Aulojoppoides, Riedel, 2017

Riedel, Matthias, 2017, Contribution to the Ichneumoninae (Hymenoptera, Ichneumonidae) of Southeastern Asia: 4. Platylabini, Eurylabini, and Oedicephalini, Linzer biologische Beiträge 49 (2), pp. 1275-1307 : 1289-1291

Aulojoppoides nov.gen.

Type species: Aulojoppoides mandibulator nov.sp.

The tribal position of this new genus is questionable. Although the sickle-shaped mandibles are an unusal feature for the tribe Oedicephalini , the structure and coloration of the head is typical for this tribe. In my opinion, this new genus should be included into the Oedicephalini . As already discussed by HEINRICH (1934: 118), Nesostenodontus CUSHMAN might also belong to the Oedicephalini , but has been placed in the tribe Ichneumonini so far ( YU et al. 2016). A detailed analysis of the correct tribal status of both genera will be necessary in the future to solve the taxonomical problems.

Habitus and colour pattern of the type-species of Aulojoppoides strongly resembles Aulojoppa spilocephala CAMERON. It differs from Aulojoppa by the following features:

 Mandibles sickle-shaped, lower tooth very small and completely turned inwards,

 mesoscutum with notauli,

 propodeal spiracles short-oval, not slit-shaped,

 postpectal carinae widely interrupted in front of hind coxae,

 metasoma strongly oxypygous, ovipositor sheaths reaching widely beyond metasomal apex (by the length of hind metatarsi).

From Notosemus FÖRSTER and Imeria CAMERON it can be differentiated by the sickleshaped mandibles which resemble the genus Nesostenodontus CUSHMAN. Aulojoppoides differs from Nesostenodontus CUSHMAN mainly by the wide genae, present epomia, and very large thyridia.

Aulojoppoides mandibulator nov.sp. ( fig. 1 , 3, 4) H o l o t y p u s (♀) Laos, Hua Phan Prov., Phon-Pan, Umg. Ort Ban Saleui, 2013´30´´N

10359´26´´E, 1350-1900 m, 13.VI.2013 KJa, leg. C. Holzschuh & natives (Linz).

Descr iption:

Body length 10.5 mm. Flagella with 30 segments, almost filiform; 1 st flagellomeres length 4.4x width, widest flagellomeres about square. Head large. Ocelli small, distance of lateral ocelli to eyes 1.9x their diameter. Temples wide, parallel behind eyes and slightly narrowed caudally, length c.0.7x eye width. Occiput strongly and concavely excised medially. Vertices and temples with scattered punctures, shining; frons with coarse rugose-punctation medially. Face densely punctate, punctures more scattered laterally. Clypeus large, c.2x wider than long, moderately convex, with scattered punctures; apical margin sharp, slightly rugose (not dentate as in Nesostenodontus), almost straight. Clypeal grooves large. Mandibles strongly sickle-shaped; lower teeth very small and completely turned inwards. Genae wide, seen from lateral as wide as eyes, with very scattered punctures, strongly shining. Malar spaces length c.0.6x width of mandibular bases. Genal carinae reaching mandibular bases.

Mesosoma covered with short whitish hairs. Sides of pronotum almost smooth, upper margins strongly bulging, epomia present. Mesoscutum without notauli, coarse and densely punctate, smooth and shining between punctures. Mesopleura coarsely punctate, specula smooth; metapleura with finer punctures, coxal carinae present. Sternauli impressed in frontal 0.5 of mesopleura; epicnemial carina narrow, reaching center of mesopleura; postpectal carinae widely interrupped in front of hind coxae. Scutellum wider than long, moderately elevated, with very scattered punctures, without lateral carinae. Propodeum completely carinate, without apophyses; spiracles short-oval, c.1.5x longer than wide. Area superomedia halfmoon-shaped, wider than long, costulae in the middle. Area petiolaris slighly impressed, with strong transverse rugae. Hind coxae densely punctate, without scopa or carinae; hind femora densely punctate, length 4.2 height; hind tibiae long, narrowed in basal 0.2, inner apical spurs of hind tibiae c.0.34x as long as hind metatarsi; claws strongly bent apically, not pectinate. Areolets quadrangular, 2 nd recurrent veins in their middle; nervuli interstitial.

Metasoma rather stout, strongly oxypygous; hypopygium very short. Petiolus slightly flattened. Postpetiolus slightly widened, punctate, without lateral or dorsal carinae. 2 nd tergite 1.3x wider than long, 3 rd tergite 2x wider than long. Gastrocoeli slightly impressed, with fine oblique striation; thyridia very large, oblique, width c.3x width of their interval. 2 nd to 5 th tergites coarsely and densely punctate, punctation of 5 th tergite superficial; 6 th and 7 th tergites almost smooth and shining. Ovipositor sheaths reaching widely beyond metasomal apex, seen from dorsal length about as long as 2 nd hind tarsomeres.

Colour: Black. Flagella black; scapes, pedicels and 4 basal flagellomeres reddish brown; flagellomeres 7-13 with ivory stripes. Head black; ivory are palps, mandibles except teeth, clypeus except blackish upper margin, face except short blackish median stripe, frontal orbits, large triangular spots on vertices, malar spaces, wide genae and outer orbits. Mesosoma black; tegulae brownish; ivory are frontal and upper margins of pronotum, wing bases, subtegular ridges, ventral wide band on mesopleura, scutellum, carinal triangles, apical third of metapleura and apical half of propodeum except black area petiolaris. Metasoma black; 1 st metasomal segment red, postpetiolus with ivory apical margin; 2 nd tergite with narrow reddish frontal and caudal margins; 3 rd tergite ivory except black caudal-median band, 6 th and 7 th tergites completely ivory. Legs reddish; fore and middle coxae and trochanters ivory; hind coxae red, with ivory spots dorsally; all tibiae and fore and middle tarsi reddish-brown; hind tibiae diffusely blackish basally and apically; hind tarsi ochreous, apical hind tarsomeres brown. Wings hyaline, pterostigmas dark brown.

3 and hosts unknown.

D i s t r i b u t i o n: Only known from Laos.

Figures

Fig. 1-2: Heads from dorsal of (1) Aulojoppoides mandibulator nov.sp. HT; (2) Aulojoppa spilocephala CAMERON ♀. Fig. 3-4: Aulojoppoides mandibulator nov.sp. HT (3) habitus; (4) head from frontal with sickle-shaped mandibles. Fig. 5: Hind coxa of Satrius spinicoxis nov.sp. HT.