Cochlostoma (Clessiniella) stelucarum, Zallot & Mattia & Fehér & Gittenberger, 2021

Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán & Gittenberger, Edmund, 2021, Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 762 (1), pp. 49-95: 78-81

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Cochlostoma (Clessiniella) stelucarum

sp. nov.

Cochlostoma (Clessiniella) stelucarum   sp. nov.

Figs 1 View Fig , 3E View Fig , 14B View Fig , 15B View Fig , 16B, 15–27 View Fig View Fig View Fig View Fig View Fig View Fig View Fig View Fig View Fig View Fig View Fig 25 View Fig 26 View Fig


Slender shell with 8½–10½ rather widely ribbed whorls. Spots always visible. Poorly developed lip with FF or AF external lobe. Visceral oviduct simplified and seminal receptacle sometimes oval.


The epithet ‘ steluca ’ is an acronym noun formed from the initial syllables of the names of the wife and two daughters (Stefania, Ludovica and Camilla Zallot) of the senior author.

Type material ( Fig. 27 View Fig )

Holotype ITALY • ♀ (in ethanol); Friuli Venezia Giulia , Udine, Cornino ; 46.2297° N, 13.0250° E; 180 m a.s.l.; 5 Jun. 2002; Zallot leg.; rocks surrounding the lake; RMNH.MOL.347617. GoogleMaps  

Paratypes ITALY • 1 ♂ (in ethanol); same collection data as for holotype; RMNH.MOL.347618 GoogleMaps   1 ♀ (empty shell); same collection data as for holotype; GenBank voucher H3: KX120830 View Materials ; RMNH.MOL.347619 GoogleMaps   .

Other material

ITALY • Anduins-cliffs ; 46.23755° N, 12.97106° E; 1997; Zallot leg.; EZ0102 GoogleMaps   Campone ; 46.26194° N, 12.83333° E; 2010; Zallot leg.; EZ0098 GoogleMaps   Castelmonte ; 46.094° N, 13.52095° E; Oct. 2010; Margelli leg.; EZ1020 GoogleMaps   Cjanet-Peonis ; 46.25917° N, 13.04111° E; 2002; Zallot leg.; EZ0105 GoogleMaps   Cornino ; 46.22972° N, 13.025° E; 2002; Zallot leg.; EZ0104 GoogleMaps   F. Piccola Mt Valinins ; 46.245° N, 12.80111° E; 2000; Zallot leg.; EZ0095 GoogleMaps   Mt Bernadia ; 46.23361° N, 13.25556° E; 2005; Zallot leg.; EZ0107 GoogleMaps   Mt Ciaurlec ; 46.21226° N, 12.88018° E; 2005; Zallot leg.; EZ0099 GoogleMaps   Mt Prat ; 46.24577° N, 12.9907° E; 2002; Zallot leg.; EZ0103 GoogleMaps   Mt Valinis ; 46.23417° N, 12.8075° E; 2002; Zallot leg.; EZ0096 GoogleMaps   San Vito D’asio ; 46.2337° N, 12.94531° E; 1997; Zallot leg.; EZ0101 GoogleMaps   Tramonti-Campone ; 46.27833° N, 12.81917° E; 2000; Zallot leg.; EZ0097 GoogleMaps   Val di Torre-all’imbocco ; 46.23861° N, 13.23111° E; 1998; Zallot leg.; EZ0106 GoogleMaps   Valle D’Arzino ; 46.30278° N, 12.92972° E; 2000; Zallot leg.; EZ0100 GoogleMaps   .


SHELL. The shell has prominent spots and strong and rounded ribs, which are regularly and widely spaced and coloured like the background. The ribs become less prominent on the body whorl. The body whorl slightly and gradually enlarges while approaching the aperture ( Fig. 4A View Fig : FF, AF). In comparison with the other species of the subgenus, the whorls increase more slowly in size so that, given the same shell height, there is a higher number of whorls ( Fig. 9 View Fig ).

FEMALE GENITALIA. The visceral oviduct has only 1–2 superficial loops, what makes it one of the simplest in the genus Cochlostoma   . The seminal receptacle is thin and may be oval, thus not always club-shaped as in the other species.

MALE GENITALIA. The penis is longer than the body and inflated, although less so than in the other species of Clessiniella   . The penial spermiduct is gently twisted. The body spermiduct is straight. There is a long groove, ending in a well-formed sperm pocket, delimited at the frontal side by an ascending sperm funnel.


The species inhabits a restricted area in the eastern Southern Pre-Alps. Our sampling localities are located between Val Tramontina to the west and Val di Torre to the east (Friuli region, Italy).

Habitat preference

Like in most species of Cochlostoma   , these are obligate rock-dwelling snails but, rather than on the surface of limestone cliffs, their main habitat is on smaller surfaces of stones and boulders which are normally found in large numbers at the foot of the cliffs.


Cochlostoma (Clessiniella) stelucarum   sp. nov. was introduced as Cochlostoma sp.   by Zallot (2002: 98). At most of the localities, it lives syntopically with either Co. (Eupomatias) philippianum (Gredler, 1853)   or Co. (Eupomatias) henricae ( Strobel, 1851)   . Near the village of Anduins it has been found a few centimeters apart from both Co. (Eu.) henricae   and Co. (Cochlostoma) septemspirale   on stones and boulders. This is one of the rare cases where 3 species of Cochlostoma   , belonging to different subgenera, live syntopically. The range of the species is situated in between the range of Co. (Cl.) villae   in the west and that of the other species of Clessiniella   in the east.