Lindholmiola lens (A. FéRUSSAC, 1832)

Subai, Peter & Neubert, Eike, 2014, Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae), Contributions to Natural History 23, pp. 1-94 : 5-33

Lindholmiola lens (A. FéRUSSAC, 1832)

Figs 3–6 , 7–11 , 12 , 15

1821 Helix barbata A. FéRUSSAC , Tableaux systématiques des animaux mollusques classés en familles naturelles: 41 (Folio edition) or 37 (Quarto edition), Nr. 152 [partim].

1832 Helix lens A. FéRUSSAC (in A. Férussac & Deshayes), Histoire naturelle générale et particulière des mollusques terrestres et fluviatiles, (Atlas), 22–27: Taf. 66, F. 2 [Lectotype, design. Gittenberger & Groh 1986].

1832 Helix barbata , – Deshayes, Exped. Moree Moll., p. 162 (non barbata FéRUSSAC ).

1838 Helix lens , – Rossmässler, Iconographie der Land- & Süsswasser-Mollusken mit vorzüglicher Berücksichtigung der europäischen noch nicht abgebildeten Arten, (1) 2 (1/2): 10, Taf. 32, Fig. 450.

1846 Helix lens , – L. Pfeiffer, In: Martini & Chemnitz: Systematisches Conchylien-Cabinet, (1) 12 (II): 101, Taf. 13, Fig. 16–17 .

1848 Helix lens , – L. Pfeiffer, Monographia heliceorum viventium, 1: 209.

1850 Helix lens , – Deshayes (in Férussac & Deshayes), Histoire naturelle générale et particulière des mollusques terrestres et fluviatiles, (Text), 1: 110 [partim, nec descr. et terr. typ.].

1853 Helix lens , – L. Pfeiffer, Monographia heliceorum viventium, 3: 162.

1854 Helix lens , – Reeve, Conchiologica Iconica, 7: species 1221.

1855 Helix lens , – Roth, Malakozoologische Blätter, 2: 30.

1859 Helix lens , – Mousson, Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich, 4: 258.

1859 Helix lentiformis MOUSSON , Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich, 4: 258. Locus typicus: " Thessalie et Attique ".

1859 Helix lens , – L. Pfeiffer, Monographia heliceorum viventium, 4: 187.

1868 Helix lens , – L. Pfeiffer, Monographia heliceorum viventium, 5: 260.

1868 Helix lentiformis , – L. Pfeiffer, Monographia heliceorum viventium, 5: 260.

1873 Helix barbata , – Martens, Malakozoologische Blätter, 20: 32.

1873 Helix lens , – Martens, Malakozoologische Blätter, 20: 32.

1874 Helix lens , – Martens, Malakozoologische Blätter, 21: 122.

1876 Helix lens , – L. Pfeiffer, Monographia heliceorum viventium, 7: 295.

1876 Helix lens , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, Prodromus, I: 40.

1876 Helix lens var. aliostoma WESTERLUND , Fauna der in der paläarctischen Region lebenden Binnenconchylien, Prodromus, I: 41.

1876 Helix lens var. lentiformis , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, Prodromus, I: 41.

1877 Helix lens , – L. Pfeiffer, Monographia heliceorum viventium, 8: 576.

1879 Helix (Trigonostoma) lens , – Westerlund & Blanc, Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 34.

1879 Helix (Trigonostoma) lens var. lentiformis , – Westerlund & Blanc, Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 35.

1879 Helix (Trigonostoma) lens var. piligera WESTERLUND & BLANC , Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 36. Taf. 1, Fig. 10 .

1879 Helix (Trigonostoma) lens var. callojuncta WESTERLUND & BLANC , Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 36. Taf. 2, Fig. 11 .

1882 Helix lens , – Hesse, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 9: 320.

1882 Helix lens var. lentiformis , – Hesse, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 9: 320.

1883 Helix (Anchistoma) lens , – O. Boettger, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 10: 323.

1883 Helix (Anchistoma) lens var. elia O. BOETTGER , Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 10: 330.

1884 Helix lens , – Hesse, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 11: 233, Taf. 4, Fig. 5 , Fig. 5 a .

1885 Helix (Gonostoma) lens var. piligera , – O. Boettger, in Stussiner & O. Boettger, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 12: 165.

1885 Helix (Gonostoma) lens var. lentiformis , – O. Boettger, in Stussiner & O. Boettger, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 12: 165.

1887 Helix (Caracolina) lens , – Tryon, Manual of Conchology, 3: 119.

1887 Helix (Caracolina) lens var. lentiformis , – Tryon, Manual of Conchology, 3: 119.

1887 Helix (Caracolina) lens var. piligera , – Tryon, Manual of Conchology, 3: 119.

1887 Helix (Caracolina) lens var. callojuncta , – Tryon, Manual of Conchology, 3: 119.

1887 Helix (Caracolina) lens var. aliostoma , – Tryon, Manual of Conchology, 3: 119.

1889 Helix (Caracollina) lens , – Martens, Archiv für Naturgeschichte, 55 (1): 172.

1889 Helix (Gonostoma) lens var. lentiformis , – Martens, Archiv für Naturgeschichte, 55 (1): 173.

1889 Helix (Caracolina) lens , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 22.

1889 Helix (Caracolina) lens var. lentiformis , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 22.

1889 Helix (Caracolina) lens var. piligera , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 23.

1889 Helix (Caracolina) lens var. callojuncta , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 23.

1889 Helix (Caracolina) lens var. elia , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 23.

1891 Helix (Gonostoma) lens var. callojuncta , – O. Boettger, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 23 (5/6): 83.

1892 Helix lens , – Schuberth, Archiv für Naturgeschichte, 58 (1): 8, Taf. 1 Fig. 6 .

1892 Helix lens var. lentiformis , – Schuberth, Archiv für Naturgeschichte, 58 (1): 8, Taf. 1, Fig. 7–8 .

1894 Helix (Caracollina) lens , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) lens var. lentiformis , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) lens var. piligera , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) lens var. callojuncta , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) lens var. aliostoma , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) lens var. elia , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Anchistoma) lens , – O. Boettger, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 26 (1–2): 2.

1902 Helix (Gonostoma) lens , – Sturany, Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse Wien, 52: 403.

1902 Helix (Gonostoma) barbata , – Sturany, Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse Wien, 52: 403 (non barbata FéRUSSAC ).

1918 Caracollina lens , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1918 Caracollina lens lentiformis , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1918 Caracollina lens aliostoma , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1918 Caracollina lens piligera , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1918 Caracollina lens callojuncta , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1918 Caracollina lens elia , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1928 Helicodonta lens , – Adensamer & Käufel, Sitzungsberichte der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 137 (10): 793.

1928 Helicodonta lens piligera , – Adensamer & Käufel, Sitzungsberichte der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 137 (10): 793.

1928 Helicodonta lens lentiformis , – Adensamer & Käufel, Sitzungsberichte der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 137 (10): 793.

1929 Helicodonta (Caracollina) lens , – Gambetta, Archivio Zoologico Italiano, 13 (1/2): 51.

1930 Caracollina lens , – Käufel, Sitzungsberichte der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 139: 184.

1931 Lindholmiola lens , – Hesse, Zoologica, 31 (81) 1/2: 50.

1936 Lindholmiola lens , – Fuchs & Käufel, Archiv für Naturgeschichte, N.F. 5 (4): 651. Fig. 77 [partim, not in the Angista canyon close to Drama]

1941 Lindholmiola (Helicodonta) lens lens , – Käufel & Fuchs, Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 88/89: 201.

1960 Lindholmiola lens , – Zilch, Euthyneura, p. 693, Abb. 2419.

1961 Lindholmiola lens , – Jaeckel & Plate, Zoologische Abhandlungen, Abhandlungen and Berichte aus dem Staatlichen Museum für Tierkunde in Dresden, 26 (1): 3.

1962 Lindholmiola lens , – Klemm, Sitzungsberichte der Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abt. I, 171 (6/7): 252.

1976 Lindholmiola lens , – Frank, Mitteilungen der Zoologischen Gesellschaft Braunau, 2 (9/11): 261.

1986 Lindholmiola lens , – Gittenberger & Groh, Archiv für Molluskenkunde, 116 (4/6): 220. Abb. 1, a (= 2).

1986 Lindholmiola lens , – Reischütz, Malakologische Abhandlungen aus dem Staatlichen Museum für Tierkunde Dresden, 11 (9): 94.

1990 Lindholmiola lens , – Fechter & Falkner, Steinbachs Naturführer, p. 222 Abb. 1–3, 5.

1996 Lindholmiola lens , – Welter-Schultes, Schriften zur Malakozoologie, 9: 22, 31.

1996 Lindholmiola lens , – Dhora & Welter-Schultes, Schriften zur Malakozoologie, 9: 156, 190, Taf. 13, Fig. 167.

2002 Lindholmiola lens , – Dhora, Studime mbi molusqet e Shqipërisë: 99: 193.

Diagnosis: shell flat, lower whorls with a blunt to sharp keel, umbilicus cylindrical to broadly perspective, penial lumen with two longitudinal pilasters.

Description of shell: shell almost flat, shell colour light brown to yellowish brown; protoconch whorls smooth; initial whorls of the teleoconch with fine radial stripes, developing to small riblets on the subsequent whorls, but decreasing in height on the last teleoconch whorls; on the subsurface, radial riblets weak to almost missing; sculpture of teleoconch of fine granules; below the suture a small stripe devoid of hairs, the remaining shell including the umbilicus covered by usually impermanent hairs (often only the periphery of the shell with hairs); hairs yellowish-brown, 0.56–0.75 mm long; 5½–7¾ regularly increasing whorls; lower whorls usually flattened with a blunt to sharp keel (sometimes the whorls are more rounded forming an inconspicuous keel); last whorl only slightly descending, then shortly bent towards the aperture; suture moderately deep; umbilicus shape variable, either initially cylindrical and rapidly increasing with the last whorl, or initially broadly perspective and regularly increasing, reaching a diameter of 1.6–3.9 mm; aperture oblique, in lateral view slightly to strongly bent; peristomial insertions 2.2–5.9 mm apart, connected by an inconspicuous callus (except in callojuncta); peristome sharp to blunt, sometimes slightly reflected; columellar peristomial callus weak, covering the umbilicus ca 10%.

Measurements: H: 3.6–7; D: 7.2–15.6; aH: 2–3.9 (3–6.1); aW: 3.4–7.8.

Details of body (n = 12 from 5 populations: 1 specimen from 2 km N of Neohóri/1; 4 specimens from the pass at Haniá; 1 specimen from E of Glifáda; 2 specimens from 3 km S of Ptéri; and 4 specimens between Olymbía and Kréstena): head and dorsum grey to brown, more bright at the flanks and the posterior body; foot sole cream, sometimes darker at the fringes; mantle on the last 1½ whorls with small irregular dots; dots are more elongate between lung vein and suture and larger than those on the lower side of the mantle; intensity of pigmentation increasing towards the mantle collar, pigments often fuse to a large brown spot on the last whorl; ureter opens 1.5– 1 mm before the respiratory pore.

Morphology of the genital organs: penis relatively short, moderately thickened, glandular tissue covering one third to half of the total of the penial tube; penial lumen filled by two longitudinal pilasters of equal length starting as thin threads on the distal end of the penis and ending in the proximal third of the penis with thickened tissue; sometimes a few small folds at the distal end of the penis parallel to the major pilasters; vagina moderately long, usually thickened; glandula with a short stem and a long distal part with deep crossing furrows; bursa copulatrix as long as or somewhat longer than the glandula, the pedunculus reaches ca 60% of the total length of the organ, usually well separated from the vesicle; atrium with a broad and flat pilaster starting at the genital pore, often subdivided by deep longitudinal furrows; vagina filled by a single sometimes broadened pilaster.

Differential diagnosis: Its sisterspecies seems to be L. spectabilis , which has a shell that is similar to the form callojuncta of L. lens . It differs from this form by its more slowly increasing and less perspective umbilicus. Generally, the shell surface in L. spectabilis is more coarsely granulated, hairs are almost missing (if present, then below the shell's periphery), its aperture is narrower but higher, its lower rim is more wavy and larger, and its peristomial insertions are closer. In the penial lumen, L. spectabilis shows three longitudinal pilasters (instead of two in L. lens ); according to Fechter & Falkner (1990: 222), its pre-mating behaviour is shorter, and the lack of several behavioural elements proved to establish a mating barrier.

Broader specimens of L. girva are similar to that form of L. lens , which inhabits the Island Évvia and the easternmost parts of continental Greece (= lentiformis ). However, the shell of L. girva is less flattened, it has more rounded and less shouldered whorls; its radial riblets are less regular, and its aperture is more strongly bent in lateral view; its penis is longer, but the glandular tissue is less developed; the penial lumen is filled with 2–3 pilasters which are not broadened at their proximal end.

Shells of L. barbata are usually more rounded with a blunt keel. Shells of the same size differ by having an additional whorl, and the umbilicus is almost completely covered by the peristomial columellar reflection (for more details refer to L. barbata ).

Type specimens: Greece, Ilía (= Elis), Koúmanis (= Kumani), UTM EG 68, 37.78°N 21.74°E, leg. Brenske, 1883, syntypes lens var. elia : SMF 101961/6 , 225646/1 , 225647/2 . — Etoloakarnanía, Náfpaktos (= Lèpante , Lepanto), UTM EH 74, 38.39°N 21.82°E, syntype lens var. callojuncta : NHMG 372 .

Additional specimens examined:

Greece, Makedonía, Vérmio Mountains , about 6 km from the highway Véria – Kozáni direction Séli (= Áno Vérmio), conglomerate rocks at river bed, 530 m alt., UTM EK 98 , 40 .4971°N 22.1253°E, leg. Fauer & Subai 24.09.1989, S 15686 /1 ; gorge 2 km N of Neohóri (= NE of Aridéa ), on limestone rocks, 250 m alt., UTM EL 94 , 41 .0475°N 22.0916°E, leg. Subai 09.05.1995, S 15922 /13 + 1 (alk.) ; leg. Subai & Szekeres, 19.5.1997, S 15923 /5 ; 8.5 km NNE of Aridéa, on limestone rocks 3 roadkm. N of Vorinó , 250 m alt., UTM EL 94 , 41 .06°N 22.09°E, leg. Gittenberger & Uit de Weerd 9.5.2000, NNM/6 ; 9 km NNE of Aridéa, isolated rocks 6.3 roadkm. N of Vorinó , 440 m alt., UTM EL 94 , 41 .08°N 22.06°E, leg. Gittenberger & Uit de Weerd 9.5.2000, NNM/2 + 1 (juv.) ; in Véria, on loess wall beside the rivulet, 135 m alt., UTM FK 08 , 40 .5260°N 22.1997°E, leg. Fauer & Subai 24.09.1989, S 15925 /3 + 1 (juv.) ; forest in rocky gorge near Néa Pandeleimónas (= 18 km NW of Stómio), 100 m alt., UTM FK 32 , 40 .00°N 22.59°E, leg. Gittenberger & Uit de Weerd 24.5.1999, NNM/1 (juv.) ; Halkidikí Peninsula, 500 m N of Eleohória , on low limestone rocks, 170 m alt., UTM FK 86 , 40 .3434°N 23.1613°E, leg. Subai 05.05.1995, S 15684 /11 ; Halkidikí Peninsula, in valley beside the cave of Petrálona , 360 m alt., UTM FK 87 , 40 .3759°N 23.1681°E, leg. Subai 05.05.1995, S 15924 /24 ; leg. Subai 29.5.1997, S 15682 /2 ; rocky mountain slope 11.5 km ESE of Aridéa, NE border of Theodorákio , 450 m alt., UTM FL 03 , 40 .95°N 22.19°E, leg. Gittenberger & Uit de Weerd 10.5.2000, NNM/2 ; 500 m W of Gríva (= WSW of Axioúpoli ), on limestone rocks, 540 m alt., UTM FL 13 , 40 .9562°N 22.4009°E, leg. Subai & Szekeres 20.05.1997, S 15683 /4 ; N border of Asvestohóri (= E of Thessaloníki), on dry limestone rocks, 400 m alt., UTM FL 70 , 40 .6466°N 23.0236°E, leg. Subai & Szekeres 8.05.1997, S 15688 /1 ; between Rizá and Paleohóra (= 12.5 km W of Arnéa), 500 m alt., UTM GK 08 , 40 .50°N 23.44°E, leg. Gittenberger & Uit de Weerd 22.5.1999, NNM/1 (juv.) ; Kassándra Peninsula, limestone rocks near Loutrá, near sea, 20–25 m alt., UTM GK 22 , 39 .9235°N 23.5915°E, leg. Subai 05.05.1995, S 15681 /66 ; Sithonía Peninsula, mountain at S border of Porto Koufós , on limestone rocks in quarry, 30 m alt., UTM GK 42 , 39 .9576°N 23.9263°E, leg. Subai 06.05.1995, S 15631 /1 ; Sithonía Peninsula , mountain 1.5 km S of Porto Koufós, 10 m alt., UTM GK 42 , 39 .9494°N 23.9348°E, leg. Subai 05.05.1995, S 15629 /110. — Thessalía, E border of Theópetra (= 6 km SE of Kalambáka ), surrounding of church, UTM EJ 59 , 39 .67°N 21.68°E, leg. Fauer 21.8.1982, Maa /7, S 12218 /5 ; Theópetra (= Kuvelci , 6 km SSE of Kalambáka), on limestone blocks of rocks at river bank, 260 m alt., UTM EJ 59 , 39 .6798°N 21.6810°E, leg. E. & P. Subai 26.07.1975, S 3781/145, 3784/14 ; leg. Neuteboom 25.7.1979, NNM/2 ; leg. Gittenberger & Uit de Weerd 25.5.1999, NNM/16 + (alk.) ; Kalambáka, on conglomerate rocks beside the highway from Kalambáka to the Meteóra monasteries, 280 m alt., UTM EJ 59 , 39 .7103°N 21.6164°E, leg. Pintér, E. & P. Subai 29.07.1976, S 3785/5 ; N border of Morfovoúni , on E exposed limestone rocks, 840 m alt., UTM EJ 65 , 39 .3567°N 21.7464°E, leg. Subai & Szekeres 12.05.1997, S 15928 /2 + 1 (juv.) ; road border in Petrotó , 100 m alt., UTM EJ 87 , 39 .58°N 21.95°E, leg. Gittenberger & Uit de Weerd 25.5.1999, NNM/4 ; W border of Petrotó, on low limestone rocks, 120 m alt., UTM EJ 88 , 39 .5840°N 21.9584°E, leg. Subai, 18.05.1995, S 15691 /1 (juv.) ; 10 km E of Kefalóvrisso , 250 m alt., UTM EK 81 , 39 .92°N 21.98°E, leg. Gittenberger & Uit de Weerd 25.5.1999, NNM/9 ; between Elassóna and Deskáti (= 9.5 km W of Elassóna), 250 m alt., UTM EK 91 , 39 .88°N 22.08°E, leg. Gittenberger & Uit de Weerd 25.5.1999, NNM/2 ; 17 km W of Lárissa , bifurcation to Temkat, UTM FJ 08 , 39 .61°N 22.20°E, leg. Erőss 9.8.1993, S, 18258/1 ; between Tírnavos and Mesohóri , 8 km along road SE of Tírnavos, 125 m alt., UTM FJ 09 , 39 .71°N 22.24°E, leg. Gittenberger & Uit de Weerd 25.5.1999, NNM/19 + (alk.) ; road border 1 km W of Damássi , 150 m alt., UTM FJ 09 , 39 .71°N 22.16°E, leg. Gittenberger & Uit de Weerd 25.5.1999, NNM/1 ; between Tírnavos and Damássi , crossing to Tríkala, on low limestone rocks, 120 m alt., UTM FJ 09 , 39 .6991°N 22.2134°E, leg. Pintér, E. & P. Subai 28.07.1976, HNHM 85641 /1, S 3782/16 ; 1 km E of Fársala , UTM FJ 15 , 39 .28°N 22.40°E, leg. Kiss & Pintér 8.7.1986, HNHM 29905 /6 ; 37.5 km SSE of Lárissa, on limestone rocks 3.5 roadkm. bevor Fársala, 200 m N of road Domokós – Lárissa , 425 m alt., UTM FJ 15 , 39 .28°N 22.32°E, leg. Gittenberger & Uit de Weerd 21.5.2000, NNM/1 ; 2 km E of Fársala (= direction Ambelía), UTM FJ 25 , 39 .30°N 22.41°E, leg. Neuteboom 21.7.1979, NNM/2 ; Míra, N slope, on rocks, UTM FJ 25 , 39 .45°N 22.54°E, leg. Neuteboom 21.7.1979, NNM/1 ; Míra, path to Sofó, UTM FJ 36 , 39 .45°N 22.53°E, leg. Neuteboom 21.7.1979, NNM/12 ; 6 km SE of Almirós , W border of junction, 25 m alt., UTM FJ 53 , 39 .14°N 22.82°E, leg. Gittenberger & Uit de Weerd 29.5.1999, NNM/24 ; 3 km S of Aerinó , W rim of highway, 150 m alt., UTM FJ 55 , 39 .30°N 22.74°E, leg. Gittenberger & Uit de Weerd 29.5.1999, NNM/4 ; Néa Ionía (= Vólos, Volo), UTM FJ 65 , 39 .38°N 22.92°E, HNHM 39177 /3, SMF 101969 /5 ; leg. Stussiner, 1886, SMF 101964 /4 ; leg. C. R. Boettger, 1906, SMF 101978 /2 ; Néa Ionía , near of beach, UTM FJ 65 , 39 .34°N 22.92°E, leg. Schütt, 19.5.1962, SMF 279326 /7 ; road border 500 m E of Ahíllio (= Akhilion), 25 m alt., UTM FJ 71 , 39 .01°N 22.97°E, leg. Gittenberger & Uit de Weerd 29.5.1999, NNM/12 ; surroundings of Agía Kiriakí Village (= 32 km SE of Néa Ionía), 50–100 m alt., UTM FJ 72 , 39 .09°N 23.07°E, leg. Gittenberger & Uit de Weerd 28.5.1999, NNM/6 ; 1 km E of Agriá (= 5 km ESE of Néa Ionía ), 125 m alt., UTM FJ 75 , 39 .33°N 23.04°E, leg. Gittenberger & Uit de Weerd 26.5.1999, NNM/9 ; SE border of Néa Ionia (= Volos) W of the freight harbor, on rocks, among weeds, UTM FJ 75 , 39 .3492°N 22.9768°E, leg. Subai 30.04.1991, S 15926 /11 ; Makrinítsa at Pílio [Mountains], UTM FJ 76 , 39 .40°N 22.98°E, leg. Stussiner, 1887, SMF 284934 /3, 284955/1 ; 500 m SSW of Makrinítsa , 275 m alt., UTM FJ 76 , 39 .39°N 22.97°E, leg. Gittenberger & Uit de Weerd 27.5.1999, NNM/33 ; Pílio (= Pelion) [Mountains], UTM n. det., SMF 101870 /20 ; foot of Pílio [Mountains] (= Pílios), UTM n. det., leg. Stussiner, 1888, SMF 284933 /3 ; Pílio, 10 min. below peak, UTM FJ 76 , 39 .43°N 23.04°E, leg. Stussiner, 1888, SMF 284931 /4 ; Pílio, 1 hour below peak, UTM FJ 76 , 39 .42°N 23.05°E, leg. Stussiner, 1888, SMF 284932 /2 ; Survias monastery, N side of Pílio, UTM FJ 76 , 39 .4620°N 22.9847°E, leg. Stussiner, 1888, SMF 284935 /2 ; 4 km above Portaria (= direction Hánia), karst slope, 840 m alt., UTM FJ 76 , 39 .3704°N 23.0290°E, leg. Subai & Szekeres 9.05.1997, S 15929 /2 + 1 (juv.) + 1 (alk.) ; near Hánia (= ENE of Néa Ionia = Volos), in forest beside the Hotel "Hani Zisi", in leaf litter, 1150 m alt., UTM FJ 76 , 39 .3949°N 23.0619°E, leg. Subai & Szekeres 9.05.1997, S 15913 /2 (juv.) + 11 (alk.) ; road border at Kóttes (= 30.5 km SE of Néa Ionía), 70 m alt., UTM FJ 83 , 39 .11°N 23.09°E, leg. Gittenberger & Uit de Weerd 28.5.1999, NNM/7 ; 4 km WSW of Marathéa (= 32 km SE of Néa Ionía), 15 m alt., UTM FJ 83 , 39 .11°N 23.12°E, leg. Gittenberger & Uit de Weerd 28.5.1999, NNM/35 + 7 + (alk.) ; Olimbos (= Olympos) [Mountain], UTM n. det., leg. Maltzan, 1883, SMF 284954 /1 ; leg. Mahunka 15.9.1984, HNHM 23886 /14 + 8 (juv.) ; Témbi (= Tempe valley , approx. 30 km NE of Lárissa), on limestone rocks, UTM FK 31 , 39 .87°N 22.58°E, leg. Stussiner, 1884, SMF 284939 /4 ; Témbi (= Tempe valley , approx. 30 km NE of Lárissa), on limestone rocks and from the "Daphne" spring, UTM FK 31 , 39 .8796°N 22.5870°E, leg. Pintér, E. & P. Subai 28.07.1976, HNHM 39188 /12, S 3783/14 + 1 (juv.) ; leg. Neuteboom, 19.7.1979, NNM/2 + 3 ; leg. Maassen 7.1980, Maa /3 + 26 ; leg. Subai 24.7.1990, S 15927 /5 ; small valley along old road 500 m SW of Omólion (= Moli ), 50 m alt., UTM FK 31 , 39 .89°N 22.63°E, leg. Gittenberger & Uit de Weerd 26.5.1999, NNM/1 ; Ossa Mountains , Profítis Ilías peak region, on marble rocks, 1550 m alt., UTM FK 40 , 39 .7948°N 22.6710°E, leg. Subai 23.07.1990, S 15930 /4 + 1 (juv.) ; Ossa Mountains, Bugari Valley , UTM n. det., leg. Stussiner, leg., SMF 284938 /2 ; Thessaloníki (= Saloniki), UTM n. det., leg. Haussknecht, 1885, SMF 284941 /1 ; Langadás near Thessaloníki , UTM FL 71 , 40 .74°N 23.06°E, SMF 207803 /9. — Fthiótida, plane between Brálos and Parnassós [Mountains], UTM FH 28, 38.69°N 22.45°E, leg. Pfeiffer, 1926, SMF 101876 /6 ; Brálos – Loutrá Thermopilón highway, 8 km from bifurcation Moni Damasta direction Loutrá Thermopilón , on rocks, 110 m alt., UTM FH 29 , 38 .8008°N 22.4973°E, leg. Subai 10.04.1986, S 15795 /1 ; entrance of gorge at SE border of Amfíklia , in the vicinity of a small chapel, on limestone rocks, 500 m alt., UTM FH 37 , 38 .6306°N 22.5963°E, leg. Subai 10.04.1986, S 15696 /2 ; Loutrá Thermopilón , monument, UTM FH 39 , 38 .79°N 22.53°E, leg. Kiss & Pintér 9.7.1986, HNHM 29917 /1 ; gorge W of Tithoréa and in the big cave, 850 m alt., UTM FH 47 , 38 .5702°N 22.6602°E, leg. Subai 11.04.1986, S 15694 /4 ; SW border of Atalánti , on limestone rocks, 190 m alt., UTM FH 77 , 38 .6534°N 22.9849°E, leg. Subai 11.04.1986, S 15695 /3 ; Atalánti (= Atalandi ), UTM FH 78 , 38 .65°N 22.99°E, SMF 284950 /4 ; leg. Thiesse, 1890, SMF 284897 /6, 284937/6 ; leg. Clessin, 1879, SMF 101859 /1 ; Ipáti, on limestone rocks beside the waterfall, 380 m alt., UTM FJ 00, 38.8710°N 22.2440°E, leg. Subai 16.05.1991, S 15711 /2 ; 1.5 km NE of Lamía (= highway direction Lárissa), stone mine beside Xerias river , on limestone rocks, 615 m alt., UTM FJ 20 , 38 .9673°N 22.3874°E, leg. Subai 10.04.1986, S 15697 /6 ; 9 km from Glífa direction Pelasgía , on limestone rocks, 100 m alt., UTM FJ 61 , 38 .9309°N 22.9229°E, leg. Subai 30.04.1991, S 15706 /19. — Etoloakarnanía, Veloútsa Mountain SE slope above Astakós , on limestone rocks, approx., 185 m alt., UTM EH 06 , 38 .5370°N 21.0692°E, leg. Subai, 19.05.1991, S 15715 /3 ; Messolóngi , UTM EH 34 , 38 .37°N 21.43°E, leg. Krüper, 1901, SMF 101866 /4 ; above Kefalóvrisso , on limestone rocks, 30 m alt., UTM EH 35 , 38 .4598°N 21.3670°E, leg. Fauer & Subai 30.09.1989, S 15736 /18 ; ruins of Ancient Kalydon (= 2 km from Evinos bridge direction Messolóngi ), under stones, UTM EH 44 , 38 .3725°N 21.5332°E, leg. Fauer & Subai 01.10.1989, S 15732 /42 ; Krionéri , on limestone rocks on Varássova Mountain, SW slope, 50–220 m alt., UTM EH 54 , 38 .3477°N 21.5973°E, leg. Fauer & Subai 01.10.1989, S 15731 /5 ; in Riza village, on limestone rocks and mountain slope W of Riza, beside the old road to Messolóngi , 70 m alt., UTM EH 64 , 38 .3541°N 21.6978°E, leg. Fauer & Subai 02.10.1989, S 15735 /13 ; Náfpaktos (= Lepanto ), UTM EH 74 , 38 .39°N 21.82°E, SMF 285762 /7 ; leg. Krüper SMF 101895 /23, 101898/6 ; leg. Thiesse, 1890, SMF 284956 /6 ; on limestone rocks in Náfpaktos, 20 m alt., UTM EH 74 , 38 .3929°N 21.8259°E, leg. Pintér, E. & P. Subai 31.07.1976, HNHM 39184 /2 + 3 (juv.), S 3799/3. — Fokída, mountain W of Glifáda (= opposite of Trizónia Island ), on limestone rocks, 100 m alt., UTM EH 94 , 38 .3883°N 22.0711°E, leg. Subai 14.04.1986, S 15720 /5 ; leg. Kiss & Pintér 23.7.1986, HNHM 29900 /8 + 6 (juv.) ; leg. Riedel & Subai 15.4.1988, S 15739 /36 ; border of Ágios Spirído Village (= E of Glifáda), on dry limestone rocks, 30 m alt., UTM EH 94 , 38 .3679°N 22.1169°E, leg. Fauer & Subai 01.10.1989, S 15734 /16 ; 6 km from Erátini direction Náfpaktos , on rocks at sea shore, 40 m alt., UTM FH 04 , 38 .3316°N 22.1907°E, leg. Subai 14.04.1986, S 15723 /5 ; 4 km S of Malandrino (= S of Lidoríki ), on limestone rocks, 550 m alt., UTM FH 05 , 38 .4178°N 22.2498°E, leg. Subai 13.4.1986, S 15730 /7 ; approx. 200 m from the crossing Lidoríki–Erátini–Amfissa direction Amfissa , on limestone rocks, 640 m alt., UTM FH 15 , 38 .4052°N 22.2550°E, leg. Fauer & Subai 01.10.1989, S 15733 /8 ; 9 km from Mórnou dam direction Lévka, W side of a peninsula on S banks of the dam reservoir, on limestone rocks, 550 m alt., UTM FH 06 , 38 .5342°N 22.1634°E, leg. Subai 13.04.1986, S 15729 /3 ; 3 km from Penteória direction Erátini , on limestone rocks, 750 m alt., UTM FH 15 , 38 .4155°N 22.2967°E, leg. Subai 14.04.1986, S 15722 /8 ; Lidorikíou Mountains, 28 km from Amfissa direction Lidoríki (= 19 km S of Lidoríki), 600 m alt., UTM FH 15 , 38 .3992°N 22.2696°E, leg. Subai 13.04.1986, S 15728 /2 ; 7.5 km from Itéa direction Galaxídi , UTM FH 25 , 38 .42°N 22.39°E, leg. Neuteboom 22.7.1979, NNM/8 + 1 (juv.) ; 2 km S of Ámfissa (= direction Vouníhora), on limestone rocks, 300 m alt., UTM FH 26 , 38 .5087°N 22.3821°E, leg. Subai 16.05.1991, S 15718 /16 ; Eptálofos (= Agóriani), Parnassós Mountains , UTM FH 27 , 38 .59°N 22.49°E, SMF 284867 /3 ; leg. Krüper, 1888, SMF 101984 /4 ; leg. Krüper, 1899 and, 1901, SMF 101923 /8 ; leg. Schlüter, 1888, SMF 284864 /1 ; 2.5 km of Graviá direction Amfissa , on limestone rocks, 540 m alt., UTM FH 27 , 38 .6607°N 22.4253°E, leg. Subai 12.04.1986, S 15737 /4 ; E-exponated rocks at S border of Marioláda , 450 m alt., UTM FH 27 , 38 .65°N 22.46°E, leg. Gittenberger & Uit de Weerd , 19.5.2000, NNM/18 ; 2 km SE von Marioláda , an Kalkfelsen, 780 m alt., UTM FH 27 , leg. Gittenberger & Uit de Weerd , 19.5.2000, NNM/5 ; E border of Desfína, UTM FH 35 , 38 .41°N 22.53°E, leg. Hoenselaar 27.5.1996, Maa /2 ; NE of Delfí, ruins of ancient Delphi , 650 m alt., UTM FH 36 , 38 .4824°N 22.4981°E, SMF 207815 /9 ; leg. Müller, SMF 285174 /1 ; leg. Krüper, SMF 101896 /7 ; leg. Pfeiffer, 1926, SMF 101892 /3, 101893/10, 101894/1 ; leg. Schütt 4.6.1969, S 3798/3 ; leg. Neuteboom 22.7.1979, NNM/7 + 3 (juv.) ; leg. Drimmer 23.5.1982, HNHM 33703 /5 ; leg. Podani 30.7.1984, HNHM 21467 /3 ; leg. Neuteboom 26.10.1984, NNM/3 ; leg. Perjesi 29.4.1985, HNHM 23902 /1 ; leg. Kiss & Pintér 23.7.1986, HNHM 29895 /18 + 2 (juv.) ; leg. Sín 6.7.1987, HNHM 44341 /5 + 3 (juv.) ; leg. Erőss 6.4.1989, S, 18259/3 ; leg. Subai 16.5.1991, S 15719 /35 ; leg. Hoenselaar 26–27.5.1996, Maa /1 + 2 ; 2.5 km E of Amfíklia, UTM FH 37 , 38 .64°N 22.62°E, leg. Neuteboom 25.10.1984, NNM/1 + 2 (juv.) ; Polídrosos , UTM FH 37 , 38 .63°N 22.53°E, leg. Neuteboom 25.10.1984, NNM/1 ; Parnassós Mountains (= Parnass ), UTM n. det., SMF 101858 /7, 101882/1 ; leg. Krüper, 1883, SMF 101872 /3 ; leg. Schmidt, 1935, SMF 101878 /3 ; Parnassós Mountains , tree-line 1.7 km of ski-lift, UTM FH 46 , 38 .55°N 22.57°E, leg. Neuteboom 23.7.1979, NNM/2 + 3 (alk.). — Viotía, Tríodos, bifurcation after Dístomo , UTM FH 45 , 38 .44°N 22.68°E, leg. Neuteboom 23.7.1979, NNM / 25 km from the crossing Dístomo – Delfí–Livadiá direction Livadiá , on limestone rocks, 310 m alt., UTM FH 45 , 38 .4627°N 22.7161°E, leg. Subai 14.04.1986, S 15794 /1 ; Elikón Mountains, 6 km from Kiriáki direction Dístomo and 4 km southward on a small path, upper border of a gorge, on limestone rocks, 100 m alt., UTM FH 54 , 38 .3625°N 22.7208°E, leg. Subai 14.04.1986, S 15721 /1 ; 3 km from Paralía Sarandi direction Pródromos , on limestone rocks and in debris among rocks, 220 m alt., UTM FH 63 , 38 .2572°N 22.8756°E, leg. Subai 15.04.1986, S 15725 /14 ; 4.3 km from Paralía Sarandi direction Pródromos , on limestone rocks and in an olive tree plantation, 225 m alt., UTM FH 63 , 38 .2578°N 22.8826°E, leg. Subai 15.04.1986, S 15724 /5 ; rock gorge at SE border of Livadiá , on limestone rocks and in organic flotsam of creek, 220 m alt., UTM FH 65 , 38 .4329°N 22.8920°E, leg. Subai 15.04.1986, S 15796 /13 ; Lafísti (= Lofistion , E of Livadiá), on limestone rocks, 310 m alt., UTM FH 65 , 38 .4281°N 22.9189°E, leg. Subai 11.04.1986, S 15799 /3 ; Herónia , ancient theatre 6 km NW of Livadiá, 250 m alt., UTM FH 66 , 38 .49°N 22.84°E, leg. Gittenberger & Uit de Weerd , 19.5.2000, NNM/1 ; 2 km E of Alíartos , on limestone rocks, approx. 80 m beside highway, 115 m alt., UTM FH 84 , 38 .4351°N 23.1997°E, leg. Riedel & Subai 09.04.1988, S 15761 /1 + 3 (juv.) ; 13 km W of Thíva, 9.9 roadkm. of Alíartos additional 2 km on a secondary road, 250 m alt., UTM FH 94 , 38 .33°N 23.14°E, leg. Gittenberger & Uit de Weerd 20.5.2000, NNM/5 ; on limestone rocks beside the motor way, at the bifurcation to Akréfnio (= 5.5 km SE of the crossing near Kástro ), 120 m alt., UTM FH 95 , 38 .4500°N 23.2092°E, leg. Subai & Szekeres 11.05.1997, S 15813 /6 ; 7 km N of Thíva, on rocks 1 roadkm. along a secondary road to Movríki , UTM GH 05 , 38 .38°N 23.32°E, leg. Gittenberger & Uit de Weerd 20.5.2000, NNM/2 + 1 (juv.) ; Skoponéri Village , about 50 m S of the tavern, on low limestone rocks, approx. 10 m alt., UTM GH 06 , 38 .5032°N 23.3549°E, leg. Subai & Szekeres 11.05.1997, S 15690 /47 + 2 (juv.) + 9 (alk.) ; above Skoponéri , E border of the village, on limestone rocks, 270 m alt., UTM GH 06 , 38 .5013°N 23.3675°E, leg. Subai & Szekeres 11.05.1997, S 15689 /8 + 1 (juv.) ; mountain slope 10 km from Halkída (= Chalkis) direction Thiva (= Thebe ), on limestone rocks, 270 m alt., UTM GH 25 , 38 .4282°N 23.5309°E, leg. Riedel & Subai 09.04.1988, S 15698 /7. — Atikí (= Attika), UTM n. det., SMF 284881 /7, 284893/2, 284894/1, 284980/3 ; leg. Krüper, 1883, SMF 101861 /8 ; approx. 8.5 km from Paralía Shínou direction Káto Alepohóri, on limestone rocks, approx. 10 m alt., UTM FH 81 , 38 .0514°N 23.0571°E, leg. Subai 14.05.1991, S 15712 /7 + 1 (juv.) ; Aigósthena (= Porto Germeno), on rocks below the castle ruins, 45 m alt., UTM FH 92 , 38 .1483°N 23.2321°E, leg. Subai 16.05.1991, S 15693 /2 (fragment) ; 11 km from the highway Vília–Aigósthena direction Kithairón peak, on limestone rocks, 1300 m alt., UTM FH 92 , 38 .1835°N 23.2583°E, leg. Subai 16.05.1991, S 15915 /7 + 1 (juv.) ; Vouliagméni (= opposite of Fléves Island ), on limestone rocks beside the highway, 20 m alt., UTM GG 48 , 37 .8086°N 23.7841°E, leg. Subai, 19.07.1979, S 15701 /51 ; 3.4 km SE of Varkíza (= coast road direction Agía Marína ), on limestone rocks, 30 m alt., UTM GG 48 , 37 .8127°N 23.8314°E, leg. Subai 07.05.1991, S 15705 /1 (fragment) ; "Varkíza Beach" camping near Varkíza , on limestone rocks, 30 m alt., UTM GG 48 , 37 .8226°N 23.8170°E, leg. Subai 20.07.1979, S 6644/10 ; 3 km from Vári direction Koropí , on limestone rocks, 50 m alt., UTM GG 49 , 37 .8415°N 23.8215°E, leg. Subai 20.07.1979, S 6700/2 ; coastal strip near Paleá Fókaia , UTM GG 57 , 37 .71°N 23.94°E, leg. Neuteboom 20.10.1984, NNM/1 ; 3 km N of Legrená , UTM GG 67 , 37 .69°N 24.00°E, leg. Neuteboom 20.10.1984, NNM/3 ; coast near Megara , UTM GH 00, 37.97°N 23.32°E, leg. Pfeiffer, 1926, SMF 101844 /1 ; Sborder of Megara , UTM GH 00, 37.97°N 23.35°E, leg. Kiss & Pintér, 19.7.1986, HNHM 29918 /2 + 1 (juv.) ; mountain slope at W border of Mégara , on limestone rocks, 65 m alt., 37.9962°N 23.3240°E, UTM GH 00, leg. Subai 07.07.1979, S 6515/2 + 2 (juv.) ; mountain slope beside Ágios Ierothéou monastery (= W of Mégara), 300 m alt., UTM GH 01 , 38 .093N, 23.2677°E, leg. Subai 07.07.1979, S 15702 /11 + 1 (juv.) ; 3.5 km from the highway Vília–Aigósthena direction Kithairón peak, rocks in coniferous forest, 850 m alt., UTM GH 02 , 38 .1706°N 23.2728°E, leg. Subai 16.05.1991, S 15692 /2 (fragment) ; Elefsína–Thíva country road, mountain behind Mándra , UTM GH 11 , 38 .08°N 23.48°E, leg. Pfeiffer, 1926, SMF 101875 /4 ; Pérama, on rocks at coastal strip direction Piráeus , UTM GH 20 , 37 .96°N 23.60°E, leg. Neuteboom 4.9.1982, NNM/1 ; Elefsína, dry mountain slope beside a quarry, 130 m alt., UTM GH 21 , 38 .0508°N 23.4950°E, leg. Subai 07.07.1979, S 6559/1 ; Athína (= Athen ), UTM n. det., SMF 284878 /9, 284879/2, 284946/2 ; leg. Krüper, 1883, SMF 101863 /5, 101972/11 ; leg. Heldreich, SMF 284882 /3 ; Piráeus, UTM GH 30 , 37 .94°N 23.63°E, leg. Krüper, 1883, SMF 101983 /3 ; Athína, Paleo Falíro (= Phaleron, part of Athína ), UTM GH 30 , 37 .92°N 23.70°E, leg. Krüper, SMF 101975 /39 ; Athína , Akropolis, UTM GH 30 , 37 .97°N 23.72°E, leg. Pfeiffer, 1926, SMF 101843 /24 ; Párnis Mountain (= Parnés) Chasia , UTM n. det., leg. Krüper, 1883, SMF 101971 /1 ; W side of Párnitha Mountain (= N of Athína), river valley along the circular path on top of the mountain, on limestone rocks, 1000 m alt., UTM GH 32 , 38 .1631°N 23.7020°E, leg. Subai 21.04.1988, S 15916 /13 ; SW border of Avlóna , on limestone rocks, 300 m alt., UTM GH 33 , 38 .2454°N 23.6855°E, leg. Subai 16.04.1986, S 15801 /2 ; leg. Riedel & Subai 9.4.1988, S 15804 /15 ; leg. Gittenberger & Uit de Weerd 20.5.2000, NNM/13 ; Likavittós (= Lykabettos) Mountain in Athína , on limestone rocks, UTM GH 40 , 37 .98°N 23.74°E, SMF 101874 /2, 285169/24, 285170/2 ; leg. Krüper, 1883, SMF 101860 /1 ; leg. Heldreich, SMF 284892 /3 ; leg. Coneménos, 1889, SMF 284886 /9 ; leg. Krüper, 1888, SMF 101985 /9 ; leg. Pintér 21.8.1976, HNHM 39181 /2 + 1 (juv.) ; leg. Perjesi 22.4.1985, HNHM 23890 /1 ; top of Likavittós Mountain in Athína , on limestone rocks, 250 m alt., UTM GH 40 , 37 .9826°N 23.7436°E, leg. Subai, 19.7.1979, S 6249/12 ; leg. Subai 22.04.1988, S 15704 /2 ; Hyméttos Mountain , UTM GH 40 , 37 .96°N 23.81°E, leg. Schmidt, SMF 285168 /2 ; leg. Köhler, SMF 285167 /2 ; leg. Knobbe, SMF 284885 /2 ; leg. Brenske, 1883, SMF 101963 /2, 284888/2 ; leg. Krüper, 1883, SMF 101862 /6 ; leg. Krüper, 1899, SMF 101967 /10 ; leg. Pfeiffer, 1926, SMF 101954 /1 ; N side of mount Hyméttos , on limestone rocks, approx. 800 m alt., UTM GH 40 , 37 .9604°N 23.8162°E, leg. Subai 4.4.1986, S 15713 /5 + 1 (juv.) 15714/100 + 3 (juv.) ; leg. Subai 21.4.1988, S 15710 /3 ; Kifissiá (= Kephissia ), UTM GH 41 , 38 .07°N 23.82°E, leg. Pfeiffer, 1926, SMF 101886/3 ; mountain SW of Anatolí (= W of Néa Mákri ), on limestone rocks, approx. 200 m alt., UTM GH 51 , 38 .0895°N 23.9468°E, leg. Subai 02.05.1991, S 15808 /14 ; Pendéli (= Pentelikon) [Mountains], UTM n. det., SMF 207814 /2 ; leg. Krüper, SMF 101974 /40 ; leg. Pfeiffer, 1926, SMF 101888 /1 ; Pendéli monastery, UTM GH 51 , 38 .05°N 23.86°E, leg. Pfeiffer, 1926, SMF 101887 /1 ; approx. 4 km NE of Pendéli, Pendéli (= Pentelikon) Mountains W side, on crystalline rocks, 700 m alt., UTM GH 51 , 38 .0653°N 23.9081°E, leg. Subai 02.05.1991, S 15809 /12 ; Soúnio , UTM KB 37 , 37 .65°N 24.02°E, leg. Pfeiffer, 1926, SMF 101885 /29 ; 4 km from Lavrio direction Markópoulo, on limestone rocks, 125 m alt., UTM KB 38 , 37 .7523°N 24.0260°E, leg. Subai 05.05.1991, S 15703 /15 ; road Athína–Soúnio , before Lavrio, UTM KB 48 , 37 .74°N 24.05°E, KB 38, leg. Pfeiffer, 1926, SMF 101884 /5 ; road Athína – Marathónas, UTM n. det., leg. Pfeiffer, 1926, SMF 101890 /9 ; road Athína – Soúnio, UTM n. det., leg. Pfeiffer, 1926, SMF 101883 /7. — Égina Island , UTM n. det., SMF 207811 /2 ; Mesagrós, Aphaia holy place, UTM GG 28 , 37 .75°N 23.53°E, leg. Erőss 5.4.1989, S, 18261/10. — Méthana Island , Méthana Town, UTM GG 25 , 37 .58°N 23.39°E, leg. Haussknecht, 1885, SMF 284887 /2. — Kíthira Island (= Cerigo), "Keramoto", UTM n. det., leg. Krüper, 1893, SMF 101869 /8, 284913/2. — Kefaloniá Island , UTM n. det., HNHM 39171 /1, 39175/2, 39179/2, SMF 101857 /6, 101881/4, 84910/9, 284911/1 ; leg. Müller, SMF 285162 /2 ; Argostóli Peninsula, UTM DH 52 , 38 .18°N 20.48°E, leg. Visser 1.11.1985, NNM/1 (alk.). — Zakinthos Island (= Zante), UTM n. det., SMF 284884 /5 ; above Chiliomenon, UTM n. det., HNHM 39236 /3 + 1 (juv.). — Ilía (= Elis), theatre 15 km E of Gastoúni , UTM EG 39 , 37 .89°N 21.37°E, leg. Hoenselaar 17.4.1996, Maa /5 ; Kaíafás (= NW of Zaháro), on limestone rocks and from the thermal well, UTM EG 55 , 37 .5193°N 21.6022°E, leg. Subai 07.04.1986, S 15798 /11 ; ruins of Arhea Olympia , 35 m alt., UTM EG 56 , 37 .6384°N 21.6307°E, HNHM, 18649/1; SMF 279327 /4, 284900/10, 285173/6 ; leg. Schlüter, SMF 284901 /15 ; leg. Pfeiffer, 1926, SMF 101851 /7 ; leg. Kühnel 4.1933, HNHM 9784 /2 ; leg. Hansen, 1937, HNHM 39187 /2 ; leg. Jaeckel, 1958, SMF 207816 /10 ; leg. Seidl 3.10.1965, NNM/4 ; leg. Guyat & Dawber 8.1973, HNHM 85645 /1, S 3793/12 ; leg. Pintér & Subai 2.8.1976, S 3494/61 ; leg. Sin 4–5.7.1985, HNHM 24029 /17 + 3 (juv.), 24069/25 + 4 (juv.) ; leg. Kiss & Pintér 23.7.1986, HNHM 29897 /9 + 3 (juv.) ; leg. Hoenselaar, 19.4.1996, Maa /7 ; between Arhea Olympia and Kréstena, beside the channel of Alfiós River, in grass and under stones, UTM EG 56 , 37 .6279°N 21.5923°E, leg. Subai 10.07.1979, S 15882 /36 + 17 (alk.) ; Giannitsohóri , Camping "Apollo", UTM EG 63, 37.39°N 21.69°E, leg. Hoenselaar 21–23.4.1996, Maa /2 + 3 ; river gorge approx. 1.5 km E of Néa Figália, on limestone rocks, approx. 550 m alt., UTM EG 64 , 37 .4469°N 21.7890°E, leg. Riedel & Subai, 18.04.1988, S 15762 /1 ; road Tholon– Andrítsena , border of Lépreo, UTM EG 64 , 37 .43°N 21.72°E, leg. Hoenselaar 22.4.1996, Maa /2 ; Koúmanis (= Kumani), UTM EG 68 , 37 .79°N 21.74°E, SMF 225648 /3 ; river gorge between the two bifurcations of the Lámbia – Tripótama highway to Orini , on limestone rocks, 630 m alt., UTM EG 79 , 37 .8608°N 21.8612°E, leg. Riedel & Subai 17.04.1988, S 15760 /9 ; leg. Hadjicharalambous & Subai 12.5.1991, S 15756 /1. — Ahaïa , 4.1 km from the crossing Kalávrita–Trípoli–Tripótama direction Tripótama, on limestone rocks, 1120 m alt., UTM EG 99 , 37 .8992°N 22.0087°E, leg. Hadjicharalambous & Subai 14.05.1991, S 15716 /5 ; dry meadow 2 km S of Klitoria (= direction Trípoli ), 500 m alt., UTM EG 99 , 37 .8808°N 22.1355°E, leg. Subai 09.07.1979, S 15738 /3 ; Araxos , on limestone rocks beside the stone mine at banks of Lake Kalogera, 20 m alt., UTM EH 32, 38.1642°N 21.4011°E, leg. Subai 07.04.1986, S 15727 /9 ; Skollis Mountain W slope above Sandoméri, on limestone rocks, in rock debris, 550 m alt., UTM EH 40 , 37 .9899°N 21.5781°E, leg. Riedel & Subai 16.04.1988, S 15766 /4 ; W slope of Skollis Mountain , NE of Sandoméri, on limestone rocks, 530 m alt., UTM EH 50 , 37 .9929°N 21.5778°E, leg. Subai 07.04.1986, S 15797 /9 ; Erímanthos Mountains above Kaléntzi, on limestone rocks, 1060 m alt., UTM EH 60 , 37 .9447°N 21.7650°E, leg. Hadjicharalambous & Subai 13.05.1991, S 15921 /4 ; Pátra (= Patras), UTM EH 63 , 38 .24°N 21.74°E, SMF 284890 /2, 284908/5, 284953/4 ; leg. Coneménos, SMF 101854 /5 ; leg. Brenske, 1883, SMF 101968 /6 ; leg. Krüper, 1895, SMF 284960 /6 ; leg. Krüper, 1899, SMF 101952 /11 ; leg. C. R. Boettger, 1905, SMF 101979 /5 ; Psathópirgos, UTM EH 74 , 38 .32°N 21.87°E, SMF 225650 /19 + 20 ; river valley near Psathopírgos, among dry plants and on rocks (no limestone!), 230 m alt., UTM EH 74 , 38 .3183°N 21.8823°E, leg. Subai 07.04.1986, S 15726 /5 ; in a dry river bed S of Flámpoura village (= Metohi ), 900 m alt., UTM EH 81 , 38 .0286°N 21.9965°E, leg. Pintér, E. & P. Subai 01.08.1976, S 3791/8, HNHM 39170 /5 ; bifurcation of the highway Kalávrita – Diakoftó to Rogí, Kerpiní, on limestone rocks, 700 m alt., UTM EH 91 , 38 .0534°N 22.1384°E, leg. Hadjicharalambous & Subai 13.05.1991, S 15717 /3 ; river gorge 3 km S from Ptéri , on limestone rocks and under stones, 1150 m alt., UTM EH 92 , 38 .1418°N 22.0710°E, leg. Riedel & Subai 15.04.1988, S 15920 /3 + 1 (juv.) + 2 (alk.) ; leg. Fauer & Subai 3.10.1989, S 15919 /9 ; leg. Hadjicharalambous & Subai 13.05.1991, S 15780 /1 ; in and around the cave N of Kastría , on limestone rocks, 840 m alt., UTM FH 00, 37.9603°N 22.1404°E, UTM FH 00, leg. Hadjicharalambous & Subai 14.05.1991, S 15778 /2 ; on rocks and on walls of Mega Spileo monastery (= N of Kalávrita), 950 m alt., UTM FH 01 , 38 .0884°N 22.1746°E, leg. Pintér, E. & P. Subai 01.08.1976, S 3792/3 ; Subai 9.7.1979, S 6577/1 ; in "Lemon Beach" camping near Paralia Akráta, under stones, UTM FH 12 , 38 .1653°N 22.3408°E, leg. Subai 09.07.1979, S/2 (alk.) ; Nezerá , UTM n. det., leg. Coneménos, SMF 225649 /2. — Korinthía, Killini Mountains , in spruce forest above Goúra, on limestone rocks, 1100 m alt., UTM FG, 19, 37.9347°N 22.3488°E, leg. Subai 08.07.1979, S 6428/1 + 2 (juv.) ; Killini Mountains , 7.5 km from Goúra direction Kastanéa, on limestone rocks, 800 m alt., UTM FG, 19, 37.8866°N 22.3408°E, leg. Subai 08.07.1979, S 6573/2 ; Oligirtos Mountains , 1 km from Kartéri direction Goúra, on limestone rocks and in cave crevices, 850 m alt., UTM FG 29 , 37 .8510°N 22.3996°E, leg. Subai 08.07.1979, S 15750 /2 ; Killini Mountains , mountain slope 6 km from Kaliáni direction Kiáto, on conglomerate rocks, 710 m alt., UTM FG 39 , 37 .9236°N 22.5262°E, leg. Subai 08.07.1979, S 6661/2 ; 1 km S of Neméa , on limestone rocks, 370 m alt., UTM FG 48 , 37 .8024°N 22.6670°E, leg. Hadjicharalambous & Subai 07.05.1991, S 15781 /2 ; ancient Neméa (= near Arhéa Neméa), UTM FG 58 , 38 .82°N 22.66°E, leg. Hoenselaar 22.5.1996, Maa /5 ; 8.5 km from Ágios Vassílios direction Árgos, on limestone rocks, 290 m alt., UTM FG 58 , 37 .7779°N 22.7339°E, leg. Subai, 18.07.1979, S 6561/2 + 1(juv.) ; ruins of Arhea Kórinthos , 80 m alt., UTM FG 69 , 37 .9059°N 22.8792°E, SMF 284951 /6 ; leg. Brenske, 1883, SMF 284920 /1 ; leg. Krüper, SMF 101853 /5 ; leg. Pfeiffer, 1926, SMF 101845 /6 ; leg. Pintér & Subai 6.08.1976, HNHM 39172 /4, S 3795/4 ; leg. Nosek 3.10.1976, HNHM 21461 /2 ; leg. Hoenselaar 23.5.1996, Maa /2 ; road to Akrokorinthos, UTM FG 69 , 37 .89°N 22.87°E, leg. Pfeiffer, 1926, SMF 101846 /1 + 4 (juv.) ; leg. Neuteboom 29.8.1982, NNM/1 ; ruins of the old castle of Akrokorinthos, 400 m alt., UTM FG 69 , 37 .8900°N 22.8690°E, SMF 285172 /2 ; leg. Schlüter 27.4.1912, SMF 285765 /8 ; Tomlin, 1904, SMF 284889 /1 ; leg. Subai, 18.7.1979, S 15744 /57 + 1 (alk.) ; leg. Kiss & Pintér 21.8.1988, HNHM 44600 /24 +, 18 (juv.) ; leg. Neuteboom 29.8.1982, NNM/11 + 3 (juv.) ; highway Epidauros – Kórinthos, 8 km from the bifurcation to Sofikó direction Epidauros, 230 m alt., UTM FG 87 , 37 .7334°N 23.1010°E, leg. Pintér, E. & P. Subai 06.08.1976, HNHM 39182 /2 + 1 (juv.), S 3796/2 ; Kórfos , UTM FG 88 , 37 .76°N 23.12°E, leg. Kiss & Pintér 16.8.1988, HNHM 85644 /7 ; 2 km S of Kórfos , UTM FG 88 , 37 .73°N 23.12°E, leg. Kiss & Pintér, 19.7.1986, HNHM 29909 /1 (juv.) ; N exposed lime hill slope between Ano Loutró and Kariá, 550 m alt., UTM FH 31 , 38 .0842°N 22.5403°E, leg. Fauer & Subai 03.10.1989, S 15740 /1 ; ruins of Heraion and on limestone rocks beside Lake Heraiou , 60 m alt., UTM FH 61 , 38 .0349°N 22.8708°E, leg. Subai, 18.07.1979, S 6352/13 + 3 (alk.) ; Loutráki, UTM FH 70 , 37 .98°N 22.97°E, leg. Krüper, SMF 101973 /10 ; leg. Butot & Subai, 19.2.1981, S 17540 /12 + 3 (alk.) ; leg. Kiss & Pintér, 19.7.1986, HNHM 29901 /18 + 7 (juv.) ; leg. Erőss 22.7.1992, S, 18257/1 ; dry mountain slope at NW border of Loutráki, on limestone rocks, 80 m alt., UTM FH 70 , 37 .9836°N 22.9715°E, leg. Subai 07.07.1979, S 15751 /15 ; W border of Perahóra , UTM FH 71 , 38 .02°N 22.94°E, leg. Kiss & Pintér 16.8.1988, HNHM 44685 /15 + 6 (juv.) ; 1 km S of Perahóra , UTM FH 71 , 38 .01°N 22.94°E, leg. Kiss & Pintér 16.8.1988, HNHM 44589 /6 + 1 (juv.) ; approx. 1.5 km from Paralía Shínou direction Káto Alepohóri, on limestone rocks, approx. 100 m alt., UTM FH 81 , 38 .0570°N 23.0055°E, leg. Subai 14.05.1991, S 15914 /10. — Arkadía, 1 km from Stavrodrómio direction Arhea Olympia (= highway Arhea Olympia–Trípoli), 630 m alt., UTM EG 87 , 37 .6989°N 21.9392°E, leg. Pintér, E. & P. Subai 02.08.1976, S 3775/10 ; 2 km from Langádia direction Arhea Olympia (= highway Arhea Olympia–Trípoli), 850 m alt., UTM EG 87 , 37 .6815°N 22.0147°E, leg. Pintér, E. & P. Subai 02.08.1976, S 3777/1 ; 3 km from Kalonério direction Arhea Olympia (= highway Arhea Olympia–Trípoli), 1090 m alt., UTM EG 97 , 37 .6727°N 22.0534°E, leg. Pintér, E. & P. Subai 02.08.1976, S 3776/2 ; lime mountain N side, 6 km from Trópea direction Dáfni, approx. 500 m alt., UTM EG 87 , 37 .7573°N 21.9626°E, leg. Riedel & Subai 17.04.1988, S 15759 /1 ; 2 km from Paradísia direction Megalópoli (= highway Megalópoli – Kalamáta ), 475 m alt., UTM EG 93 , 37 .3319°N 22.0743°E, leg. Pintér, E. & P. Subai 03.08.1976, S 3779/1 ; 400 from the crossing Dimitsána–Andrítsena– Megalópoli direction Dimitsána, on limestone rocks along border of road, 355 m alt., UTM EG 94 , 37 .4810°N 22.0492°E, leg. Subai 05.04.1986, S 15784 /1 ; 12.5 km S from Dimitsána (= crossing Dimitsána–Trípoli–Megalópoli), on limestone rocks along border of road, 1130 m alt., UTM EG 95 , 37 .5453°N 22.0894°E, leg. Subai 05.04.1986, S 15786 /1 (juv.) ; S border of Magoúliana, UTM EG 97 , 37 .67°N 22.12°E, leg. Pintér & Varga 21.7.1992, HNHM 44846 /1 ; on rocks below Moni Ambeláki (= Taigetos Mountains NE border), 650 m alt., UTM FG 02 , 37 .2692°N 22.2181°E, leg. Hadjicharalambous & Subai 08.05.1991, S 15773 /2 ; 5 km from Vitína direction Arhea Olympia (= highway Arhea Olympia–Trípoli), 985 m alt., UTM FG 06 , 37 .6515°N 22.1685°E, leg. Pintér, E. & P. Subai 02.08.1976, S 3778/1 ; leg. Subai 5.4.1986, S 15800 /1 ; leg. Hadjicharalambous & Subai 8.5.1991, S 15775 /1 ; 3.7 km from Perthóri direction Moni Epáno Hrépas, on limestone rocks, 1000 m alt., UTM FG 15 , 37 .5417°N 22.3347°E, leg. Hadjicharalambous & Subai 08.05.1991, S 15774 /3 ; Levídi, UTM FG 17 , 37 .68°N 22.29°E, SMF 207810 /3 ; 4.7 km from Levídi direction Kandíla , on limestone rocks, 720 m alt., UTM FG 17 , 37 .7214°N 22.3276°E, leg. Hadjicharalambous & Subai 07.05.1991, S 15783 /10 ; 4 km from Manthiréa direction Spárti , on limestone rocks, 860 m alt., UTM FG 23 , 37 .3813°N 22.4034°E, leg. Pintér, E. & P. Subai 05.08.1976, S 3780/1 ; 1.5 km SSE of Kandíla , near Kandílas monastery, 800 m alt., UTM FG 28 , 37 .7635°N 22.3745°E, leg. Gittenberger 17.8.2000, NNM/5 ; Parnon Mountains , approx. 5.5 km SSE of Ágios Pétros (= approx. 2.5 km from road junction to Kastanitsa), on limestone rocks and among rock pebbles, 970 m alt., UTM FG 32 , 37 .3114°N 22.5775°E, leg. Riedel & Subai 20.04.1988, S 15769 /2 ; Ágios Ioánnis, UTM FG 43 , 37 .35°N 22.63°E, leg. Neuteboom 4.9.1982, NNM/1 ; 1 km from Ágios Ioánnis direction Ástros, on limestone rocks, 750 m alt., UTM FG 43 , 37 .3539°N 22.6401°E, leg. Subai 16.07.1979, S 6425/9 ; between Elónis monastery and Kosmās, small caves near Ágios Dimitri, UTM FG 50 , 37 .13°N 22.76°E, leg. Neuteboom 30.8.1982, NNM/2 (juv.) ; SW border of Ágios Andreas , on limestone rocks, 90 m alt., UTM FG 53 , 37 .3437°N 22.7690°E, leg. Subai 16.07.1979, S 6424/9 ; 1 km E of Ágios Andréas , UTM FG 53 , 37 .34°N 22.78°E, leg. Hoenselaar 14.5.1996, Maa /1 ; 2 km SE of Ágios Andréas , 25 m alt., UTM FG 53 , 37 .33°N 22.79°E, leg. Neuteboom 30.8.1982, NNM/1 + 1 (juv.) ; 3 km SE of Ágios Andréas , UTM FG 53 , 37 .32°N 22.79°E, leg. Kiss & Pintér, 18.8.1988, HNHM 44594 /2 + 1 (juv.) ; 5 km SE of Ágios Andréas , opposite of Camping "Arkadia", UTM FG 53 , 37 .33°N 22.80°E, leg. Hoenselaar 12.5.1996, Maa /1 ; leg. Neuteboom 30.8.1982, NNM/1 + 2 (juv.) ; 3 km SE of Ástros , UTM FG 53 , 37 .38°N 22.77°E, leg. Kiss & Pintér, 18.8.1988, HNHM 44593 /39 + 7 (juv.) ; 7 km from Ástros direction Árgos, on limestone rocks, 75 m alt., UTM FG 54 , 37 .4663°N 22.7421°E, leg. Subai 16.07.1979, S 15753 /14 ; leg. Neuteboom 4.9.1982, NNM/3 + 3 (juv.) ; on rocks of coastal road Kivéri– Ástros , 7.4 roadkm. N of bifurcation to Parálio Ástros, 25 m alt., UTM FG 54 , 37 .49°N 22.72°E, leg. Gittenberger 25.7.2000, NNM 71842/2 ; 1 km S von Kivéri, an Kalkfelsen , UTM FG 55 , Subai 17.7.1979, S 15755 /13 + 2 (juv.) ; leg. Kiss & Pintér 17.8.1988, HNHM 44592 /1 + 3 (juv.) ; Pirgoúdi, UTM FG 60 , leg. Kiss & Pintér, 18.8.1988, HNHM 85639 /4 ; on rocks near Livádi (= 5 km NE of Leonídio ), 25 m alt., UTM FG 61 , 37 .19°N 22.89°E, leg. Gittenberger 20.7.2000, NNM 71861/1 ; on rocks along the road, 1.7 km NE of Tsitália , UTM FG 61 , 37 .13°N 22.87°E, leg. Gittenberger 21.7.2000, NNM 71858/3 ; on conglomerate rocks at W border of Leonídio , 100 m alt., UTM FG 61 , 37 .1694°N 22.8529°E, leg. Subai 16.07.1979, S 6690/39 ; leg. Butot & Subai, 19.2.1981, S 15806 /25 + 4 (juv.) ; 1 km W of Leonídio , UTM FG 61 , 37 .16°N 22.83°E, leg. Kiss & Pintér, 18.8.1988, HNHM 44595 /14 + 4 (juv.) ; Pláka, UTM FG 61 , 37 .14°N 22.89°E, leg. Kiss & Pintér, 18.8.1988, HNHM 85640 /1 ; gorge 4 km WSW of Leonídio , 285 m alt., UTM FG 61 , 37 .15°N 22.79°E, leg. Neuteboom 30.8.1982, NNM/6 ; leg. Kiss & Pintér, 18.8.1988, HNHM 44596 /1 ; 5.5 km from Ágios Andréas direction Tirós, UTM FG 62 , 37 .25°N 22.85°E, leg. Neuteboom 30.8.1982, NNM/1 ; 5 km from Ágios Andreas direction Tiros, on limestone rocks, 25 m alt., UTM FG 63 , 37 .3242°N 22.8161°E, leg. Subai 16.07.1979, S 6426/57. — Argolída, mountain pass 7 km from Kandíla direction Psári, Olígirtos Mountains, on limestone rocks, 1400–1450 m alt., UTM FG 28 , 37 .7944°N 22.3979°E, leg. Hadjicharalambous & Subai 07.05.1991, S 15782 /9 ; 3 km NW of Ahladókambos , UTM FG 35 , 37 .54°N 22.53°E, leg. Kiss & Pintér 20.7.1986, HNHM 29907 /2 (juv.) ; 8 km from Ahladókambos direction Trípoli, on limestone rocks, 720 m alt., UTM FG 35 , 37 .5356°N 22.5136°E, leg. Pintér, E. & P. Subai 05.08.1976, S 3772/1 ; mountain slope 12.5 km from Agiorgítika direction Árgos (= environment of Ahladókambos ), on limestone rocks, 800 m alt., UTM FG 35 , 37 .5385°N 22.5741°E, leg. Subai 05.04.1986, S 15787 /7 ; on rocks 1.3 roadkm. E of Andrítsa , 120 m alt., UTM FG 44 , 37 .47°N 22.62°E, leg. Gittenberger 24.7.2000, NNM 71848/2 ; Trípoli – Nafplio Landstrasse, highway Trípoli – Nafplio , 4 km from the bifurcation to Spiliotákis direction Trípoli, on limestone rocks, 600 m alt., UTM FG 45 , 37 .5232°N 22.6404°E, leg. Pintér, E. & P. Subai 05.08.1976, HNHM 39176 /4, S 3774/5 ; mountain slope 7 km from Ahladókambos direction Árgos, on limestone rocks, 650 m alt., UTM FG 45 , 37 .5181°N 22.6173°E, leg. Subai 05.04.1986, S 15788 /3 ; leg. Riedel & Subai 20.4.1988, S 15768 /1 ; in Kefalari, on limestone rocks and from a rich spring, 30–35 m alt., UTM FG 46 , 37 .5959°N 22.6882°E, leg. Subai, 18.07.1979, S 15743 /7 ; on rocks 12 roadkm. E of Andrítsa (= 3 km W of Kivéri), 55 m alt., UTM FG 55 , 37 .50°N 22.68°E, leg. Gittenberger 24.7.2000, NNM 71853/1 ; Nafplio (= Nauplia, Nauplion), UTM FG 55 , 37 .56°N 22.81°E, SMF 101867 /6, 101880/1, 207813/2, 284883/6, 284898/3, 284899/3, 284904/2 ; leg. Subai, 18.7.1979, S 15742 /19 ; leg. Sin 6–7.7.1985, HNHM 24037 /11 + 2 (juv.), 24042/6 ; leg. Kiss & Pintér, 19.7.1986, HNHM 29891 /122 + 46 (juv.) ; on ruins of the old castle of Tiryns, 15 m alt., UTM FG 56 , 37 .5990°N 22.8001°E, leg. Pintér, E. & P. Subai 06.08.1976, HNHM 39180 /12 + 1 (juv.), S 3769/20 ; leg. Neuteboom 30.8.1982, NNM/5 ; leg. Mészáros 1.5.1986, HNHM 44342 /2 ; leg. Kiss & Pintér, 19.7.1986, HNHM 29913 /8 + 4 (juv.) ; Árgos, on ruins of the old castle, 260 m alt., UTM FG 56 , 37 .6384°N 22.7150°E, leg. Pintér, E. & P. Subai 05.08.1976, S 3773/15 ; Subai , 18.7.1979, S 6699/97 + 1 (alk.) ; Mykenai, UTM FG 57 , 37 .72°N 22.75°E, SMF 284961 /8, 285171/6 ; leg. Pfeiffer, 1926, SMF 101847 /4 ; leg. Erőss 21.7.1992, S, 18260/1 ; ruins of the old castle of Mykene and in valley beside it, 220 m alt., UTM FG 57 , 37 .7291°N 22.7577°E, leg. Pintér, E. & P. Subai 06.08.1976, HNHM 39168 /5, S 3770/5 ; leg. Nosek 29.9.1976, HNHM 23962 /1 ; leg. Sín 8.7.1985, HNHM 24046 /5 + 1 (juv.) ; leg. Kiss & Pintér 20.7.1986, HNHM 29912 /16 + 4 (juv.) ; leg. Subai 20.4.1988, S 15767 /24 ; Toló, UTM FG 65 , 37 .52°N 22.85°E, leg. Kiss & Pintér 17.8.1988, HNHM 44591 /58 + 13 (juv.) ; Toló, Camping "Kastráki", UTM FG 65 , 37 .52°N 22.85°E, 37.52°N 22.85°E, leg. Hoenselaar 15.5.1996, Maa/9 ; surroundings of Drépano, UTM FG 65 , 37 .53°N 22.89°E, leg. Mészáros 30.4.1986, HNHM 44340 /41 + 30 (juv.) ; ancient Asine (= near Asíni, SE of Nafplio ), UTM FG 65 , 37 .52°N 22.87°E, leg. Hoenselaar 15.5.1996, Maa/2 ; from road Pirgiótika–Arkadiko towards Ágios Dimítrios , UTM FG 66 , 37 .58°N 22.93°E, leg. Neuteboom 4.9.1982, NNM/3 ; mountain slope 2 km from Agionóri direction Límnes, on limestone rocks, 700 m alt., UTM FG 67 , 37 .7439°N 22.8739°E, leg. Subai 04.04.1986, S 15792 /22 ; mountain slope 3 km from Agionóri direction Límnes, on limestone rocks, 730 m alt., UTM FG 67 , 37 .7388°N 22.8772°E, leg. Subai 04.04.1986, S 15793 /1 + 1 (juv.) ; Néa Epídavros , bifurcation to Ligoúrio, UTM FG 87 , 37 .68°N 23.09°E, leg. Neuteboom 4.9.1982, NNM/8 ; 1 km N of Néa Epídavros , UTM FG 87 , 37 .68°N 23.13°E, leg. Kiss & Pintér 16.8.1988, HNHM 85642 /19 ; 1 km N of Ermióni , UTM FG 94 , 37 .39°N 23.24°E, leg. Kiss & Pintér 17.8.1988, HNHM 44590 /3 + 2 (juv.) ; steps of the amphitheatre in Epidauros, UTM FG 86 , 37 .6334°N 23.1599°E, leg. Pintér, E. & P. Subai 06.08.1976, HNHM 39173 /4, S 3771/6 ; 1 km N of Trahiá , UTM FG 96 , 37 .58°N 23.15°E, leg. Kiss & Pintér 16.8.1988, HNHM 85638 /12 ; 1 km W von Eleónas , UTM n. det., leg. Kiss & Pintér 24.7.1986, HNHM 29898 /12 + 3 (juv.) ; Ília bei Kastrí, UTM n. det., leg. Philippson 8.9.1887, SMF 284896 /7 + 1 (juv.). — Messenía, ruins of the old Navarino palace (= Paleókástro, S of Petrohóri ), under stones, on limestone rocks, 120 m alt., UTM EF 58 , 36 .9582°N 21.6568°E, leg. Riedel & Subai, 18.04.1988, S 15764 /6 ; on walls of the old castle in Pílos , 30 m alt., UTM EF 68 , 36 .9118°N 21.6909°E, leg. Subai 06.04.1986, S 15790 /8 ; E border of Pílos , on limestone rocks, 50 m alt., UTM EF 68 , 36 .9204°N 21.7052°E, leg. Subai 11.07.1979, S 15748 /3 ; 6 km from Pílos direction Kalamáta, on limestone rocks, 70 m alt., UTM EF 68 , 36 .9373°N 21.7429°E, leg. Subai 06.04.1986, S15791 /2 ; Camping "Navarino Beach" NE of Pílos , UTM EF 68 , 36 .93°N 21.71°E, leg. Hoenselaar 26.4.1996, Maa/2 ; cave niches 8 km from Methóni direction Pílos , on limestone rocks, UTM EF 68 , 36 .8877°N 21.7044°E, leg. Subai 11.07.1979, S 15749 /6 ; 8 km from Handrinós direction Pílos (= highway Pílos–Messíni), on limestone rocks, 50 m alt., UTM EF 68 , 36 .9373°N 21.7423°E, leg. Subai 12.07.1979, S 6572/3 + 5 (alk.) ; Néstor’s Palace between Hóra and Korifási , tomb of Néstor, UTM EF 69 , 37 .04°N 21.70°E, leg. Hoenselaar 27.4.1996, Maa /2 ; mountain SW of Hrisokellariá , on limestone rocks, 400 m alt., UTM EF 77 , 36 .7864°N 21.8801°E, leg. Hadjicharalambous & Subai 11.05.1991, S 15777 /10 ; Kastánia near Petalidi, UTM EF 89 , 36 .95°N 21.88°E, leg. Philippson 26.10.1887, SMF 284917 /1, 284918/3 ; Kalamáta , UTM EF 99 , EG 90 , 37 .02°N 22.11°E, leg. Philippson 14.10.1887, SMF 284895 /1 ; cave niches 1 km from Gargaliáni direction Kiparissía , on limestone rocks, 200 m alt., UTM EG 50 , 37 .0667°N 21.6199°E, leg. Subai 10.07.1979, S 6666/8 ; Filiatrá, UTM EG 51 , 37 .15°N 21.58°E, leg. Thiesse, 1890, SMF 284903 /17 ; 2.9 km from Filiatrá direction Pláti, on limestone rocks, 140 m alt., UTM EG 51 , 37 .1657°N 21.6212°E, leg. Hadjicharalambous & Subai 12.05.1991, S 15757 /11 ; in Touloupa Hani Village (= E of Hóra and Metamórfosi), on limestone rocks, 470 m alt., UTM EG 60 , 37 .0621°N 21.7804°E, leg. Riedel & Subai, 18.04.1988, S 15763 /5 ; E border of Rodiá (= SE of Kiparissía), on limestone rocks and along river bank, 420 m alt., UTM EG 61 , 37 .2106°N 21.7336°E, leg. Riedel & Subai, 18.04.1988, S 15765 /2 ; 1.2 km E of Rodiá , on low limestone rocks, 390 m alt., UTM EG 61 , 37 .2129°N 21.7400°E, leg. Hadjicharalambous & Subai 12.05.1991, S 15758 /8 ; ruins of the old castle in Kiparissía , 100 m alt., UTM EG 62 , 37 .2506°N 21.6783°E, leg. Subai 10.07.1979, S 6411/42 ; on dry limestone rocks 13 km N of Kiparissía (= highway Pírgos – Pílos ), 50 m alt., UTM EG 63 , 37 .3473°N 21.7015°E, leg. Subai 10.07.1979, S 15881 /8 ; Ithómi (= antic Messene), on ruins of the town walls, 350 m alt., UTM EG 81 , 37 .1784°N 21.9184°E, leg. Pfeiffer, 1926, SMF 101850 /7 ; leg. Pintér & Subai 4.8.1976, HNHM 39190 /4, S 3788/1 ; leg. Subai 12.07.1979, S 6410/1 ; leg. Kiss & Pintér 22.7.1986, HNHM 29908 /5 + 2 (juv.) ; leg. Hoenselaar 24.4.1996, Maa /2 ; 2 km W of the crossing near Lámbena (= N of Messíni ), 230 m alt., UTM EG 81 , 37 .1485°N 21.9418°E, leg. Pintér, E. & P. Subai 04.08.1976, S 3787/10 ; mountain slope 6.6 km from Lámbena direction Mavrongáti, on limestone rocks, 280 m alt., UTM EG 81 , 37 .1517°N 21.9299°E, leg. Subai 05.04.1986, S 15785 /2 ; 5.5 km from Ithómi direction Lámbena, on limestone rocks along border of road, 275 m alt., UTM EG 81 , 37 .1511°N 21.9299°E, leg. Subai 12.07.1979, S 15754 /3 ; 3 km N of Bassai , UTM EG 84 , 37 .45°N 21.92°E, leg. Kiss & Pintér 20.8.1988, HNHM 44599 /2 + 1 (juv Málta village (= SE of Kalamáta ), on medieval castle ruins, 370 m alt., UTM FF 08 , 36 .9313°N 22.1915°E, leg. Pintér, E. & P. Subai 04.08.1976, HNHM 39178 /9, S 3789/7 ; 4 km of Kalamáta , near Camping "Maria’s See and Sun", UTM FF 09 , 37 .02°N 22.02°E, leg. Hoenselaar 28.4.1996, Maa /4 ; above Mikra Mandínia (= Taigetos Mountains SE of Kalamáta), 120 m alt., UTM FF 09 , 36 .9845°N 22.1619°E, leg. Pintér, E. & P. Subai 04.08.1976, HNHM 39181 /4, S 3790/5 ; N of Kámbos (= Taigetos Mountains SE of Kalamáta), on limestone rocks, 210 m alt., UTM FF 09 , 36 .9556°N 22.1913°E, leg. Pintér, E. & P. Subai 04.08.1976, HNHM 39183 /1 ; limestone rocks in valley near Langáda, 430 m alt., UTM FF 17 , 36 .7801°N 22.3383°E, leg. Butot & Subai 16.02.1981, S 17531 /13 ; leg. Neuteboom 31.8.1982, NNM/4 ; Roman stone highway near Plátsa, on limestone rocks, 370 m alt., UTM FF 17 , 36 .8074°N 22.3205°E, leg. Butot & Subai 16.02.1981, S 17532 /11 ; border of Plátsa direction Nomítsis, UTM FF 17 , 36 .80°N 22.31°E, leg. Neuteboom 31.8.1982, NNM/1 ; 15 km from Pigí direction Kalamáta , on limestone rocks, 25 m alt., UTM FF, 18, 36.8592°N 22.2564°E, leg. Subai 13.07.1979, S 6427/3 ; S border of Kardamili, on limestone rocks, UTM FF, 18, 36.8849°N 22.2356°E, leg. Butot & Subai 16.02.1981, S 17530 /1 ; leg. Neuteboom 31.8.1982, NNM/5 ; in valley W of Kéndro , on limestone rocks and from a spring at river bank, 575 m alt., UTM FF, 18, 36.9504°N 22.2502°E, leg. Subai 06.04.1986, S 15789 /1 ; gorge 3.8 km E of Kéndro (= Chóra Gaitses), approx. 720 m alt., UTM FF, 19, 36.9749°N 22.2596°E, leg. Hadjicharalambous & Subai 10.05.1991, S 15770 /4 ; highway Kalamáta–Spárti , start of Artemíssia gorge (= Taigetos Mountains NE of Kalamáta), 710 m alt., UTM FG 00, 37.0298°N 22.1865°E, leg. Pintér, E. & P. Subai 05.08.1976, HNHM 39185 /15 + 2 (juv.), S 3786/16 ; Artemissía gorge 9.4 km from Kalamáta direction Artemissía, on limestone rocks, 310 m alt., UTM FG 00, 37.0840°N 22.1575°E, leg. Hadjicharalambous & Subai 11.05.1991, S 15776 /41 + 4 (juv.) ; Dimióvis monastery (= Taigetos Mountains NE of Kalamáta), on limestone rocks, 820 m alt., UTM FG 00, 37.0412°N 22.2001°E, leg. Pintér, E. & P. Subai 03.08.1976, HNHM 39189 /4 ; on limestone rocks at border of Eleohóri Village (= Janitsa), 545 m alt., UTM FG 00, 37.0408°N 22.1817°E, leg. Hadjicharalambous & Subai 10.05.1991, S 15771 /10 ; Taigetos Mountains , at the governmental border of Messinía to Lakonía, on marble rocks, 880 m alt., UTM FG 10 , 37 .0745°N 22.2955°E, leg. Pintér, E. & P. Subai 05.08.1976, HNHM 39169 /2 ; leg. Neuteboom 31.8.1982, NNM/1 ; Kalamáta–Areópoli highway, chapel St. Nicholas Kabinari, UTM n. det., leg. Hoenselaar 28.4.1996, Maa/1. — Lakonía, on limestone rocks at W border of Geroliménas, 25 m alt., UTM FF 23 , 36 .4842°N 22.3993°E, leg. Butot & Subai 14.02.1981, S 17527 /8 + 5 (juv.) ; leg. Kiss & Pintér 21.7.1986, HNHM 29904 /1 + 1 (juv.) ; Geroliménas, road to Areópoli , UTM FF 23 , 36 .48°N 22.39°E, leg. Neuteboom 2.9.1982, NNM/2 + 1 (juv.) ; Geroliménas, road to Koúnos, UTM FF 23 , 36 .49°N 22.39°E, leg. Neuteboom 2.9.1982, NNM/1 + 1 (juv.) ; W of Pírrihos (= between Kótronas and Areópoli), UTM FF 25 , 36 .65°N 22.43°E, leg. Neuteboom 2.9.1982, NNM/2 (juv.) ; Pirgos (= S of Areópoli ), on limestone rocks beside the "Diros" cave, 5–10 m alt., UTM FF 25 , 36 .6382°N 22.3792°E, leg. Subai 13.07.1979, S 15812 /18 ; leg. Butot & Subai 14.2.1981, S 15807 /71 + 4 (juv.) + 1 (alk.) ; leg. Kiss & Pintér 21.7.1986, HNHM 29916 /6 + 3 (juv.) ; 2 km W of Vahós , UTM FF 26 , 36 .68°N 22.40°E, leg. Kiss & Pintér 21.7.1986, HNHM 29899 /3 ; on limestone rocks near Liméni (= N of Areópoli ), 40 m alt., UTM FF 26 , 36 .6787°N 22.3778°E, leg. Butot & Subai 16.02.1981, S 17529 /21 ; S border of Néo Ítilo, on limestone rocks, 30 m alt., UTM FF 26 , 36 .6877°N 22.3862°E, leg. Hadjicharalambous & Subai 10.05.1991, S 15772 /2 ; mountain slope 11 km from Petrina direction Árna, on low limestone rocks, 760 m alt., UTM FF 28 , 36 .8785°N 22.4254°E, leg. Butot & Subai 15.02.1981, S 17528 /3 ; between Ágios Dimítrios and Kokála, UTM FF 33 , 36 .51°N 22.47°E, leg. Neuteboom 2.9.1982, NNM/3 + 2 (juv.) ; mountain slope at border of Githio direction Spárti , on limestone rocks, 30 m alt., UTM FF 36 , 36 .7594°N 22.5562°E, leg. Butot & Subai, 18.02.1981, S 17537 /2 ; 8.5 km from Githio direction Areópoli , in forest S of the highway, on limestone rocks, 150 m alt., UTM FF 36 , 36 .7398°N 22.4936°E, leg. Hadjicharalambous & Subai 10.05.1991, S 15880 /3 ; gorge 10 km from harbor of Githio direction Areópoli , on limestone rocks, 40 m alt., UTM FF 36 , 36 .7299°N 22.5055°E, leg. Butot & Subai, 18.02.1981, S 17536 /7 + 3 (juv.) ; gorge 15 km from port of Githio direction Areópoli , on limestone rocks, 100 m alt., UTM FF 36 , 36 .7023°N 22.4725°E, leg. Butot & Subai leg. 14.02.1981, S 17526 /4 ; gorge 12 km from Githio direction Areópoli , on limestone rocks and from a spring, 30 m alt., UTM FF 36 , 36 .7178°N 22.4998°E, leg. Subai 13.07.1979, S 15752 /1 ; approx. 14 km from Githio direction Areópoli , on limestone rocks beside the highway, 100 m alt., UTM FF 36 , 36 .7017°N 22.4713°E, leg. Hadjicharalambous & Subai 10.05.1991, S 15810 /1 + 1 (juv.) ; 1 km S of Goritsá , UTM FF 39 , 37 .01°N 22.56°E, leg. Kiss & Pintér, 19.8.1988, HNHM 44597 /36 + 7 (juv.) ; Spárti – Geráki country road, 1 km E of Kefalās, UTM FF 39 , 37 .02°N 22.53°E, leg. Hoenselaar 4.5.1996, Maa/1 ; Stefaniá, UTM FF 47 , 36 .84°N 22.60°E, leg. Kiss & Pintér 20.7.1986, HNHM 29914 /2 + 2 (juv.) ; 6.5 km from Skála direction Githio, on limestone rocks beside a bridge and from the creek, UTM FF 47 , 36 .8048°N 22.6179°E, leg. Subai 14.07.1979, S 6578/4 ; 13 km from Skála direction Githio, on limestone rocks, 50 m alt., UTM FF 47 , 36 .7959°N 22.5957°E, leg. Subai 14.07.1979, S 6660/2 ; 3 km from Skála direction Spárti , on limestone rocks beside highway, 100 m alt., UTM FF 47 , 36 .8594°N 22.6058°E, leg. Subai 14.07.1979, S 15747 /76 ; 4 km SE of Agii Anárgiri , UTM FF 49 , 37 .00°N 22.66°E, leg. Kiss & Pintér, 19.8.1988, HNHM 44598 /1 ; Ortsrand von Geráki in Richtung Spárti , unter Steinen, UTM FF 59 , leg. Butot & Subai, 18.02.1981, S 17538 /1 ; 5 km from the crossing Geráki – Spárti , Skála – Kosmás direction Kosmás, on limestone rocks, 800 m alt., UTM FF 59 , 37 .0607°N 22.7071°E, leg. Butot & Subai, 18.02.1981, S 17539 /3 + 2 (juv.) ; 4 km N from the bifurcation to Mólai , on limestone rocks 500 m E of the highway, 250 m alt., UTM FF 67 , 36 .8456°N 22.8448°E, leg. Subai 14.07.1979, S 15745 /12 ; 4 km N of Mólai , UTM FF 67 , 36 .84°N 22.83°E, leg. Neuteboom 1.9.1982, NNM/4 + 1 (juv.) ; Apídia, road direction Spárti–Monemvassía , UTM FF 68 , 36 .88°N 22.79°E, leg. Neuteboom 1.9.1982, NNM/2 (juv.) ; rock gorge 10.5 km from the bifurcation to Mólai direction Skála, 250 m alt., UTM FF 68 , 36 .8635°N 22.8154°E, leg. Subai 14.07.1979, S 6802/1 ; highway Skála – Mólai , on limestone rocks near bifurcation to Niáta, 230 m alt., UTM FF 68 , 36 .8683°N 22.8145°E, leg. Butot & Subai 17.02.1981, S 17534 /11 + 2 (juv.) ; 700 m N of Nómia (= road direction Kardenitsa), UTM FF 75 , 36 .65°N 23.01°E, leg. Neuteboom 2.9.1982, NNM/1 + 2 (juv.) ; Elika, before junction to Arhángelos, UTM FF 75 , 36 .62°N 22.89°E, leg. Neuteboom 1.9.1982, NNM/5 + 2 (juv.) ; 9 km from Lira direction Nómia , on limestone rocks, 85 m alt., UTM FF 75 , 36 .6521°N 23.0139°E, leg. Butot & Subai 17.02.1981, S 17533 /7 + 1 (juv.) ; 9 km from Monemvassía direction Skála, on limestone rocks along border of road, 100 m alt., UTM FF 76 , 36 .7301°N 22.9882°E, leg. Subai 14.07.1979, S 15746 /8 ; Sikía– Monemvassía highway after junction to Veliés, UTM FF 76 , 36 .72°N 22.97°E, leg. Neuteboom 1.9.1982, NNM/3 + 3 (juv.) ; Veliés – Angelóna road, 3 km before Sikía, UTM FF 76 , 36 .74°N 22.95°E, leg. Neuteboom 1.9.1982, NNM/4 + 1 (juv.) ; Talanda, before junction to Ágios Nikólaos, UTM FF 76 , 36 .67°N 22.94°E, leg. Neuteboom 1.9.1982, NNM/3 ; Ágios Apóstoli (= 16 km SE of Pandanassa), 36.55°N 23.02°E, UTM FF 84 , leg. Neuteboom 1.9.1982, NNM/7 + 3 (juv.) ; mountain near Dermatiánika (= Kabos, N of Neapolis ), on limestone rocks and on house ruins, 70 m alt., UTM FF 84 , 36 .5451°N 23.0533°E, leg. Butot & Subai 17.02.1981, S 15805 /53 + 4 (juv.) + 12 (alk.) ; 1 km N of Monemvassía , UTM FF 86 , 36 .68°N 23.03°E, leg. Kiss & Pintér 21.7.1986, HNHM 29906 /4 ; in the old town of Monemvassía, on wall remains and under stones, 50 m alt., UTM FF 86 , 36 .6888°N 23.0572°E, leg. Subai 14.07.1979, S 6579/11 ; leg. Butot & Subai 17.2.1981, S 17525 /6 + 1 (juv.) + 3 (alk.) ; leg. Neuteboom 1.9.1982, NNM/10 + 3 (juv.) ; leg. Kiss & Pintér 20.7.1986, HNHM 29893 /50 + 4 (juv.) ; 2 km from Monemvassía direction Nómia, on rocks at the sea shore beside the highway, UTM FF 86 , 36 .6710°N 23.0306°E, leg. Butot & Subai 17.02.1981, S 17535 /6 + 1 (alk.) ; leg. Kiss & Pintér 21.7.1986, HNHM 29892 /28 + 14 (juv.) ; well of Trípi Village , UTM FG 10 , 37 .09°N 22.35°E, leg. Neuteboom 31.8.1982, NNM/1 (juv.) ; Mistra, byzantine ruin town W of Spárti , 410 m alt., UTM FG 20 , 37 .0739°N 22.3691°E, leg. Pintér, E. & P. Subai 05.08.1976, HNHM 39167 /6, S 3797/7 ; leg. Neuteboom 3.9.1982, NNM/1 (juv.) ; leg. Kiss & Pintér 22.7.1986, HNHM 29910 /1 + 1 (fragment) ; Spárti , UTM FG 20 , 37 .07°N 22.43°E, leg. Pfeiffer, 1926, SMF 101849 /1 ; 4 km from Spárti direction Hrísafa, UTM FG 20 , 37 .12°N 22.49°E, leg. Neuteboom 3.9.1982, NNM/2 ; 2 km S of Kástro , UTM n. det., leg. Kiss & Pintér 9.7.1986, HNHM 29902 /3. — Adelfí Island , NW part of the Island, 20 m alt., UTM GJ 53 , 39 .11°N 23.97°E, leg. Uit de Weerd 5.6.1999, NNM/2 . — Adelfópoulo Island , S part of central area, 20 m alt., 39.12°N 23.98°E, UTM GJ 53 , leg. Uit de Weerd 5 6.1999, NNM/5 . — Alónissos Island , Patitíri–Miléa road, 1.2–1.6 km NE of harbour Patitíri (= approx. 2.5–2.8 km NE of Alónissos town), 210 m alt., 39.15°N 23.87°E, UTM GJ 43 , leg. Uit de Weerd 4.6.1999, NNM/1 + 3 ; 150 m from road Patitíri – Miléa towards Leftogiálo (= 4.5 km NE of Alónissos town ), 160 m alt., UTM GJ 44 , 39 .16°N 23.88°E, leg. Uit de Weerd 4.6.1999, NNM/2 ; Ormos Tsortsi (= 4.2 km NE of Alónissos town ), 4 m alt., UTM GJ 53 , 39 .16°N 23.88°E, leg. Uit de Weerd 6.6.1999, NNM/2 ; Patitíri – Gerakis road, 2.1 km NE from turning point to Leftos Giakos (= 6.5 km NE of Alónissos town ), UTM GJ 54 , 39 .17°N 23.90°E, leg. Uit de Weerd 6.6.1999, NNM/1 . — Gioúra Island , UTM KD 56 , 39 .39°N 24.16°E, leg. Storch 7.7.1978, SMF 284930 /1 ; Gioúra Island , E coast, UTM KD 56 , 39 .38°N 24.18°E, leg. Storch 6.7.1978, SMF 284929 /1. — Kirá Panagiá Island , UTM KD 45 , 39 .32°N 24.07°E, leg. Storch 1.7.1978, SMF 284928 /3. — Pipéri Island , UTM KD 65 , 39 .35°N 24.32°E, leg. Storch 4.7.1978, SMF 284927 /1. — Skándzoura Island , UTM KD 52 , 39 .08°N 24.11°E, leg. Storch 9.7.1978, SMF 284925 /9 ; Skándzoura monastery, UTM KD 52 , 39 .08°N 24.11°E, leg. Storch 10.7.1978, SMF 284926 /3. — Skíathos Island , UTM n. det., SMF 284945 /2 ; Skíathos town , "Odos Filokleus Georgiadou" 200 m NE of harbour, 20 m alt., UTM GJ 13 , 39 .16°N 23.49°E, leg. Gittenberger & Uit de Weerd 29.5.1999, NNM/3 ; Evangelistrías monastery (= 4 km NNW of Skíathos town), 80 m alt., UTM GJ 14 , 39 .1904°N 23.4806°E, leg. Gittenberger & Uit de Weerd 30.5.1999, NNM/2 + 1 (juv.) ; 400 m N of Evangelistrías monastery, 150 m alt., UTM GJ 14 , 39 .19°N 23.48°E, leg. Gittenberger & Uit de Weerd 30.5.1999, NNM/4 ; 1 km N of Evangelistrías monastery, 230 m alt., 39.19°N 23.48°E, UTM GJ 14 , leg. Gittenberger & Uit de Weerd 29.5.1999, NNM/4 . — Skópelos Island , UTM n. det., leg. Banzhaf, 1929, SMF 284876 /1 (juv.) ; Kafasi, 1 km ESE of Metamórfosis monastery (= 3 km ESE of Skópelos town), 200 m N of junction, 300 m alt., UTM GJ 33 , 39 .11°N 23.75°E, leg. Uit de Weerd 3.6.1999, NNM/1 (juv.) ; Koprisies Mountain , 400–1300 m from peak (along the road), 320–370 m alt., UTM GJ 33 , GPS n. det., leg. Uit de Weerd 1.6.1999, NNM/1 + 2 + 24 ; 500 m NNE of church Ágios Konstantínos (= 1.2 km NW of Skópelos town), S slope, UTM GJ 33 , 39 .13°N 23.71°E, leg. Uit de Weerd 2.6.1999, NNM/1 . — Évvia Island (= Euböa), N border of Ágios Geórgios, in valley, on dry limestone rocks, 50 m alt., UTM FJ 60 , 38 .8410°N 22.8920°E, leg. Subai & Szekeres 10.05.1997, S 15687 /2 ; spruce forest 5 km from Gregolimano direction Ágios Geórgios, approx. 200 m N of the highway, on dry limestone rocks, 50 m alt., UTM FJ 60 , 38 .8401°N 22.9054°E, leg. Subai & Szekeres 10.05.1997, S 15699 /14 + 3 (juv.) + 1 (alk.) ; Loutrá Edipsoú , near church, UTM FJ 70 , 38 .85°N 23.04°E, leg. Neuteboom 25.10.1984, NNM/1 (juv.) ; Límni, UTM GH 09 , 38 .76°N 23.32°E, leg. Krüper, 1903, SMF 101970 /3 ; Mantoúdi, UTM GH, 19, 38.79°N 23.48°E, leg. Ottermann, 1912, SMF 284860 /22 ; S border of Mantoúdi , on limestone rocks at river side, 20 m alt., UTM GH, 19, 38.7942°N 23.4770°E, leg. Subai 01.05.1991, S 15708 /2 ; leg. Gittenberger & Uit de Weerd 10.6.1999, NNM/2 ; Halkída (= Chalcis), UTM GH 26 , 38 .46°N 23.58°E, SMF 284943 /3, 284944/4, 285166/2 ; leg. Fitz-Gerard, 1881, SMF 101986 /3 ; Kiparíssi (= 7 km N of Psahná ), UTM GH 27 , 38 .62°N 23.59°E, leg. Neuteboom 23.10.1984, NNM/1 + 4 (juv.) ; 1.5 km S of Néa Pagóntas , UTM GH 28 , 38 .65°N 23.56°E, leg. Neuteboom 24.10.1984, NNM/1 ; approx. 1–2 km N of Néa Artáki , approx. 200 m E of the highway, on low limestone rocks, UTM GH 26 , 38 .5404°N 23.6344°E, leg. Riedel & Subai 09.04.1988, S 15803 /4 + 2 (alk.) ; Dírfis Mountains (= Monte Delphi), UTM GH 47 , 38 .62°N 23.84°E, SMF 101879 /1, 284921/5, 284922/4, 284923/3, 285764/8 ; Stení, UTM GH 47 , 38 .58°N 23.83°E, SMF 101856 /4, 101871/15, 284924/3 ; above Áno Steni , on limestone rocks, 520 m alt., 520 m alt., UTM GH 47 , 38 .5847°N 23.8349°E, leg. Subai & Szekeres 10.05.1997, S 15917 /1 + 3 (juv.) ; Dírfi Mountains , N of the highway Steni Dirfios–Strópones, 200 m NE of the E. O.S. refuge hut, on limestone rocks, 1150 m alt., UTM GH 47 , 38 .6142°N 23.8597°E, leg. Subai & Szekeres 10.05.1997, S 15918 /1 ; 9 km from Stení Dírfios towards Strópones, UTM GH 47 , 38 .60°N 23.87°E, leg. Neuteboom 23.10.1984, NNM/2 + 3 (juv.) ; 3 km from Strópones towards Stení Dírfios, UTM GH 57 , 38 .60°N 23.88°E, leg. Neuteboom 23.10.1984, NNM/1 (juv.) ; small rock gorge at W border of Strópones (= Dírfis Mountains), on limestone rocks, 360 m alt., UTM GH 57 , 38 .6205°N 23.8853°E, leg. Subai 01.05.1991, S 15709 /6 ; S of Vassiliká , UTM GJ 01 , 38 .96°N 23.35°E, leg. Neuteboom 24.10.1984, NNM/2 + 1 (juv.) ; 1.7 km from Elliniká direction Vassiliká, on limestone rocks beside the quarry, 35 m alt., UTM GJ 01 , 38 .9979°N 23.3488°E, leg. Subai 30.04.1991, S 15707 /2 ; on rocks E of Elliniká , UTM GJ 02 , 39 .00°N 23.34°E, leg. Neuteboom 24.10.1984, NNM/1 ; W border of Alivéri , on limestone rocks and in rock debris, 350 m alt., UTM KC 45 , 38 .4022°N 24.0138°E, leg. Subai 16.04.1986, S 15883 /35 + 1 (alk.) ; leg. Neuteboom 22.10.1984, NNM/1 ; S of Kriezá , UTM KC 45 , 38 .38°N 24.14°E, leg. Neuteboom 21.10.1984, NNM/1 ; SE border of Koutoumoulas , under stones, 125 m alt., UTM KC 45 , 38 .3787°N 24.1105°E, leg. Subai 16.04.1986, S 15802 /2 ; cave niches W of Vrissi, foot of Mavrovoúni Mountains , on limestone rocks, 450 m alt., UTM KC 46 , 38 .5549°N 24.0268°E, leg. Subai 16.04.1986, S 15779 /2 ; road between Orológi and Monódrio, UTM KC 46 , 38 .53°N 24.08°E, N leg. euteboom 22.10.1984, NNM/2 ; 1 km S of Kaliáni , UTM n. det., leg. Neuteboom 22.10.1984, NNM/2 ; Konistraes, UTM KC 47 , 38 .54°N 24.06°E, SMF 101899 /4, 284957/4, 284958/2, 284959/2 ; Kími, UTM KC 48 , 38 .63°N 24.10°E, SMF 101873 /2 ; 8.1910, SMF 285164 /4, 285165/40 ; leg. Holtz 4.1926, SMF 207818 /7 ; between Néa Stíra and Stíra, UTM KC 52 , 38 .16°N 24.22°E, leg. Pintér 22.8.1976, HNHM 39166 /14 + 3 (juv.) ; Kapsála (= 3 km of Stíra ), UTM KC 52 , 38 .14°N 24.24°E, leg. Neuteboom 21.10.1984, NNM/5 ; "Malithianon", "Malithianu", "Malithiedin", UTM n. det., leg. Krüper, SMF 101855 /13, 284152/9, 285162/2 ; leg. Krüper, 1890, SMF 284940 /4, 284949/7 ; Fleva, UTM n. det., leg. Krüper 7.1928, SMF 101962 /5 ; "Achmet Aga", UTM n. det., leg. Godet, SMF 284942 /4 ; between Lépoura and Ag. Nikolaou monastery, UTM n. det., leg. Krüper, SMF 101868 /20. — Kea Island (= Keos), approx. 1 km E of Otziás, on limestone rocks on top of a slate mountain, 180–200 m alt., UTM KB 67 , 37 .6774°N 24.3625°E, leg. Subai 06.05.1991, S 15700 /34.

— Páros Island , UTM n. det., leg. Krüper, 1883, SMF 101864 /4. — Lipsí Island (= Lipsos, S of Sámos), surrounding Lipsí [Village], UTM MB 72 , 37 .29°N 26.77°E, leg. Becker & Wedekind 9.1984, S 15741 /1. — Sámos Island , UTM n. det., SMF 101981 /3 ; leg. Paravicini, 1933, SMF 284912 /2 ; between Marathókambos and Ágios Theódori, UTM MB 77 , 37 .73°N 26.71°E, leg. Neuteboom 8.5.1978, NNM/1 (juv.) ; Balos (= coast above Kouméika), UTM MB 77 , 37 .69°N 26.74°E, leg. Neuteboom 10.5.1978, NNM/1 ; Ireon, Hera Temple, UTM MB 86 , 37 .66°N 26.87°E, leg. Neuteboom 7.5.1978, NNM/1 (juv.) ; 4 km from Spatharéi direction Pagóndas, UTM MB 86 , 37 .66°N 26.79°E, leg. Neuteboom 10.5.1978, NNM/6 ; leg. Maassen 5.1993, Maa /7 ; 2 km N of Mavratzéi, UTM MB 87 , 37 .73°N 26.66°E, leg. Maassen 5.1993, Maa /1 ; Mavratzéi , UTM MB 87 , 37 .71°N 26.86°E, leg. Maassen 5.1993, Maa /13 ; 2 km SE of Míli, UTM MB 87 , 37 .67°N 26.87°E, leg. Maassen 5.1993, Maa /8 ; 1.5 km SE of Stravrinídes, UTM MB 88 , 37 .78°N 26.82°E, leg. Maassen 5.1993, Maa /1 ; 3 km E of Avlákia (= W of Kokkári), UTM MB 88 , 37 .79°N 26.86°E, leg. Maassen 5.1993, Maa /2 ; river valley 0.5 km from Platanakia direction Manolates, UTM MB 88 , GPS n. det., leg. Maassen 5.1993, Maa /9 ; 0.5 km W of Vourliotes, along path, UTM MB 88 , 37 .78°N 26.83°E, leg. Maassen 5.1993, Maa /3 ; wells N of Vourliotes, UTM MB 88 , 37 .78°N 26.84°E, leg. Maassen 5.1993, Maa /2 ; Sámos , UTM MB 97 , 37 .76°N 26.97°E, leg. Neuteboom 5.5.1978, NNM/11 (alk.) ; Pithagório , near Hotel "Dorissa Bay", under stones, UTM MB 97 , 37 .69°N 26.92°E, leg. Neuteboom 7.5.1978, NNM/9 + 1 (juv.) ; between Pithagório and Vathí, before pass, under stones, UTM MB 97 , 37 .72°N 26.96°E, leg. Neuteboom 7.5.1978, NNM/9 + 3 (juv.) ; Vathí (= between Sámos and Pithagório), UTM MB 97 , 37 .74°N 26.98°E, leg. Neuteboom 7–9.5.1978, NNM/6 + 3 (juv.) ; Thios Mountain , 1 km NE of Sámos, 200 m alt., UTM MB 97 , 37 .75°N 26.98°E, leg. Maassen 5.1993, Maa /1 ; Platanaki (= 1.5 km NW of Sámos ), UTM MB 97 , GPS n. det., leg. Maassen 5.1993, Maa /15, S 15937 /3 ; 3 km S of Kokkári (= direction Mitilinií ), 200 m alt., UTM MB 97 , 37 .75°N 26.90°E, leg. Maassen 5.1993, Maa /6 ; 1 km N of Mitilinií (= direction Kokkári ), UTM MB 97 , 37 .74°N 26.89°E, leg. Maassen 5.1993, Maa /2 ; 0.5 km from Pithagório direction Spilianis , UTM MB 97 , 37 .69°N 26.93°E, leg. Maassen 5.1993, Maa /3 ; above Spilianis monastery (= N of Pithagório), under stones, UTM MB 97 , 37 .69°N 26.93°E, leg. Maassen 5.1993, Maa /12 + 1 (juv.) S 15938 /3 ; 1 km W of Kokkári, UTM MB 98 , 37 .78°N 26.86°E, leg. Maassen 5.1993, Maa /1 ; 0.5 km S of Kokkári (= direction Mitilinií ), 50 m alt., UTM MB 98 , 37 .77°N 26.89°E, leg. Maassen 5.1993, Maa /3 ; 1 km S of Kokkári (= direction Mitilinií ), 150 m alt., UTM MB 98 , 37 .76°N 26.89°E, leg. Maassen 5.1993, Maa /1 ; Kótsikas monastery (= NE of Sámos), UTM MB 98 , 37 .78°N 26.96°E, leg. Maassen 5.1993, Maa /4 ; Ágios Paraskeví (= NE of Sámos), along beach, UTM MB 98 , 37 .78°N 26.99°E, leg. Maassen 5.1993, Maa /1 + 1 (juv.) ; 0.5 km S of Ágios Paraskeví, Thios Mountain , 150 m alt., 37.77°N 26.99°E, UTM MB 98 , leg. Maassen 5.1993, Maa /3 ; 1 km S of Ágios Paraskeví, Thios Mountain , 300 m alt., UTM MB 98 , 37 .76°N 26.99°E, leg. Maassen 5.1993, Maa /2 ; 3 km W of Ágios Paraskeví, UTM MB 98 , 37 .78°N 26.95°E, leg. Maassen 5.1993, Maa /16 ; Thios Mountain (= NE of Sámos), Profítis Ilías peak, 430 m alt., 37.76°N 26.99°E, UTM MB 98 , leg. Maassen 5.1993, Maa /3 ; 0.5 km N of Profítis Ilías peak, 350 m alt., 37.77°N 26.99°E, UTM MB 98 , leg. Maassen 5.1993, Maa /27 + 1 (juv.) S 15939 /3 ; W border of Kokkári , UTM MB 98 , 37 .78°N 26.87°E, leg. Neuteboom 8.5.1978, NNM/1 ; 1 km W of Agia Zóni and walls of monastery, UTM NB 07 , 37 .75°N 26.98°E, leg. Neuteboom 9.5.1978, NNM/13 + 37 (alk.) ; approx. 2–3 km E of Sámos , on rocks along sea shore, UTM NB 07 , 37 .76°N 27.02°E, leg. Maassen 5.1993, Maa /13 ; coast 1 km W of Psilí Ámmos (= E of Pithagório ), UTM NB 07 , 37 .70°N 26.99°E, leg. Maassen 5.1993, Maa /12 ; 1.5 km N of Psilí Ámmos , along path, UTM NB 07 , 37 .72°N 27.01°E, leg. Maassen 5.1993, Maa /9. — Kálimnos Island (= Calino), UTM n. det., SMF 284915 /18, 284916/2 ; Kálimnos town (= Pothia), UTM MA 98 , 36 .95°N 26.98°E, leg. Neuteboom 24.10.1979, NNM/10 ; gorge behind harbor of Kálimnos, UTM MA 98 , 36 .95°N 26.99°E, 1934, SMF 101877 /6 ; Kástro Hrisohórias, UTM MA 98 , 36 .96°N 26.96°E, leg. Neuteboom 24.10.1979, NNM/6 + 1 (juv.) + 15 (alk.) ; behind harbor of Vathis (= Vati), UTM NA 09 , 36 .97°N 27.02°E, leg. K. L. Pfeiffer, 1934, SMF 101976 /3. — Psérimos Island , UTM NA, 18, SMF 284914 /2 ; leg. K. L. Pfeiffer, 1934, SMF 207817 /2 ; leg. K. L. Pfeiffer, 1940, SMF 285175 /4 ; leg. Neuteboom 20.10.1979, NNM/18 ; on rocks near harbour, UTM NA, 18, 36.93°N 27.13°E, leg. K. L. Pfeiffer, 1934, SMF 101977 /30. — Kós Island , UTM n. det., SMF 207812 /2. — Sphanteria Island near Tilos Island, UTM n. det., leg. Philippson 2.4.1887, SMF 284891 /1.

Turkey, Çanakkale County, Gökçeada Island , UTM n. det., leg. Kleefisch 10.9.1985, (ex. coll. Schütt Nr. 1180) S 20789 /1 ; Çanakkale, Kilidürbahir , UTM ME 44 , 40 .17°N 26.37°E, leg. Schütt 28.4.2002, (ex. coll. Schütt Nr. 1751) S 20790 /29. — Manisa County, Cornana, UTM n. det., HNHM 39174 /2 ; Sochia , UTM n. det., SMF 101865 /4 ; Manisa (= ancient Magnesia ), UTM NC 37 , 38 .60°N 27.37°E, SMF 284909 /2. — AydIn County, DIlek YarImadasI MIllI ParkI, Kanyon 7 km direction GüzelçamlI, UTM NB 17 , 37 .68°N 27.16°E, leg. Schütt 23.4.2000, (ex. coll. Schütt Nr. 1686) S 20793 /15 + 2 (juv.) ; ruins of Milet (= Miletos), UTM NB 25 , 37 .52°N 27.27°E, leg. Puts, 1983, Maa/1 ; Samsun Mountains, Priene near Güllübahçe , above ancient ruins, UTM NB 26 , 37 .65°N 27.29°E, leg. Schlickum, 19.9.1968, SMF 279324 /3 ; leg. Neuteboom 24.4.1989, NNM/1 + 1 (juv.) ; leg. Erőss 28.7.1992, coll Erőss /12 + 2 (juv.) ; leg. Schütt 24.4.2000, (ex coll. Schütt Nr. 1687) S 20794 /22 ; KuşadasI, UTM NB 29 , 37 .85°N 27.26°E, leg. Neuteboom 6.5.1978, NNM/2 (juv.). — Izmir County, Efes (= Ephesos), among ruins, UTM NB 29 , 37 .93°N 27.34°E, leg. Kaysser, 1904, SMF 284902 /1 ; leg. Schlickum 15.9.1968, SMF 279323 /1 ; leg. Papp 10.10.1972, HNHM 33704 /1 ; leg. Seidl 7.5.1975, HNHM 39186 /2, 49552/2 ; leg. Neuteboom 6.5.1978, NNM/11 + 3 (juv.) ; leg. Erőss 30.7.1992, E&F/31 ; leg. Bánki 26.7.1995, HNHM 41708 /3 + 3 (juv.) ; surroundings hous of death of Saint Maria on Ala Dag Mountain (= 5 km S of Efes), UTM NB 29 , 37 .91°N 27.33°E, leg. Seidl 7.5.1975, Maa /5 ; Pamucak, Orman Kampi, UTM NB 29 , 37 .93°N 27.28°E, leg. Bilgin, (ex. coll. Schütt) S 20791 /1 ; surroundings of Izmir (= Smyrna), UTM n. det., leg. Basoglu 2.12.1956, SMF 166916 /4 ; Izmir – Bornova , UTM NC 15 , 38 .47°N 27.24°E, SMF 101956 /5 ; halfway between Izmir and ManIsa, UTM NC 26 , 38 .54°N 27.30°E, leg. Schütt 4.9.2004, (ex coll. Schütt Nr., 1842) S 20792 /1.

Remarks: Lindholmiola lens is very variable in shell shape and size, a fact which caused authors to describe several varietal taxa, which here are all synonymised under L. lens . The nominotypical subspecies is known from the Pelopónnisos and the Ionian islands of Kefallinia and Zakinthos and is characterized by a blunt keel, a narrow and only slightly perspective umbilicus, the moderately enlarged peristome, the medium shell size and narrow whorls. The mountains in the north of the Pelopónnisos are inhabited by a form which differs by shells with slightly broader whorls, an even more blunt keel, and a larger perspective umbilicus (= elia). However, this form also lives on the Island Évvia and close to Neohóri (= NE of Aridéa) in Makedonía. This form is intermediate between the nominotypical subspecies and another, quite remarkable form (= callojuncta). This very distinct form has a shell with a sharp keel, broad whorls, a wide perspective umbilicus, and a surface sculpture of small riblets. Its shells are of medium size (sometimes scalarid shells occur), and the peristomial insertions are connected by a more conspicuous parietal callus. The peristome of this form is wider than in any other form of L. lens . The form callojuncta lives around the Bay of Korinthos, but also in Argolída and in Arkadía.

The eastern part of continental Greece, Atikí, the Island Évvia, Thessalía and the Halkidikí Peninsula is inhabited by a small to medium-sized form (= lentiformis ). It can be characterised by its moderately broad whorls, the blunt keel, and the narrow to slightly perspective umbilicus. A similar but unnamed form inhabits the Dodecanese Islands and the western coast of Turkey. It has a somewhat sharper keel and a wider umbilicus if compared to lentiformis . The form piligera is intermediate between the forms elia and lentiformis . It lives in Viotía, Fthiótida and on the Island Évvia. It differs by a more tightly coiled shell, the whorls which are more rounded (surface and subsurface), and the only slightly perspective umbilicus.

Distribution ( Fig. 15 ): Lindholmiola lens inhabits quite a large area. It is known from the Ionian Islands of Kefallinia and Zakinthos, the Pelopónnisos, the islands of the Aegean Sea to the western part of Turkey. In the north, it is recorded from the Bay of Korinthos, and the eastern part of continental Greece to the western part of the Halkidikí Peninsula.

Figures

Figs 3–6. Lindholmiola lens. Fig. 3: syntype Helix lentiformis, ZMZ 508656, "Thessalie et Attique", D = 11.2 mm; Fig. 4: NMBE 515542, Attika, Methana, 500 m before village, 12.07.1996, leg. E. Neubert, D = 12.77 mm; Fig. 5: syntype Helix lens var. callojuncta, NHMG 372, D = 12.0 mm; Fig. 6: syntype Helix lens var. elia, SMF 101961, Kumani, Elis, coll. Jetschin ex Boettger, 1883, D = 11.94 mm. — All photos Neubert/Bochud, × 3.

Figs 7–11. Lindholmiola lens. Fig. 7: NMBE 515616, Greece, Ilía (= Elis), Alfiós canal between Olimbía and Kréstena (nominotypical form), D = 11.78 mm; Fig. 8: NMBE 515617, Greece, Fokída, Delphi, archaeological site (flat keeled form,broad whorls, wide umbilicus), D = 14.85 mm; Fig. 9: NMBE 515614, Greece, Thessalía, W boundary of Morfovoúni (densely coiled, blunt keel, narrow umbilicus), D = 12.88 mm; Fig. 10: NMBE 515615, Greece, Makedonía, Kassándra Peninsula, Loutrá (small form, blunt keel, moderately wide umbilicus), D = 7.95 mm; Fig. 11: NMBE 515629, Greece, Makedonía, Canyon 2 Km N of Neohóri (= NE of Aridéa) (flat form, blunt keel, wide umbilicus), D= 14.16 mm. — All photos Neubert/Bochud, × 3.

Figs 12–14. 12: Genital organs of Lindholmiola species. Fig. 12: Lindholmiola lens, a: situs, Greece, Ilía (= Elis) between Olimbía and Kréstena, b: lumina of genital organs, Greece, Ahaïa, 3 Km S of Ptéri; Fig. 13: Lindholmiola spectabilis, a: situs, b: lumina of genital organs, Greece, Makedonía, Falakró Mts., Ágios Pnevma plateau; Fig. 14: Lindholmiola girva, a: situs, b: lumina of genital organs, Greece, Makedonía, Taxiárhis.

Fig. 15. Distribution of Lindholmiola lens and L. spectabilis.

Figs 16–18. Lindholmiola barbata. Fig. 16: NMBE 515636, Island Kríti, Nomos Iráklio, valley E from the airport Iráklio (small rounded form), D = 9.66 mm; Fig. 17: NMBE 515635, Island Kríti, Nomos Haniá, canyon ca. 4 Km N of Thériso (large flat form), D = 13.78 mm; Fig. 18: NMBE 515581, Rethimnon, Zousidi, 09.04.1993, leg. N. Chalwatzis, D = 12.49 mm. — All photos Neubert/Bochud, × 3.