Coffea littoralis A.P. Davis & Rakotonas., 2001

Davis, Aaron P. & Rakotonasolo, Franck, 2001, Three new species of Coffea L. (Rubiaceae) from NE Madagascar, Adansonia (3) 23 (1), pp. 137-146 : 138

Key to three new species of Coffea and C. pervilleana

1. Leaf venation on abaxial surface manifest to prominent ............................................................................ 2

1’. Leaf venation on abaxial surface obscure .................................................................................................... 3

2. Leaf apex shortly caudate to caudate; pedicel ± absent, or 0.1-0.5 mm long, bractlets absent; calyx limb shallowly 5-crenate to undulate, NW Madagascar (Island of Nosy Be) .................................. C. pervilleana

2’. Leaf apex acute to ± rounded, rarely very shortly caudate; pedicel always present, 3-5 mm long, bractlets present; calyx limb truncate to slightly undulate; NE Madagascar.................................................. C. littoralis

3. Leaves broadly obovate, broadly elliptic, or ± orbicular; calyx limb with colleters; calyx and disc prominent on fruit (at apex) ................................................................................................................ C. mcphersonii

3’. Leaves obovate or elliptic; calyx limb without colleters; calyx and disc not prominent on fruit (at apex) .............................................................................................................................. C. ratsimamangae

1. Coffea littoralis A.P. Davis & Rakotonas. , sp. nov.