Achiderbe obrienae Emeljanov et Shcherbakov, 2020

Emeljanov, A. F. & Shcherbakov, D. E., 2020, The first Mesozoic Derbidae (Homoptera: Fulgoroidea) from Cretaceous Burmese amber, Russian Entomological Journal 29 (3), pp. 237-246 : 242

Achiderbe obrienae Emeljanov et Shcherbakov , sp.n.

Figs 21–24 , 28 .

MATERIAL. Holotype PIN 5608 /126, ♂, most of head and dorsal part of thorax eaten by some arthropod scavenger, distal parts of hind legs polished away; syninclusions: 2 Empidoidea — Burmese amber, Hukawng Valley, Kachin State, Myanmar; mid-Cretaceous (probably Albian–Cenomanian).

DESCRIPTION. Tegmen 3.9 mm long, elongate, 2.8:1, fuscous with pale: irregular transverse bands proximally, crossveins and vein apices distally; stigmal cell pale at nodus, distally of same shade of brown as adjacent cells. Anterior margin shallowly convex near base and at nodus, almost straight between; apical margin obliquely rounded. Costal vein thickened near base, turning flat, slender distad of ScR+M stalk; C and ambient vein with sensory pits, better developed at base, traceable up to apex of tegmen. ScR fork much distad of, and CuA fork distad of, union of claval veins. Nodal r-m level with nodal m-cu. MA fork about halfway from nodal r-m to 2rm. Medial cell long and broad, much larger than cell distal to it or cell anterodistal to it. Clavus truncate at apex, without crossvein. Hind wing ca. 2.9 mm long, 1.2 mm wide, not large, 0.75 as long as tegmen, and narrow, 2.4:1, apex considerably not reaching apex of tegmen in repose; membrane fuscous, veins brown. Costal margin biconvex, coupling lobe in concavity beyond wing midlength. ScRA apex proximad of level of M fork. M fork twice as long as CuA1 fork. Body 3.6 mm long; body and appendages various shades of brown. Anterodorsal part of head not preserved. Scape elongate, obliquely produced anterodorsally, pedicel thrice longer than scape, paddle-shaped, flattened, scape + pedicel about as long as clypeus. Pedicel covered with cuticular denticles, with many plaque organs. Subantennal ridge absent. Postclypeus strongly narrowed towards unicarinate, laterally flattened anteclypeus. Rostrum reaching hind coxae, apical segment dark, about 4 times as long as wide. Pronotum not preserved. Mesonotum with lateral discal carinae strong, subparallel, median carina well-developed. Legs rather short and stout, tibiae tetrahedral, somewhat flattened, hind tibiae without lateral teeth in proximal half (their distal half and hind tarsi not preserved), claws strong. Abdomen slightly inflated, laterotergites rectangular. Male genital block rather small; pygofer ventrally sinuate with median depression between two oblique carinae; anal tube rather long, with posterolateral lobes; styles short and broad, their apices bent mediad.

ETYMOLOGY. Named in honour of Dr Lois B. O’Brien, a worldwide known entomologist, specialist on Fulgoroidea.

Figures

Figs 21–24. Achiderbe obrienae gen. et sp.n., holotype male, Burmese amber: 21 — habitus, ventral; 22 — head, lateroventral; 23 — abdomen, lateroventral; 24 — thorax and bases of wings, laterodorsal (inset, base of tegmen, rotated 180°; arrows, sensory pits on base of C). Scale bars: 21 — 1 mm, 22–24 — 0.5 mm. Рис. 21–24. Achiderbe obrienae gen. et sp.n., голотип, самец, бирманский Янтарь: 21 — обЩий вид, сниЗу; 22 — голова, сниЗу- сбоку; 23 — брЮШко сниЗу-сбоку; 24 — грудь и основаниЯ крыльев, сверху-сбоку (вреЗка, основание переднего крыла, повернуто на 180°; стрелки — сенсорные Ямки на основании C). Длина масШтабной линейки: 21 — 1 мм, 22–24 — 0,5 мм.

Figs 25–28. Venation of Derbachilini: 25–26 — Derbachile aschei sp.n.: 25 — tegmen; 26 — hind wing; 27 — Derbachile hochae gen. et sp.n., tegmen; 28 — Achiderbe obrienae gen. et sp.n., tegmen. Scale bars: 1.0 mm. Рис. 25–28. Жилкование Derbachilini: 25–26 — Derbachile aschei sp.n.: 25 — переднее крыло; 26 — Заднее крыло; 27 — Derbachile hochae gen. et sp.n., переднее крыло; 28 — Achiderbe obrienae gen. et sp.n., переднее крыло. МасШтабные линейки: 1,0 мм.