Elmomorphus umphangicus Kodada, Selnekovič & Jäch, 2024

Selnekovič, Dávid, Jäch, Manfred A. & Kodada, Ján, 2024, Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae), European Journal of Taxonomy 957, pp. 1-229 : 97-101

Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov.

urn:lsid:zoobank.org:act:309D3D42-6B56-4D27-B75A-7B8F80FD8B03

Figs 44–45 , 111C

Differential diagnosis

Elmomorphus umphangicus sp. nov. ( Fig. 44 ) is characterised by having a plastron on the cranial surface (except for a small triangular area on the vertex) and on the lateral sides of the pronotum and elytra. Elytral punctures are scattered over the entire surface and longitudinal striae are absent. Such a combination of characters is also present in E. bispinosus sp. nov., E. longitarsis sp. nov., E. parvulus sp. nov., E. sausai sp. nov., and E. yunnanensis sp. nov. The elytral plastron bands are slenderer than in E. sausai . Elytra are widest behind the middle in E. umphangicus , while in the rest of the species, they are widest around the middle. Elmomorphus umphangicus is smaller (TL: ♂♂ 2.64–2.90 mm (2.76±0.10, n=5), ♀♀ 2.46– 3.12 mm (2.83 ±0.30, n=6)) than E. bispinosus (TL: ♀♀ 3.38–3.54 mm (n=3)), E. longitarsis sp. nov. (TL: ♂♂ 2.93–3.09 mm (n =3)), E. sausai (TL: ♂♂ 3.07–3.15 mm (3.10±0.03, n=4)), or E. yunnanensis (TL: ♂♂ 3.29–3.67 mm (3.37 ±0.15, n=4)) and slightly larger than E. parvulus (TL: ♂♂ 2.41–2.62 mm (2.54±0.07, n=6), ♀♀ 2.60–2.81 mm (2.71± 0.07, n=6)). Pronotal and elytral surface with distinct bronze lustre ( Fig. 44 ), not present in E. yunnanensis ( Fig. 50 ). Aedeagus as in Fig. 45 .

Etymology

The epithet is a Latin adjective in the nominative singular referring to the type locality of the species.

Type material

Holotype

THAILAND – Tak Province • ♂; “THAI, 26.IV.-6.V. 1991 UMPHANG 500 m 16°04′N 98°53′E Vít Kubáň leg.”; NMW.

Paratypes

THAILAND – Tak Province • 2 ♂♂, 6 ♀♀; same collection data as for holotype; CKB, NMW • 3 ♀♀; with similar labels, but “ David Král lgt.”; NMW .

Remarks

A series of 67 specimens (NMW) labelled “ Thailand 2003 Prov. Suratthani [Surat Thani] Khao Sok N.P. 15.1. leg.: Horst Forster”, closely resembles the type specimens of E. umphangicus sp. nov., but they differ in the slightly more convex elytra and the slightly narrower elytral plastron bands. Although we could not detect any apparent differences in the male genitalia, we are not including these specimens from Khao Sok N.P. (Surat Thani Province) in the type series or as “additional material examined” of E. umphangicus ; molecular data will be necessary to reveal the identity of these specimens.

Type locality

Thailand, Tak Province, Umphang, 16°4′ N, 98°53′ E.

Description

Measurements (mm): TL: ♂♂ 2.64–2.90 (2.76± 0.10, n= 5), ♀♀ 2.46–3.12 (2.83 ± 0.30, n =6); PL: ♂♂ 0.61–0.66 (0.63± 0.02, n =5), ♀♀ 0.53–0.71 (0.63± 0.08, n =6); PW: ♂♂ 1.06–1.17 (1.11 ±0.04, n=5), ♀♀ 0.96–1.26 (1.11 ±0.13, n=6); EL: ♂♂ 2.03–2.24 (2.13± 0.07, n=5), ♀♀ 1.92– 2.42 (2.20 ± 0.22, n =6); EW: ♂♂ 1.30–1.43 (1.35± 0.05, n=5), ♀♀ 1.17–1.51 (1.36 ± 0.14, n =6); PhL: 0.53–0.64 (0.60± 0.05, n =5); PrL: 0.31–0.34 (0.32 ±0.01, n =5).

Body elongate oval, widest behind midlength of elytra, moderately convex dorsally ( Fig. 44 ). Colouration black; mouthparts, antennae, trochanters, and tarsi reddish brown, remaining parts of legs dark brown. Dorsal pubescence consists of thin short decumbent setae arising from small round punctures. Dorsal plastron on cranium except for triangular area on vertex, on lateral portions of pronotum and elytra. Plastron present on ventral surface, except for prosternal process, median part of metaventrite, and middle of ventrites 1–2.

Cranium with round setiferous punctures smaller than eye facet and separated by 0.5–1.0 × puncture diameter. Plastron on dorsal cranial surface, except for triangular area on vertex. Labrum transverse with straight anterior margin; exposed portion microreticulate with small setiferous punctures; setae concentrated mainly along anterior margin. Anterior margin of clypeus straight. Eyes large, oval, moderately protruding, ID: ♂♂ 0.37–0.42 mm (0.40± 0.02, n =5), ♀♀ 0.34–0.45 mm (0.41± 0.04, n= 6). Antennae 10-segmented, densely setose.

Pronotum transverse, weakly convex, widest at base, PW/PL: ♂♂ 1.74–1.77 (1.76± 0.01, n=5), ♀♀ 1.71– 1.83 (1.77± 0.05,n =6). Plastron forming medially constricted band along lateral pronotal side. Remaining pronotal surface smooth, with round setiferous punctures slightly larger than those on head and subequal elytral punctures. Anterior pronotal angles protruding and deflexed; lateral pronotal sides convergent, straight, or weakly rounded. Prosternal process with lateral and posterior edges rounded; lateral portions weakly raised, without clusters of long setae; median keel flat. Scutellum longer than wide, smooth, with several setiferous punctures. Metaventral process with lateral margins slightly raised; metaventral disc flat, without clusters of long setae; posteriorly widened median portion smooth, without plastron. Elytra ovate, moderately convex, widest behind midlength, EL/EW: ♂♂ 1.56–1.63 (1.58 ±0.03, n=5), ♀♀ 1.58–1.64 (1.62 ± 0.02, n= 6). Plastron in two posteriorly expanded lateral bands, each covering about half of elytron width (along elytral midlength); remaining elytral surface weakly microreticulate, with round setiferous punctures. Tibiae straight; protibia as long as protarsus; PrTL/PL: ♂♂ 1.17–1.28 (1.21 ± 0.05, n=5), ♀♀ 1.17–1.34 (1.25± 0.07, n= 6). Terminal protarsomere as long as all preceding segments combined. Male foreclaws narrow, strongly curved, similar to those of female.

Ventrites covered with plastron, except for middle of ventrites 1–2. Admedian keels on ventrite 1 inapparent. Ventrite 5 evenly convex in both sexes; apex in males slightly excised, in females rounded with short longitudinal keel. Aedeagus ( Fig. 45 ): phallobase rather short, PhL/PrL: 1.69–2.00 (1.85– 0.14, n=5); parameres narrowed and rounded apically (lateral view), weakly curved; penis narrowly rounded at apex; sclerotised fibula slender.

Secondary sexual dimorphism

Apex of male ventrite 5 shallowly excised, in female rounded, with short longitudinal keel.

Distribution

Thailand ( Fig. 111C ).

Figures

Fig. 44. Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov., paratype, male from type locality (CKB), TL: 2.80 mm.

Fig. 45. Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov., aedeagus. A. Holotype (NMW), lateral aspect. B. Holotype (NMW), ventral aspect. C. Paratype from type locality (CKB), lateral aspect. Scale bar: 0.1 mm.

Fig. 111. Distribution maps. A. Elmomorphus paramontanus Kodada, Selnekovič & Jäch sp. nov. B. E. cuneatus sp. nov. C. E. umphangicus Kodada, Selnekovič & Jäch sp. nov. D. E. parvulus sp. nov. E. E. sausai Kodada, Selnekovič & Jäch sp. nov. F. E. longitarsis sp. nov.

Fig. 50. Elmomorphus yunnanensis Kodada, Selnekovič & Jäch sp. nov., paratype, male from type locality (CKB), TL: 3.37 mm.