Elmomorphus bispinosus, Selnekovič & Jäch & Kodada, 2024

Selnekovič, Dávid, Jäch, Manfred A. & Kodada, Ján, 2024, Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae), European Journal of Taxonomy 957, pp. 1-229 : 115-119

Elmomorphus bispinosus sp. nov.

urn:lsid:zoobank.org:act:B9F8E28F-FD2D-4D67-99E4-580CD3200EBA

Figs 54 , 55D , 112A

Differential diagnosis

Elmomorphus bispinosus sp. nov. ( Fig. 54 ) is characterised by the presence of a plastron on the dorsal cranial surface (except on the anteriorly narrowed area on the vertex) and on the lateral sides of the pronotum and elytra. Elytral punctures are scattered, and striae are absent. Similar characters are also present in E. sausai sp. nov., E. umphangicus sp. nov., E. yunnanensis sp. nov., and E. longitarsis sp. nov. Elmomorphus sausai differs in the distinctly wider elytral plastron bands, each band occupying ca two thirds of elytron width, while in E. bispinosus it is ca one-third. Elmomorphus umphangicus differs in smaller body dimensions (TL in E. umphangicus : ♂♂ 2.64–2.90 mm (2.76 ±0.10, n= 5), ♀♀ 2.46– 3.12 mm (2.83 ±0.30, n=6), in E. bispinosus : ♀♀ 3.38–3.54 mm (n =3)) and the elytra being widest behind midlength. Elmomorphus bispinosus differs from E. yunnanensis in its less convex body and in the presence of two microsclerites on the bursa copulatrix versus several microsclerites in E. yunnanensis ( Fig. 55D ). From E. longitarsis it differs in the larger (TL in E. bispinosus : ♀♀ 3.38–3.54 mm (n=3), in E. longitarsis : ♂♂ 2.93–3.09 mm (n =3), ♀♀ 3.15–3.32 mm (n= 2)), broader, and less convex body and the presence of two spiny microsclerites on the bursa copulatrix.

Etymology

The epithet is a Latin adjective in the nominative singular, referring to the presence of two spiny microsclerites on the bursa copulatrix.

Type material

Holotype

CHINA – Hunan Province • ♀; “CHINA: SW Hunan 1993 SW Huitong , 7. 11. Umg. Guangping , 400m leg. Schönmann (14) [CWBS 35]”; IAECAS.

Paratypes

CHINA – Hunan Province • 2 ♀♀; same collection data as for holotype, but one specimen “leg. L. JI ”; NMW.

Type locality

China, Hunan Province, Huaihua Prefecture, Huitong county, Guangping Township, near Moshao village, small stream, 0.5–1.0 m wide, partly canalised, partly vanishing beneath the gravel, ca 400 m a.s.l. (CWBS 35; Jäch & Ji 1995).

Description

Measurements (mm): TL: ♀♀ 3.38–3.54 (n =3); PL: ♀♀ 0.75 (n =3); PW: ♀♀ 1.44–1.48 (n =3); EL: ♀♀ 2.67–2.86 (n= 3); EW: ♀♀ 1.72–1.82 (n =3).

Body oval, moderately convex dorsally ( Fig. 54 ). Integument black; mouthparts, antennae, and tarsi reddish brown. Pubescence consisting of short yellowish setae. Plastron present on cranial surface except anteriorly narrowed area on vertex, on lateral sides of pronotum and elytra, and on ventral surface except for prosternal process, median part of metaventrite and median part of ventrites 1 and 2.

Head with round punctures separated by 0.5–1.0 × puncture diameter. Plastron present on entire cranial surface except on anteriorly narrowed area on vertex. Labrum transverse, anterior edge straight, exposed portion microreticulate; setae concentrated along anterior margin. Anterior margin of clypeus straight. Eyes oval, moderately protruding, ID: ♀♀ 0.48–0.50 mm (n =3). Antennae 10-segmented, densely setose.

Pronotum transverse, widest at base, moderately convex, PW/PL: ♀♀ 1.92–1.97 (n =3); anterior angles protruding, acute; lateral sides convergent, rounded; surface smooth, with round setiferous punctures; plastron bands present along entire lateral sides. Lateral and posterior edges of prosternal process rounded; lateral portions raised; median keel arcuate. Lateral sides of metaventral process weakly raised. Disc of metaventrite weakly convex, without clusters of long setae. Elytra oval, moderately convex, widest around midlength, EL/EW: ♀♀ 1.55–1.61 (n =3); surface weakly microreticulate with irregular meshes, with weak bronze lustre; round setiferous punctures scattered; lateral plastron bands widened posteriorly, covering about one-third of elytron width at elytral midlength. Tibiae straight, protibia ca 1.2 × as long as protarsus, PrTL/PL: ♀♀ 1.16–1.26 (n =3). Terminal protarsomere longer than all preceding segments combined.

Ventrites covered with plastron, except on middle of ventrites 1 and 2. Ventrite 5 with short longitudinal apical keel. Bursa copulatrix on each side with a rather large microsclerite bearing one prominent spine ( Fig. 55D ).

Secondary sexual dimorphism

Male unknown.

Distribution

China (Hunan) ( Fig. 112A ).

Figures

Fig. 54. Elmomorphus bispinosus sp. nov., holotype, female (IAECAS), TL: 3.54 mm.

Fig. 55. Bursa copulatrix and vagina. A. Elmomorphus parvulus sp. nov., paratype from type locality (CKB). B. E. yunnanensis Kodada, Selnekovič & Jäch sp. nov., paratype from type locality (CKB). C. E. longitarsis sp. nov., paratype from type locality (NMW). D. E. bispinosus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm.

Fig. 112. Distribution maps. A. Elmomorphus bispinosus sp. nov. B. E. nepalensis Satô, 1981. C. E. punctulatus sp. nov. D. E. simplipes sp. nov. E. E. donatus Kodada, Selnekovič & Jäch sp. nov. F. E. paradonatus sp. nov.