striatellus Delève, 1968 and E. siamensis Kodada, Selnekovič & Jäch sp. nov. were found in clumps of leaves and on rootlets of Neptunia sp. B. Vietnam, Thành SƠn (20°30′01.4″ N, 105°06′19.3″ E); E. dentipes sp. nov., and E. similis sp. nov. were found in clumps of leaves.C. Vietnam, ThànhSƠn,Kho Mường(20°28′51.3″N,105°07′51.6″E); E. paramontanus Kodada, Selnekovič & Jäch sp. nov., E. similis sp. nov., and E. cf. sulcatus were found on submerged rootlets of bamboo and banana trees. D. Same locality, showing submerged rootlets. E. Vietnam, Trung Mỹ (21°23′35.3″N, 105°42′52.7″E); E. cf. sulcatus was found in clumps of leaves and on submerged rootlets of bamboo, Bauhinia sp., and Sterculia lanceolata Cav. F. Same locality, clump of leaves and partly submerged rootlets.

Fig. 109. Distribution maps. A. Elmomorphus fusiformis sp. nov. B. E. elmoides sp. nov. C. E. calvus sp. nov. D. E. hongkong sp. nov. E. E. depressus sp. nov. F. E. sulcatus Kodada, Selnekovič & Jäch sp. nov.