Cybaeus penedentatus Bennett, 2009

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade, Zootaxa 4965 (3), pp. 401-436 : 417

Cybaeus penedentatus Bennett

Figs 42–54 , 56 , 84

Bennett in Copley et al. 2009: 386, figs 67–78. World Spider Catalog 2021

Type material examined. U.S.A.: California: Holotype male. Santa Clara County, 2 miles north of Holy City, Highway 17, 6.xii.1966, V. D. Roth & Ferguson ( CAS) . Paratypes: Specimens and locality data as in Copley et al. (2009).

Other material examined. U.S.A.: California: Monterey. 2♀, San Juan Grade Rd. , 5.i.1997, D. Ubick ( CAS); Santa Cruz. 2♀, Felton, 24.ii.1985, V . D. Roth & B. Roth ( CAS) .

Diagnosis. Both sexes of C. penedentatus are distinctive and unlikely to be confused with any of the other species in the consocius group. The male is diagnosed by its elongate patellar apophysis (as long as the width of the patella) with very few peg setae ( Fig. 43 ) as well as the posteriorly directed basal flange, longitudinal ventral keel, and acuminate slightly twisted tip of the proximal arm of the tegular apophysis ( Figs 44–45 ). The female is diagnosed primarily by the very small, transverse, medially located atrium ( Figs 46–47, 49–50 , 52 ) and secondarily by the membranous copulatory ducts extending contiguously or in close proximity from the atrium to the anterior margin of the vulva before diverging and heading laterally and posteriorly to merge with the spermathecal heads and stalks ( Figs 47–51 , 53–54 ).

Description. See Copley et al. (2009).

Distribution. Coastal central California west of San Francisco Bay south to Monterey Bay in San Mateo, Santa Clara and Santa Cruz Counties ( Fig. 86 ). The holotype, the single known male specimen of C. penedentatus , was collected in December.

Figures

FIGURE 84. Distribution of Cybaeus consocius, C. hesper, C. hummeli spec. nov., C. penedentatus, and C. ubicki spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized.

FIGURE 86. Distributions of Cybaeus pan spec. nov., C. vulpinus, and C. opulentus spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized.

FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis.

FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium.

FIGURES 52–54. Cybaeus penedentatus, female from Brookdale, California, copulatory organ. 52 Epigynum, ventral. 53–54 Vulva (53 ventral, 54 dorsal). AT—atrium.

FIGURES 55–58. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis (55 C. opulentus spec. nov. from Riverton, 56 C. penedentatus from Holy City, 57 C. vulpinus from Columbia, 58 C. pan from Calpine) (55–56 ventral, 57–58 prolateral). Arrows indicate diagnostic characters (55 slightly twisted tip and prominent basal protrusion; 56 acuminate slightly twisted tip and longitudinal ventral keel; 57 smoothly curved, twisted tip; 58 triangular keel dorsally on tip).