Gymnetis goryi Janson, 1877

Iorio, Osvaldo Di, 2013, A review of the Cetoniinae (Coleoptera: Scarabaeidae) from Argentina and adjacent countries: systematics and geographic distributions, Zootaxa 3668 (1), pp. 1-87 : 33

Gymnetis goryi Janson, 1877 , new status

Figs. 65–66 (aspects), 74 (parameres)

Gymnetis rufilateris , not Illiger, 1800: Gory & Percheron 1833: 350 (description; distribution), plate 70, Fig. 3 , fide Janson 1877b: 247 (misidentification).

Gymnetis rufilateris , not Illiger, 1800, in part: Blanchard, 1847: 193 (distribution); Bruch 1911a: 214 (catalog; distribution); Blackwelder 1944: 263 (catalog; distribution).

Gymnetis goryi Janson, 1877: 247 (description; distribution); Schoch 1898a: 79 (re-description; distribution); Schenkling 1921: 93 (catalog; distribution; references).

Paragymnetis hieroglyphica subsp. goryi Jans. : Schürhoff 1937: 67 (list; distribution); Blackwelder 1944: 263 (catalog; distribution).

Gymnetis hieroglyphica ssp. goryi Janson, 1877 : Krajčík 1998: 77 (catalog; distribution).

Distribution: BRAZIL ( Gory & Percheron 1833, Schenkling 1921). BOLIVIA: Santa Cruz; Guarayos (Blanchard 1847); Bolivia ( Janson 1877). ARGENTINA: Misiones: in Bruch Collection ( Bruch 1911a); Chaco: in Bruch Collection ( Bruch 1911a); ( Blackwelder 1944).

Material examined: BRAZIL: [Pará]: Santarém, Bosq Collection, 3 exx. [MLPA], one labeled “ Gymnetis / rufilateris / Illig.” [handwritten by Viana], other “ Gymnetis / rufilateris / G. & P. / det. B.C. Ratcliffe 2004 ”, Col. Viana, 2 exx. [MLPA], one labeled “ Gymnetosoma / rufilateris / (Illig.) / A. Martínez det. 1950”; BOLIVIA: 1 ex. [HBBA], “ rufilatris / Illig. / Bolivia ” [handwritten by Burmeister]; Beni: Santa Ana de Yacuma, 3–4-II-2012, A. Borquez leg., 1 ex. [ABJCP], 1 ex. [ODIC], C. Calderon Villavicencio leg., II-2012, 1 ex. [ABJCP], V-2012, 8 exx. [ABJCP], 3 exx. [ODIC], X-2012, 1 ex. [ABJCP], 22-XII-2012, 1 ex. [ABJCP]; Santa Cruz: “S. Cruz / Boliv.” [handwritten by Burmeister], 1 ex. [HBBA]; “Yungas de La Paz ”, F. Germain leg., 1 ex. [MLPA], “ Gymnetis sp. ”; ARGENTINA: Misiones: [without locality], Richter Collection, 1 ex. [MLPA] (black), “ Gymnetis / rufilateris / Illig.”, “ Gymnetis / goryi / Janson / det. B.C. Ratcliffe 2004 ”; Gob. Misiones, 190_, C. Bruch, 1 female [CBBA], “ Gymnetis / rufilateris / Illg.” [handwritten by Bruch], “ Gymnetis / rufilateris ” [handwritten by Moser, cut] (Fig. XX); Chaco: “Gob. Chaco / 1 1896 / C. Bruch”, 2 exx. [CBBA], one labeled “ Gymnetis / rufilateris / Illig.” [handwritten by Bruch]; Without locality: A. Breyer Collection, 2 females [MACN] (black), one labeled “ Gymnetis / rufilatris Ill. / C.B. det.” [handwritten by Bruch]; 1 male [ODIC], 1 ex. [ODIC].

Male genitalia: parameres ( Fig. 74 ).

Note 1: Janson (1877) described Gymnetis goryi from Bolivia, which corresponds to the illustration of G. rufilateris not Illiger, given by Gory & Percheron (1833) ( Fig. 64 ). As G. rufilateris sensu Gory & Percheron (1833) was recorded from Brazil, the distribution of G. goryi includes Brazil and Bolivia. While discussing G. goryi, Schürhoff (1937) mentioned that “the locality is not Brazil [error for Bolivia?] but Ecuador, instead, rufilateris G. & P. is from Brazil, and thus probably identical to rufilateris Ill. ” (translated from German). The mention of Ecuador by Schürhoff (1937), and repeated by Blackwelder (1944) and Krajčík (1998), may be erroneous and needs corroboration. Blackwelder (1944) also erroneously listed G. rufilateris Illiger from Bolivia and Argentina (Chaco). This was based on Bruch (1911a) (= Gymnetis goryi ). Gymnetis rufilateris Illiger urgently needs a redescription based on the type specimens.

Note 2: On page 67, Schürhoff (1937) listed G. goryi as a subspecies of Paragymnetis hieroglyphica ( Vigors, 1824) . The general aspect (Figs. 65–66) and the male parameres ( Fig. 74 ) of G. goryi are completely different from those of G. hieroglyphica (Figs. 67–68 and 75 respectively). Therefore, the status of G. goryi as a separate species is re-established here.

Note 3: One black specimen with interrupted, red lateral bands on the elytra in MLPA was identified by B. Ratcliffe as being to G. goryi , and was identical to another specimen in the MACN (Fig. 66). It seems that some specimens tend to be melanic, with the orange and/or red lateral band on the elytra always interrupted by the dark maculation (black background color) ( Fig. 144 ). In contrast, the light specimens, similar to that illustrated by Gory & Percheron (1833) ( Fig. 64 ), may have the red lateral bands on elytra interrupted or not by the dark maculation.

Figures

FIGURES 64–71. General aspects of Gymnetini: 64, Gymnetis rufilateris (extracted from Gory & Percheron 1833); 65–66, Gymnetis goryi (65, Chaco, Bruch Collection; 66, melanic specimen); 67–68, Gymnetis hieroglyphica (67, extracted from Vigors 1824; 68, Paraguay); 69, Gymnetis schistacea (Argentina); 70–71, Gymnetis flaveola (Brazil).

FIGURES 3–7. General aspects of Gymnetini: 3–5, Gymnetis bajula (3, darker specimen; 4, specimen from the Burmeister Collection [MACN], likely the holotype of G. perplexa; 5, lighter specimen); 6, Gymnetis bruchi (Bruch Collection); 7, Aemilius wagneri (extracted from Le Moult 1939).

FIGURES 72–78. Male parameres: 72, Gymnetis flaveola (Brazil); 73, Gymnetis schistacea (Argentina); 74, Gymnetis goryi; 75, Gymnetis hieroglyphica; 76, Hologymnetis sp. (Bruch Collection); 77, Hologymnetis undulata; 78, Heterocotinis semiopaca.

FIGURES 143–146. General aspects of Gymnetini distributed outside Argentina (Appendix II): 143, Gymnetis sp.; 144, Gymnetis chevrolati (= G. ramulosum); 145–146, Gymnetis punctipennis.