Lindholmiola corcyrensis ( ROSSMÄSSLER, 1838 )

Subai, Peter & Neubert, Eike, 2014, Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae), Contributions to Natural History 23, pp. 1-94: 39-56

publication ID

http://doi.org/ 10.5169/seals-787037

persistent identifier

http://treatment.plazi.org/id/038C1808-EF7A-FFE1-59A5-FEAB4D4588BD

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scientific name

Lindholmiola corcyrensis ( ROSSMÄSSLER, 1838 )
status

 

Lindholmiola corcyrensis ( ROSSMÄSSLER, 1838)  

Figs 20–23 View Figs 20–23 , 24–25 View Figs 24–25 , 27 View Figs 26–28 , 29 View Fig

1828 Helix contorta   , – Menke, Synopsis methodica molluscorum generum omnium et specierum earum, quae in Museo Menkeano adservantur: 10 (nomen nudum, homonym of Helix contorta LINNAEUS, 1758   ).

1837 Helix (Caracollina) corcyrensis BECK, Index   molluscorum praesentis aevi musei principis augustissimi Christiani Frederici: 28 (nomen nudum).

1838 Helix (Helix) corcyrensis   , – Anton, Verzeichniss der Conchylien welche sich in der Sammlung von Hermann Eduard Anton befinden: 39, No. 1428 (nomen nudum).

1838 Helix (Helix) canalifera ANTON, Verzeichniss der Conchylien   welche sich in der Sammlung von Hermann Eduard Anton befinden: 39, No. 1427. Locus typicus: "Corfu" (nomen oblitum).

1838 Helix contorta   [ H. corcyrensis   ] ROSSMäSSLER, Iconographie der Land- & Süsswasser-Mollusken mit vorzüglicher Berücksichtigung der europäischen noch nicht abgebildeten Arten, (1) 2 (1/2): 40, Taf. 39, Fig. 538. Locus typicus: "Corfu". Type material: lost [(partim, not the variety minor   = girva   FRIVALDSZKY, 1835) (homonym of Helix contorta LINNAEUS, 1758   )].

1839 Helix corcyrensis   , – Deshayes in Férussac & Deshayes, 1820–1851: Histoire naturelle générale et particulière des mollusques terrestres et fluviatiles, 1: 21 (Taf. 69 E, Fig. 1–5 View Figs View Figs 3–6 published in, 1840).

1846 Helix corcyrensis   , – L. Pfeiffer, In: Martini & Chemnitz, Systematisches Conchylien-Cabinet, (1) 12 (II): 103, Taf. 16 Fig. 23–24 View Figs 20–23 View Figs 24–25 .

1848 Helix corcyrensis   , – L. Pfeiffer, Monographia heliceorum viventium, 1: 415.

1848 Helix corcyrensis   var. γ. major, – L. Pfeiffer, Monographia heliceorum viventium, 1: 415.

1848 Helix canalifera   , – L. Pfeiffer, Monographia heliceorum viventium, 1: 415.

1853 Helix corcyrensis   , – L. Pfeiffer, Monographia heliceorum viventium, 3: 262.

1853 Helix canalifera   , – L. Pfeiffer, Monographia heliceorum viventium, 3: 262.

1854 Helix corcyrensis   , – Reeve, Conchologia Iconica, 7: species 1048.

1859 Helix corcyrensis   , – Mousson, Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich, 4: 17, 256.

1859 Helix corcyrensis var. octogyrata MOUSSON, Vierteljahrsschrift der Naturforschenden Gesellschaft   in Zürich, 4: 257.

1859 Helix corcyrensis var. cefalonica MOUSSON, Vierteljahrsschrift der Naturforschenden Gesellschaft   in Zürich, 4: 29.

1859 Helix corcyrensis var. canalifera   , – Mousson, Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich, 4: 17, 257.

1859 Helix corcyrensis var. girva   , – Mousson, Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich, 4: 268 [non girva FRIVALDSZKY   ].

1859 Helix corcyrensis   , – L. Pfeiffer, Monographia heliceorum viventium, 4: 312.

1859 Helix canalifera   , – L. Pfeiffer, Monographia heliceorum viventium, 4: 312.

1868 Helix corcyrensis   , – L. Pfeiffer, Monographia heliceorum viventium, 5: 415.

1873 Helix corcyrensis var. canalifera   , – Martens, Malakozoologische Blätter, 20: 33.

1876 Helix corcyrensis   , – L. Pfeiffer, Monographia heliceorum viventium, 7: 467.

1879 Helix (Trigonostoma) corcyrensis   , – Westerlund & Blanc, Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 36.

1879 Helix (Trigonostoma) corcyrensis var. cefalonica   , – Westerlund & Blanc, Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 36.

1879 Helix (Trigonostoma) corcyrensis var. octogyrata   , – Westerlund & Blanc, Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 37.

1879 Helix (Trigonostoma) corcyrensis var. canalifera   , – Westerlund & Blanc, Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 37.

1879 Helix (Trigonostoma) corcyrensis var. girva   , – Westerlund & Blanc, Aperçu sur la faune malacologique de la Grèce inclus l’Epire et la Thessalie: 37 [non girva FRIVALDSZKY   ].

1882 Helix corcyrensis   , – Hesse, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 9: 321.

1883 Helix (Anchistoma) corcyrensis   , – O. Boettger, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 10: 316, 331.

1883 Helix (Anchistoma) corcyrensis cefalonica   , – O. Boettger, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 10: 323.

1886 Helix (Gonostoma) corcyrensis var. octogyrata   , – O. Boettger, Jahrbücher der Deutschen Malakozoologischen Gesellschaft, 13: 52.

1887 Helix (Caracolina) corcyrensis   , – Tryon, Manual of Conchology, 3: 117.

1887 Helix (Caracolina) corcyrensis var. cephalonica   , – Tryon, Manual of Conchology, 3: 118.

1887 Helix (Caracolina) corcyrensis var. octogyrata   , – Tryon, Manual of Conchology, 3: 118.

1887 Helix (Caracolina) corcyrensis var. canalifera   , – Tryon, Manual of Conchology, 3: 118.

1889 Helix (Caracolina) corcyrensis   , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 19.

1889 Helix (Caracolina) corcyrensis var. octogyrata   , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 19.

1889 Helix (Caracolina) corcyrensis var. cefalonica   , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 19.

1889 Helix (Caracolina) corcyrensis var. canalifera   , – Westerlund, Fauna der in der paläarctischen Region lebenden Binnenconchylien, 2: 19.

1889 Helix (Caracollina) corcyrensis   , – Martens, Archiv für Naturgeschichte, 55 (1): 172.

1891 Helix (Gonostoma) corcyrensis   , – O. Boettger, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 23 (5/6): 83.

1894 Helix (Gonostoma) corcyrensis   , – Sturany, Annalen des Kaiserlich-Königlichen Naturhistorischen Hofmuseums, 9 (3): 371 [partim].

1894 Helix (Caracollina) corcyrensis   , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) corcyrensis var. cephalonica   , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) corcyrensis var. octogyrata   , – Pilsbry, Manual of Conchology, 9: 288.

1894 Helix (Caracollina) corcyrensis var. canalifera   , – Pilsbry, Manual of Conchology, 9: 288.

1905 Gonostoma corcyrensis   , – Sturany, in Paganetti-Hummler: Verhandlungen der kaiserlich-königlichen Zoologisch-Botanischen Gesellschaft in Wien, 1905: 119.

1915 Helicodonta (Caracollina) contorta   , – Sturany & Wagner, Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 91: 31, Taf. 2, Fig. 7 View Figs 7–11 , a–c.

1918 Caracollina corcyrensis   , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1918 Caracollina corcyrensis octogyrata   , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1918 Caracollina corcyrensis cephalonica   , – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.

1924 Helicodonta (Caracollina) contorta   , – Polinski, Annales zoologici Musei Polonici Historiae Naturalis, 3 (3/4): 130.

1924 Helicodonta (Caracollina) contorta var. canalifera   , – Polinski, Annales zoologici Musei Polonici Historiae Naturalis, 3 (3/4): 130.

1930 Caracollina contorta contorta   , – Käufel, Sitzungsberichte der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 139 (3/4): 183.

1930 Caracollina contortata cephalonica   , – Käufel, Sitzungsberichte der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 139 (3/4): 183.

1931 Lindholmiola corcyrensis   , – Hesse, Zoologica, 31 (81) 1/2: 50.

1936 Lindholmiola corcyrensis   , – Fuchs & Käufel, Archiv für Naturgeschichte, N.F. 5: 651. Fig. 79 [partim, only shells from Podgorica].

1941 Lindholmiola corcyrensis cephalonica   , – Käufel & Fuchs, Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 88/89: 201.

1957 Lindholmiola corcyrensis   , – Jaeckel, Klemm & Meise, Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden, 23 (2): 165 [partim].

1962 Lindholmiola corcyrensis   , – Klemm, Sitzungsberichte der Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abt. I, 171 (6/7): 252.

1962 Lindholmiola corcyrensis octogyrata   , – Klemm, Sitzungsberichte der Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abt. I, 171 (6/7): 253.

1962 Lindholmiola corcyrensis cephalonica   , – Klemm, Sitzungsberichte der Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abt. I, 171 (6/7): 253.

1962 Lindholmiola corcyrensis   , – Jaeckel, Die Tierwelt Mitteleuropas, p.188 (partim).

1962 Lindholmiola corcyrensis   , – Paget, Memorie di Biogeografia Adriatica, 4: 182, 184–185, 195, Fig. 3–5 View Figs 3–6 [shell].

1965 Lindholmiola corcyrensis   , – Forcart, Verhandlungen der Naturforschenden Gesellschaft in Basel, 76 (1): 141.

1971 Lindholmiola corcyrensis   , – Alzona, Atti della Società Italiana di Science Naturali e del Museo Civico di Storia Naturale di Milano, 111: 185.

1973 Lindholmiola (Lindholmiola) corcyrensis cephalonica   , – Hudec & Vašátko, Acta scientiarum naturalium Academiae Scientiarum Bohemoslovacae, 7 (9): 13.

1989 Lindholmiola corcyrensis   , – Reischütz & Stummer, Malakologische Abhandlungen aus dem Staatlichen Museum für Tierkunde Dresden, 14 (12): 106.

1990 Lindholmiola corcyrensis   , – Reischütz & Sattmann, Annalen des Naturhistorischen Museums in Wien, 91 (B): 258.

1990 Lindholmiola girva   , – Fechter & Falkner, Steinbachs Naturführer: 222, Abb. 6 (partim) (non girva FRIVALDSZKY, 1835   ).

1996 Lindholmiola girva   , – Welter-Schultes, Schriften zur Malakozoologie, 9: 23–31 (non girva FRIVALDSZKY, 1835   ).

1996 Lindholmiola girva   , – Dhora & Welter-Schultes, Schriften zur Malakozoologie, 9: 157, 190, Taf. 13, Fig. 165–166 (non girva FRIVALDSZKY, 1835   ).

2002 Lindholmiola girva   , – Dhora, Studime mbi molusqet e Shqipërisë: 99, 156–158 (non girva FRIVALDSZKY, 1835   )

Diagnosis: shell depressed, shell surface covered by granulation, whorls slightly shouldered, penial lumen with a number of small, v-shaped pilasters only.

Description of shell: shell depressed, spire only slightly raised, shell colour light to chestnut brown; first ½ to ¾ protoconch whorl smooth, then with a few fine radial stripes; the following teleoconch whorls with increasingly stronger stripes, but often reduced on the penultimate and last whorls; stripes may vary in strength to even form small riblets; on the subsurface of the shell, stripes usually weak and blurred; complete shell surface covered by a fine granulation giving the shell a dull appearance; sometimes with small hair warts (mainly on subsurface); shell covered by hairs, hairs in the umbilicus usually shorter than those on the rest of the shell, hair length at the shell's shoulder ca 0.4–0.7 mm; 5¾–7¾ regularly increasing, slightly shouldered whorls; last whorl slightly descending before the aperture, then abruptly curved downwards, the bent part in a brighter to white colour; suture deep; umbilicus of oval shape, cylindrical to slightly perspective, only slightly increasing at the beginning, with a rapid, almost double increase in the last whorl, reaching a diameter of 1.4–3.75 mm; aperture oblique, ear-shaped, apertural insertion with a distance of 2.3–5 mm, connected by a very thin, almost indiscernible callus; peristomial rim sharp, forming a strong lateral and weaker basal lip, with a small peristomial callus covering ca 1/10 of the umbilicus.

Measurements: H: 3.75–7.5; D: 7.8–15.9; aH: 3.4–7.2 (2–4.7); aW: 3.6–7.8.

Details of body (8 animals; eastern boundary of Stroúni/1, Island Lefkáda, Evgiros/3, Island Kefallinia, 1.5 km E of Sámi/4): head and dorsal body cream to grey or brown, with many intermediate colourations, becoming brighter in direction to the tail; foot sole cream to beige; mantle with small, elongate grey dots; secondary ureter opens 1–1.5 mm before the pneumostome, where it splits into two canals.

Morphology of the genital organs ( Fig. 27 View Figs 26–28 ): penis short, slightly inflated, ca half of its length is covered by penial glandular tissue, penial lumen with a number of small, v-shaped pilasters; vagina relatively long, glandula narrow, scarcely folded, reaching 2/3 of the length of the bursa copulatrix, pedunculus and vesicle of the bursa copulatrix of the same length; atrial lumen filled by two to three elevated and elongated pilasters with a few short secondary pilasters running in parallel, the major pilasters unite to form a large single pilaster filling almost the complete vaginal lumen.

Differential diagnosis: Conchologically, this species is close to L. pirinensis   . However, it differs from that species by its more flattened whorls, its shallow suture, and its less pronounced surface sculpture. Its radial riblets are quite regular, and its hairs are longer (although less permanent). As in L. pirinensis   , the aperture is only slightly bent downwards, but in frontal view, the aperture of L. pirinensis   is broader and larger. The apertural rim of L. corcyrensis   is only slightly reinforced, and the umbilical shield less pronounced. In L. pirinensis   ( Fig. 26 View Figs 26–28 ), the penial lumen shows only two large pilasters with one of them curved backwards forming a third, smaller pilaster.

Lindholmiola girva   is quite similar, but the shell is smaller on average with more flattened whorls. However, in this species, the teleoconch whorls are more distinctly shouldered with the radial riblets being more pronounced. The last whorl is narrower. The umbilicus formation in L. girva   is similar, but here it is less perspective (almost circular) and not oval as in L. corcyrensis   . It aperture is narrower and sometimes shows a tooth-like swelling. As in L. pirinensis   , L. girva   shows two to three elongated penial pilasters, and thus can easily be distinguished from L. corcyrensis   .

Similarly sized shells of L. barbata   have a half to one additional whorl, L. corcyrensis   is clearly more densely coiled. In addition, the shell of L. barbata   is covered by small hair warts, its umbilicus is partially to completely covered by a columellar shield, and the aperture is less bent downwards with a weaker labial callus. The penial lumen of L. barbata   shows two parallel elongated pilasters and thus differs profoundly from the v-shaped pilasters of L. corcyrensis   .

Type specimens: Greece, Kefallinia Island, "aux environs d’Argostoli,...à Frangata et Grisata", 38.17°N 20.49°E, syntypes var. cefalonica   : ZMZ 508737/6 GoogleMaps   ; "aux environs de Préveza ", 38.96°N 20.75°E, syntypes var. octogyrata   : ZMZ 508745/5. GoogleMaps  

Additional specimens examined:

Croatia, Pelješac–Peninsula   GoogleMaps , sea floating debris near Prapatna, UTM YH 14, 42.81°N 17.67°E, leg. Maassen 4.1989, Maa /6   ; "Dalmatien, Zirona Grande", UTM n. det., SMF 284844 View Materials /2   ; Badija Island (= Sv. Scoglia Badia) near Korčula Island, sea floating debris, UTM XH 75, 42.95°N 17.15°E, SMF 207804 View Materials /1.  

Italy, Prov. Foggia, Promontorio del Gargano, San Menáio, UTM WG 74, 41.93°N 15.94°E, ( Paget, 1962: 182; Forcart, 1965: 141)   ; Promontorio del Gargano, Péschici, UTM WG 84, 41.94°N 16.01°E, ( Forcart, 1965: 141).  

Montenegro, fortress near Vranjina, UTM CM 48, 42.27°N, 19.11°E, leg. Maassen 5.1984, Maa/1   ; Busovnik   GoogleMaps (= Mountain W of Podgorica), UTM CM 49, 42.44°N, 19.14°E, SMF 284837 View Materials /3, 284865/1   ; Ulcinj, UTM CM 54, 42.93°N, 19.21°E, leg. Podani 20.8.1976, HNHM 21472 View Materials /1 (juv.)   ; Podgorica   GoogleMaps (= Titograd), UTM CM 59, 42.44°N, 19.26°E, SMF 101901 View Materials /11   ; Šasko Lake , castle ruins, UTM CM 64, 41.97°N, 19.33°E, leg. Erőss & Fehér leg. 20.4.2000, HNHM 85624 View Materials /8   ; Spuž, UTM CN 50, 42.51°N, 19.19°E, leg. Wohlberedt, 1933, SMF 101900 View Materials /4   ; Zelenik (= Zelnik ), UTM n. det., leg. Wohlberedt, 1905, SMF 284836 View Materials /2, 284866/2.  

Albania, valley of Povlë River S of Greshicë (= 15 + 1 km SE of Ballsh), approx. 140–200 m alt., UTM CK 98, 40.54°N, 19.77°E, leg. Erőss, Fehér & Kovács 14.4.2001, HNHM 85619 View Materials /4   ; Durrës (= Durazzo), UTM CL 67–77, 41.32°N, 19.44°E, SMF 207808 View Materials /3   ; 24.9.1959, HNHM 83641 View Materials /1   ; beach of Durrës, UTM CL 77, 41.31°N, 19.47°E, leg. Clauss, SMF 207805 View Materials /3   ; Krujë , on castle walls, UTM CL 99, 41.5075°N, 19.7933°E, leg. Erőss & Fehér 2.7.1996, HNHM 85135 View Materials /1   ; Shkodër (= Skutari), UTM CM 75, 42.07°N, 19.52°E, 1905, SMF 101928 View Materials /11   ; leg. Barbanic, 1905, SMF 284873 View Materials /15   ; leg. Wohlberedt, 1905, SMF 284863 View Materials /1   ; leg. Flasch, 1906, SMF 284871 View Materials /10   ; leg. Kormos, 1907, SMF 101926 View Materials /4   ; Shkodër, castle hill, UTM CM 75, 42.0475°N, 19.4936°E, leg. Erőss & Fehér 1.7.1996, HNHM 85140 View Materials /1   ; leg. Erőss & Fehér & Kovács 7.4.2001, HNHM 85623 View Materials /3   ; Vau i Dejes (= approx. 20 km SE of Shkodër), UTM CM 85, 42.00°N, 19.63°E, leg. Erőss & Fehér 2.7.1996, HNHM/1 View Materials   ; Pllanë (= N of Milot), 4 km E from bridge of Mat River , approx. 100 m alt., UTM CM 91, 41.70°N, 19.72°E, leg. Erőss & Fehér & Kovács 9.4.2001, HNHM 85620 View Materials /3   ; rock walls at Borsh, UTM DK 03, 40.06°N, 19.85°E, leg. Erőss & Fehér 6.7.1996, HNHM 85142 View Materials /1   ; Tepelenë , on castle walls, UTM DK 16, 40.29°N 20.02°E, leg. Erőss & Fehér 6.7.1996, HNHM 85141 View Materials /1   ; leg. Erőss & Fehér & Kovács 14.4.2001, HNHM 85637 View Materials /1   ; Tepelenë, Lavdi Deshmorevë cemetery on the mountain, approx. 260 m alt., UTM DK 16, 40.29°N 20.02°E, leg. Erőss 17.8.1993, S, 18263/2   ; leg. Erőss & Fehér & Kovács 14.4.2001, HNHM 85628 View Materials /7   ; 4 km S of Tërpan (= 28 km from Berat direction Këlcyrë ), approx. 700 m alt., UTM DK, 18, 40.54°N 20.00°E, leg. Erőss & Fehér & Kovács 13.4.2001, HNHM 85625 View Materials /3   ; 4 km N of Tërpan (= 20 km from Berat direction Këlcyrë ), approx. 700 m alt., 40.57°N 20.01°E, UTM DK, 19, leg. Erőss & Fehér & Kovács 13.4.2001, HNHM 85636 View Materials /2   ; left bank of Dëshnicë River 5 km N of Këlcyrë, approx, 270 m alt., UTM DK 26, 40.35°N 20.16°E, leg. Erőss & Fehér & Kovács 13.4.2001, HNHM 85621 View Materials /8   ; Osum Gorge 4 km SE of Çorovodë (= direction Zogas), approx. 400 m alt., UTM DK 38, 40.47°N 20.25°E, leg. Erőss & Fehér & Kovács 12.4.2001, HNHM 85622 View Materials /1   ; Mal e Melesinit (= 6 km S of Leskovik), UTM DK 64, 40.10°N 20.59°E, leg. Erőss & Fehér 7.7.1996, HNHM 85132 View Materials /1   ; Leskovik (= Lescovic ), 800 m alt., UTM DK 64, 40.14°N 20.59°E, SMF 207807 View Materials /1 + 1 (juv.)   ; Mal i Dajtit , path direction relais station, on rocks, approx. 1200 m alt., UTM DL 07, 41.36°N, 19.91°E, leg. Erőss & Fehér 4.7.1996, HNHM 85131 View Materials /1   ; Mal i Dajtit , 1 km above Linzë, UTM DL 07, 41.36°N, 19.88°E, leg. Fehér 27.7.1993, HNHM 85136 View Materials /4   ; Tiranë River Valley 1 km N from Turkish bridge between Ferraj and Shtish, UTM DL 08, 41.39°N, 19.86°E, leg. Erőss & Fehér & Kovács 10.4.2001, HNHM 85630 View Materials /3   ; Tiranë River Valley 4 km N from Turkish bridge between Ferraj and Shtish, UTM DL 08, 41.43°N, 19.85°E, leg. Erőss & Fehér & Kovács 10.4.2001, HNHM 85633 View Materials /2   ; Berat, castle hill SW slope, approx. 100 m alt., UTM DL 10, 40.70°N, 19.94°E, leg. Erőss & Fehér & Kovács 12.4.2001, HNHM 85627 View Materials /7   ; Berat, left bank of Osum River opposite castle hill, approx. 100 m alt., UTM DL 10, 40.70°N, 19.94°E, leg. Erőss & Fehér & Kovács 13.4.2001, HNHM 85626 View Materials /5   ; S of Gracen (= Elbasan – Tiranë country road, 7 km N from bifurcation to Cërrik), approx. 500 m alt., UTM DL 15, 41.14°N, 19.96°E, leg. Erőss & Fehér & Kovács 14.4.2001, HNHM 85632 View Materials /2   ; Erzen Gorge NW of Ibë, approx. 300 m alt, UTM DL 16, 41.25°N, 19.91°E, leg. Erőss & Fehér & Kovács 11.4.2001, HNHM 85635 View Materials /2   ; hills S of Ibë , UTM DL 16, 41.23°N, 19.93°E, leg. Erőss & Fehér 15.9.1994, HNHM 85137 View Materials /2   ; Fangul Pass (= 9 km from Ibë direction Kllojkë), approx. 700 m alt., UTM DL 16, 41.27°N 20.05°E, leg. Erőss & Fehér & Kovács 11.4.2001, HNHM 85631 View Materials /1   ; Elbasan, UTM DL 25, 41.11°N 20.09°E, leg. Fehér 23.7.1993, HNHM 85039 View Materials /1   ; Mirakë, Ura e Kamarës , left bank of Shkumbin River , approx. 210 m alt., UTM DL 36, 41.16°N 20.24°E, leg. Erőss & Fehér & Kovács 14.4.2001, HNHM 85629 View Materials /1   ; on rocks above Tushemist at Ohrid Lake , UTM DL 72, 40.90°N 20.72°E, leg. Erőss & Fehér 7.7.1996, HNHM 85138 View Materials /1   ; Pustec, bank of Prespa Lake , UTM DL 91, 40.79°N 20.91°E, leg. Fehér, 18.9.1992, HNHM 85139 View Materials /1   ; Djelcic, UTM n. det., SMF 284845 View Materials /1.  

Fyrom, Crni Drim River left bank, N of Lukovo, on limestone rocksand in the soil, 630 m alt., UTM DL 67, 41.3544°N 20.6107°E, leg. Pintér, E. & P. Subai & Szigethy 17.07.1972, HNHM 49545 View Materials /1 (juv.)   ; Crni Drim River left bank, some km S of Lukovo, 630 m alt., UTM DL 67, 41.3398°N 20.6282°E, leg. Pintér, E. & P. Subai & Szigethy 17.07.1972, HNHM 49543 View Materials /1, 85618/1, S 3848/1   ; 1 km SE of Džepišta (= direction Lukovo   GoogleMaps ), 700 m alt., UTM DL 68, 41.4432°N 20.5374°E, leg. Pintér, E. & P. Subai & Szigethy 17.07.1972, HNHM 49544 View Materials /1, S 3847/1   ; Sv.Naum, under stones, UTM DL 72, 40.9141°N 20.7412°E, leg. Maassen 5.1976, Maa/27, NNM/3 View Materials   ; Galičica Mountains   GoogleMaps , in forest 10 km from the crossing Carina– Sv. Naum – Ohrid direction Carina, on limestone rocks, 1560 m alt., UTM DL 83, 40.9618°N 20.8165°E, leg. Pintér, E. & P. Subai & Szigethy 14.07.1972, HNHM 49542 View Materials /1 (juv.), S 3845/1   ; Galičica Mountains   GoogleMaps , pass 17 km from bifurcation at Trpejca   GoogleMaps direction Oteševo, 1600 m alt., UTM DL 83, 40.9553°N 20.8146°E, leg. Maassen 5.1976, Maa/16   ; at boat harbour of Trpejca   GoogleMaps , on limestone rocksand from Lake Ohrid, 700 m alt., UTM DL 83, 40.9584°N 20.7781°E, leg. Pintér, E. & P. Subai & Szigethy 16.07.1972, HNHM 49533 View Materials /5 + 4 (juv.+ fragment), 85634/1, S 3841/6 + 1 (juv.)   ; rocks N of boat harbour of Trpejca   GoogleMaps , UTM DL 83, 40.9591°N 20.7781°E, leg. Maassen 5.1975, Maa/1   ; Sv.Naum – Ohrid highway, bifurcation at Trpejca   GoogleMaps , UTM DL 83, 40.9439°N 20.7895°E, leg. Kiss & Pintér 4.7.1985, HNHM 30854 View Materials /1 + 4 (juv. + fragment)   ; Trpejca   GoogleMaps , UTM DL 83, 40.96°N 20.78°E, leg. Kiss & Pintér 5.7.1985, HNHM 30855 View Materials /1   ; between Trpejca and Peštani, UTM DL 83, 40.98°N 20.80°E, leg. Kiss & Pintér 5.7.1985, HNHM 30856 View Materials /1   ; on limestone rocks at shore of Lake Ohrid approx. 2.5 km S of Peštani, 710 m alt., UTM DL 83, 40.9981°N 20.8007°E, leg. Pintér, E. & P. Subai & Szigethy 16.07.1972, HNHM 49531 View Materials /1 + 4 (juv.), S 3840/3   ; leg. Maassen 5.1975, Maa/1   ; 1 km S of Peštani, 800–1000 m alt., UTM DL 83, 40.9980°N 20.8025°E, leg. Maassen 5.1975, Maa/10   ; bank of Ohrid Lake S of Peštani, on limestone rocks, UTM DL 83, 41.0065°N 20.8050°E, leg. Pintér, E. & P. Subai & Szigethy 16.07.1972, HNHM 49534 View Materials /1 + 1 (juv.), S 3843/3   ; Galičica Mountains   GoogleMaps W side, on limestone rocks in "Elešec" camping, 4 km N of Peštani and of Lake Ohrid before the camping, 750 m alt., UTM DL 84, 41.0386°N 20.8079°E, leg. Subai 11.–14.08.1971, S 3846/2   ; leg. Pintér, Subai & Szigethy 15.7.1972, HNHM 49532 View Materials /3 + 4 (juv.), S 3844/2   ; above Sveti Stefan (= S of Ohrid), on rocks, 1100 m alt., UTM DL 84, 41.0757°N 20.8289°E, leg. Maassen 5.1975, Maa /2   ; tunnel near Orce Nikolov (= 6.5 km S of Ohrid), UTM DL 84, 41.05°N 20.80°E, leg. Maassen 5.1975, Maa /1   ; mountain slope at E border of Ohrid, UTM DL 85, 41.13°N 20.85°E, leg. Mikuska & Sipos 2.10.1975, HNHM 49549 View Materials /1 (juv.).  

Greece, Macedonía, 300 m N of Pefkófito , moist place in a small river gorge, approx. 1000 m alt., UTM DK 96, 40.2972°N 20.9566°E, leg. Subai 10.05.1995, S 15826 View Materials /1   ; E slope of the mountain W of Pefkófito , on dry lime rocks, about 1200–1400 m alt., UTM DK 96, 40.2967°N 20.9433°E, leg. Subai 10.05.1995, S 15823 View Materials /2   ; mountain E slope between Epáno and Káto Aréna (= W of Pefkófito), on limestone rocks, 1650 m alt., UTM DK 96, 40.2948°N 20.9381°E, leg. Subai 11.05.1995, S 15840 View Materials /5   ; Grámmos Mountains, E side of Epáno Aréna mountain (= NW of Pefkófito), near the crest, 2000 m alt., UTM DK 96, 40.3455°N 20.8621°E, leg. Subai 15.07.1990, S 15872 View Materials /4   ; Epáno Aréna Mountain (= NW of Pefkófito), mountain slope and ridge, on limestone rocks and under stones, 1750–2100 m alt., UTM DK 96, 40.3140°N 20.8972°E, leg. Subai & Szekeres 17.05.1997, S 15848 View Materials /1   ; 15 km SW von Grevená, Landstrasse Zákas–Spíleo, an Kalkfelsen, 925 m alt., UTM EK 23, 40.0104°N 21.2832°E, leg. Gittenberger & Uit de Weerd 11.5.2000, NNM/4 View Materials   ; Kastoriá , N part of peninsula, 620 m alt., UTM EK 28, 40.52°N 21.27°E, leg. Neuteboom 28.7.1979, NNM/1 View Materials   ; Kastoriá, Monastir Panagia Mavriotissa , UTM EK 28, 40.5043°N 21.2804°E, leg. Neuteboom 28.7.1979, NNM/4 View Materials   ; on the peninsula in Kastoriá , on limestone rocks approx. 50–100 m W of Monastir Panagia Mavriotissa, 640 m alt., UTM EK 28, 40.5050°N 21.2786°E, leg. Subai 14.07.1990, S 15877 View Materials /46   ; at bridge 4 km SE of Vogatsikó , on limestone rocks, 680 m alt., UTM EK 37, 40.3983°N 21.4143°E, leg. Subai 10.05.1995, S 15824 View Materials /28   ; in valley W of Gérmas , on limestone rocks, 850 m alt., UTM EK 37, 40.4528°N 21.4149°E, leg. Subai & Szekeres 17.05.1997, S 15849 View Materials /51   ; Mikrá Prespa Lake , behind the biological station, approx. 200 m W of the W border of Mikrolímni, on limestone rocksand in a stone quarry, 885 m alt., UTM EL 01, 40.7419°N 21.1078°E, leg. Subai 13.07.1990, S 15878 View Materials /2   ; leg. Subai leg. 10.5.1995, S 15825 View Materials /2. —   Thessalía, 3–4 km from Diáva towards Koromiliá (= 4 km SW of Kalambáka), additional 400 m on a right path from the bifurcation, on limestone rocks, 550 m alt., UTM EJ 49, 39.68°N 21.58°E, leg. Gittenberger & Uit de Weerd   GoogleMaps 13.5.2000, NNM/3 View Materials   ; NW border of Píli (= 18 km SW of Tríkala), on rocks at both sides of the Turkish arched bridge, 240 m alt., UTM EJ 56, 39.4597°N 21.6121°E, leg. Pintér, E. & P. Subai 28.07.1976, S 3812/1   ; Theópetra (= Kuvelci , 6 km SSE of Kalambáka), on limestone blocks of rocks at river bank, 260 m alt., UTM EJ 59, 39.6798°N 21.6810°E, leg. E. & P. Subai 26.07.1975, S 3813/1   ; Kalambáka, on conglomerate rocks beside the highway from Kalambáka to the Meteóra monasteries, 280 m alt., UTM EJ 59, 39.7103°N 21.6164°E, leg. Pintér, E. & P. Subai 29.07.1976, S 3814/1   ; Kastráki near Kalambáka   GoogleMaps , UTM EJ 59, 39.71°N 21.62°E, leg. Neuteboom leg. 25.7.1979, NNM/2 View Materials   ; at S border of Dafnospiliá , on limestone rocks beside the cave, 260 m alt., UTM EJ 84, 39.2274°N 21.9427°E, leg. Subai & Szekeres 12.05.1997, S 15814 View Materials /2. —   Kérkira Island (= Korfu), Paleokastrítsa , limestone rocks on elevations, 30 m alt., UTM CJ 89, 39.6708°N, 19.7019°E, leg. E. & P. Subai, 18.07.1975, S 3822/5   ; Neuteboom 6.10.1991, NNM/10 View Materials + 1 (juv.)   ; Paleokastrítsa, Villa Reale, UTM n. det., SMF 284829 View Materials /5   ; Pélekas (= Pelleka), UTM CJ 98, 39.59°N, 19.82°E, SMF 101933/2   ; on limestone rocks at S border of Ermónes bay, in rock debris, UTM CJ 98, 39.6086°N, 19.7758°E, leg. Subai 03.08.1995, S 15843 View Materials /3   ; Vátos, bei Monastir Mirtiotíssa, UTM CJ 98, 39.5972°N, 19.7936°E, leg. Neuteboom 2.10.1991, NNM/1 View Materials   ; Rand von Sokráki in Richtung Zigós , UTM CJ 99, 39.7168°N, 19.7977°E, leg. Neuteboom 3.10.1991, NNM/4 View Materials + 4 (juv.)   ; Auffahrtweg zum Pantokrátor, 700 m alt., UTM CJ 99, 39.7499°N, 19.8428°E, leg. Neuteboom 3.10.1991, NNM/1 View Materials   ; Áno Korakiana, UTM CJ 99, 39.70°N, 19.79°E, leg. Neuteboom 3.10.1991, NNM/1 View Materials (juv.)   ; mountain approx. 1 km NW of Strinilas , on limestone rocks, 580 m alt., UTM CJ 99, 39.7464°N, 19.8294°E, leg. Subai 07.08.1995, S 15841 View Materials /3   ; Análipsis (= Analepsis), UTM CJ 99, 39.70°N, 19.82°E, SMF 284827 View Materials /1   ; Ágios Márkos, UTM CJ 99, 39.71°N, 19.82°E, leg. Maassen 3.7.1980, Maa /1   ; 500 m E of Troumpéta (= Pandeleimonas ), N exposed rock holes at border of highway and on S exposed limestone rocks, 460 m alt., UTM CJ 99, 39.7092°N, 19.7531°E, leg. Subai 06.08.1995, S 14275 View Materials /1 (juv.)   ; in olive grove 1 km N of Nímfes , on limestone rocks beside the highway, 150 m alt., UTM CK 90, 39.7609°N, 19.7829°E, leg. Subai 01.08.1995, S 15837 View Materials /31   ; 1.2 km S of Ág. Panteleimonas (= direction Episkepsis ), on limestone rocks beside the highway, 240 m alt., UTM CK 90, 39.7734°N, 19.8267°E, leg. Subai 01.08.1995, S 15836 View Materials /8   ; from Ágios Martínos direction Acharávi   GoogleMaps , UTM CK 90, 39.78°N, 19.83°E, leg. Neuteboom 3.10.1991, NNM/1 View Materials   ; near road between Xathátes and Nímfes , UTM CK 90, 39.75°N, 19.77°E, leg. Neuteboom 4.10.1991, NNM/1 View Materials (juv.)   ; 1.2 km ESE of Akharave , on corrugated karst rocks, 110 m alt., UTM CK 90, 39.7905°N, 19.8336°E, Subai 30.07.1995, S 15844 View Materials /25   ; Ahillio, UTM DJ 07, 39.56°N, 19.90°E, SMF 101932 View Materials /1   ; gegenüber Ahillio, UTM DJ 07, 39.56°N, 19.90°E, SMF 101930 View Materials /2   ; Gardíki, byzantinische Festung, UTM DJ 07, 39.4770°N, 19.8848°E, leg. Neuteboom 12.10.1991, NNM/1 View Materials + 2 (juv.)   ; 1.5 km N von Benítses , UTM DJ 07, 39.55°N, 19.91°E, leg. Gittenberger 9.1996, NNM/2 View Materials (alk.)   ; 2 km S of Benitzes , on rocks in the vicinity of the sea, 20 m alt., UTM DJ 07, 39.5309°N, 19.9203°E, leg. E. & P. Subai, 19.07.1975, S 3818/5   ; "Santi Deka", UTM DJ 07, 39.55°N, 19.88°E, SMF 101934 View Materials /2   ; Ag. Déka Mountain (= W of Ag. Déka Village), on limestone rocks and under stones, 290 m alt., UTM DJ 07, 39.5548°N, 19.8819°E, leg. E. & P. Subai, 19.07.1975, S 3817/5   ; Kérkira Town (= Korfu), UTM DJ 08, 39.62°N, 19.91°E, HNHM 49548 View Materials /1, SMF 284821 View Materials /11, 284822/2, 284823/5, 284824/3, 284825/1, 284826/1, 284828/8 (juv.), 284830/1, 284840/12, 284841/9, 285672/2, 285673/1   ; Brömme leg., SMF 284848 View Materials /1   ; leg. C. R. Boettger, 1906, SMF 101941 View Materials /6   ; leg. Conéménos, SMF 101937 View Materials /5   ; Krüper, 1883, SMF 101939 View Materials /1   ; Stadt Kérkira , Fort (= Fortezza), UTM DJ 08, 39.62°N, 19.91°E, SMF 101931 View Materials /2   ; Klosterruine bei Gouviá, UTM DJ 08, 39.65°N, 19.83°E, SMF 209373 View Materials /5   ; Kanóni (= " Al Kanone "), UTM DJ 08, 39.59°N, 19.92°E, SMF 284842 View Materials /8   ; "Mon Repos", UTM DJ 08, 39.60°N, 19.92°E, SMF 101935 View Materials /1   ; between Dassia and Káto Korakiána , among roots of olive trees, 40 m alt., UTM DJ 09, 39.6830°N, 19.8314°E, leg. E. & P. Subai, 18.07.1975, S 3821/23   ; Pantokrátor [Mountain], UTM DJ 09, DK 00, 39.74°N, 19.87°E, leg. Brömme, SMF 284861 View Materials /1   ; Pantokrátor mountain slope between Barbáti and Nissáki, among rock debris, 200 m alt., UTM DJ 09, 39.7249°N, 19.8776°E, leg. E. & P. Subai 20.07.1975, S 3819/4   ; Pantokrátor S slope above Nissáki , approx. 100 m W of Katsoúri cave, 200–250 m alt., UTM DJ 09, 39.7299°N, 19.8975°E, leg. Subai 06.08.1995, S 14276 View Materials /7   ; S border of road 1.5 km from Petália direction peak of Pantokrátor , limestone rocks, 720 m alt., UTM DJ 09, 39.7521°N, 19.8452°E, leg. Subai 05.08.1995, S 14250 View Materials /3 (juv.)   ; mountain SE of Loútses , on broken karst rocks, 550 m alt., UTM DK 00, 39.7722°N, 19.8928°E, leg. Subai 02.08.1995, S 15842 View Materials /3   ; Pantokrátor Mountain slope between Petália and Láfki, on limestone rocks, 480 m alt., UTM DK 00, 39.7601°N, 19.8325°E, leg. E. & P. Subai 07.08.1975, S 3820/9   ; E slope of Pantokrátor karst plateau, 700 m alt., UTM DK 00, 39.74°N, 19.88°E, leg. Gittenberger 10.1978, NNM/1 View Materials (alk.). —   Paxí Island (= Paxos), UTM n. det., SMF 284832 View Materials /2   ; Gaïou (= Gaion), UTM DJ 23, 39.19°N 20.18°E, leg. K. L. Pfeiffer 4.1932, SMF 101911 View Materials /11   ; rocks between Gaïou and Gaïou port, UTM DJ 23, 39.19°N 20.18°E, leg. Megens & Moorsel 14.5.2000, NNM/3 View Materials . —   Lefkáda Island (= Lefkas ), between road and Evgiros (= 25 km S Lefkáda), 150–175 m alt., UTM DH 67, 38.62°N 20.65°E, leg. Gittenberger 21.10.1981, NNM/4 View Materials (alk.)   ; Megáli Óros plateau, UTM DH 68, 38.72°N 20.62°E, SMF 279322 View Materials /4   ; S border of Sivros , on limestone rocks along border of road, 170 m alt., UTM DH 68, 38.6668°N 20.6433°E, leg. E. & P. Subai 03.08.1975, S 3802/59   ; valley at Stavrotas Mountain (= W of Sivros), on limestone rocks, 200 m alt., UTM DH 68, 38.6704°N 20.6373°E, leg. E. & P. Subai 03.08.1975, S 3804/25   ; 7 km SE of Sivros (= direction Póros ), on rocks beside a small chapel, 320 m alt., UTM DH 77, 38.6524°N 20.6850°E, leg. E. & P. Subai 03.08.1975, S 3806/2   ; above Póros , on limestone rocks, 300 m alt., UTM DH 77, 38.6419°N 20.7089°E, leg. E. & P. Subai 03.08.1975, S 3807/2   ; leg. Pintér & Subai 30.7.1976, S 3808/7   ; mountain slope 500–1000 m S of Nídri , under stones, 20 m alt., UTM DH 78, 38.6915°N 20.6979°E, leg. E. & P. Subai 03.08.1975, S 3805/43   ; 2 km from Kariá direction Lefkáda , on limestone rocks beside the highway, 440 m alt., UTM DH 79, 38.7756°N 20.6554°E, leg. E. & P. Subai 05.08.1975, S 3803/11   ; Fríni (= 2.5 km W of Lefkáda Town), UTM DH 79, 38.82°N 20.68°E, SMF 279321 View Materials /6   ; leg. Gittenberger 22.10.1981, NNM/2 View Materials (alk.)   ; border of Frini (= direction Tsoukaládes), on limestone rocks beside asphalt road, 140 m alt., UTM approx. DH 69, 38.8236°N 20.6800°E, leg. E. & P. Subai 05.08.1975, S 6677/38   ; Kaligóni , between roots of olive trees, UTM DH 79, 38.8148°N 20.7124°E, leg. E. & P. Subai 31.07.1975, S 3811/15   ; Kaligóni , from the spring "Megali Vrysi" and on limestone rocks beside the chapel, UTM DH 79, 38.8147°N 20.7119°E, leg. E. & P. Subai 01.08.1975, S 3809/21, 3810/8. —   Kefaloniá Island , UTM n. det., SMF 101910 View Materials /4, 284857/1   ; leg. Heldreich leg., SMF 284858 View Materials /1   ; 6 km WNW of Lixoúri , UTM DH 43, leg. Visser 4.11.1985, NNM/1 View Materials (alk.)   ; 1.5 km E of Sámi , UTM DH 73, leg. Visser 6.11.1985, NNM/5 View Materials (alk.). —   Ípiros (= Epirus), S border of Igoumenítsa , on limestone rocks and from a spring, 40 m alt., UTM DJ 37, 39.4858°N 20.2818°E, leg. E. & P. Subai 21–22.07.1975, S 3828/1   ; above Pléssio , on lime conglomerate rocks, 450 m alt., UTM DJ 38, 39.6493°N 20.2811°E, leg. Subai 13.05.1995, S 15832 View Materials /2   ; mountain slope approx. 2 km N of Kestríni (= NW of Igoumenítsa ), on limestone rocks, 20 m alt., UTM DJ 38, 39.5745°N 20.1943°E, leg. Subai, 19.07.1990, S 15876 View Materials /4   ; S border of Kótsika , on limestone rocks, under stones, 300 m alt., UTM DJ 39, 39.6592°N 20.2471°E, leg. Subai 13.05.1995, S 15834 View Materials /4   ; Párga, Valtos Beach , on rocks near castle, UTM DJ 44, 39.2832°N 20.3970°E, leg. Megens & Moorsel 14.5.2000, NNM/1 View Materials   ; 1.4 km N of Agiá (= N of Párga ), on dry limestone rocks, 300 m alt., UTM DJ 45, 39.3202°N 20.3443°E, leg. Subai 15.05.1995, S 15830 View Materials /1   ; Párga–Agiá country road, 600 m N of Anthoūssa , UTM DJ 45, 39.3044°N 20.3679°E, leg. Hoenselaar 8.6.1996, Maa /2   ; Nekromanteion (= Ephyra ), UTM n. det., leg. Hoenselaar 9.6.1996, Maa /2   ; 1 km N of Kartéri (= highway Margaríti–Igoumenítsa ), on limestone rocks, 170 m alt., UTM DJ 46, 39.3953°N 20.3702°E, leg. Subai 20.07.1990, S 15829 View Materials /2   ; leg. Subai & Szekeres 14–15.5.1997, S 15850 View Materials /3   ; 5 km E of Igoumenítsa, mountain slope between Krióvrisi and Lákka Villages , under stones, 370 m alt., UTM DJ 47, 39.4882°N 20.3274°E, leg. E. & P. Subai 22.07.1975, S 3837/7   ; 2 km from Parapótamos direction Ioánina (= highway Igoumenítsa–Ioánina ), on limestone rocks, 200 m alt., UTM DJ 47, 39.5297°N 20.3426°E, leg. E. & P. Subai 23.07.1975, S 3836/4   ; cave chapel 5 km from highway Filiátes–Igoumenítsa, direction Mavronéri , on limestone rocks, approx. 150–200 m alt., UTM DJ 48, 39.6434°N 20.3125°E, leg. Subai, 19.07.1990, S 15875 View Materials /1   ; leg. Subai 13.5.1995, S 15833 View Materials /4   ; leg. Subai & Szekeres 15.5.1997, S 15852 View Materials /4   ; about 5 km SE of the highway Filiátes–Igoumenítsa (= direction Kipárisso, Paleohóri ), approx. 100 m W of the bifurcation to Astós, beside the Thíamis river , on limestone rocks, 50 m alt., UTM DJ 48, 39.5673°N 20.3700°E, leg. Subai & Szekeres 15.05.1997, S 15853 View Materials /3   ; 9.5 km from the highway Filiátes–Igoumenítsa direction Keramitsa , 900 m from the bifurcation to Rizó (= 1 km E of Dafni), on limestone rocks in the street bend, 300 m alt., UTM DJ 48, 39.6429°N 20.4095°E, leg. Subai & Szekeres 15.05.1997, S 15851 View Materials /3   ; bifurcation of highway Filiátes–Keramítsa direction Rizó , from here still 2 km eastwards in a river bed, on limestone rocks, 250 m alt., UTM DJ 48, 39.6435°N 20.4164°E, leg. Subai, 19.07.1990, S 15874 View Materials /6   ; S of Mórfi, junction to Párga , UTM DJ 54, 39.2958°N 20.4779°E, leg. Eröss 10.8.1977, HNHM 9003 View Materials /4   ; 500 m from Ag. Kiriáki direction Párga (= bifurcation of highway Préveza–Igoumenítsa direction Párga ), dry hill slope, on rocks, 160 m alt., UTM DJ 54, 39.2900°N 20.4415°E, leg. E. & P. Subai 06.08.1975, S 3830/5   ; 13 km from Vrossína direction Igoumenítsa, E slope of limestone mountain, 650 m alt., UTM DJ 58, 39.5944°N 20.4918°E, leg. Riedel & Subai 13.04.1988, S 15820 View Materials /3   ; N of Glikí, along highway to Koukoulií , on limestone rocks, approx. 160 m alt., UTM DJ 65, 39.3322°N 20.6030°E, leg. Subai 15.05.1995, S 15831 View Materials /1   ; small chapel at N border of Tsangári , on limestone rocks, 480 m alt., UTM DJ 66, 39.4202°N 20.6127°E, leg. Subai 15.05.1995, S 15827 View Materials /3   ; river gorge at S border of Dovlá , on limestone rocks, 400 m alt., UTM DJ 68, 39.5679°N 20.5481°E, leg. Subai 13.05.1995, S 15835 View Materials /1   ; Préveza , UTM DJ 71, SMF 284846 View Materials /1, 284855/3   ; leg. Conéménos, 1883, SMF 284850 View Materials /4   ; leg. Conéménos, 1899, SMF 101912 View Materials /7   ; Camping Kalamitsi Beach , 3 km N of Préveza, UTM DJ 71, 38.98°N 20.70°E, leg. Hoenselaar 4.6.1996, Maa /5   ; wall remains of Nikopolis (= N of Préveza), 20 m alt., UTM DJ 71, 39.0235°N 20.7367°E, leg. E. & P. Subai 06.08.1975, S 3831/2   ; along old highway Préveza–Igoumenítsa, mountain slope 500 m from road junction to Zálongo direction Igoumenítsa , on limestone rocks, 90 m alt., UTM DJ 73, 39.1248°N 20.6568°E, leg. E. & P. Subai 06.08.1975, S 3829/7   ; on high rocks nearly monuments of Zálongo and on the mountain behind Moni Ágios Dimítrios (= 21.5 km NNW of Préveza), 650–725 m alt., UTM DJ 73, 39.14°N 20.68°E, leg. Gittenberger & Uit de Weerd   GoogleMaps 15.5.2000, NNM/1 View Materials   ; Skafidáki Village near Préveza and at other localities on Skafidáki Peninsula, UTM DJ 81, 38.9877°N 20.7830°E, leg. Pintér, E. & P. Subai 31.07.1976, S 3824/2   ; 2 km along the highway Ioánina–Árta opposite the rock hole near Ágios Geórgios , 130 m alt., UTM DJ 84, 39.2723°N 20.8446°E, leg. Fauer & Subai 28.09.1989, S 15867 View Materials /56   ; on limestone rocks along border of road in Nikolítsi Village , 210 m alt., UTM DJ 85, 39.3037°N 20.7676°E, leg. Subai 15.05.1995, S 15828 View Materials /4   ; gorge 5 km N of Térovo (= E of highway Ioánina–Árta ), on limestone rocks, 430 m alt., UTM DJ 86, 39.4497°N 20.8790°E, leg. E. & P. Subai 30.07.1975, S 3827/1   ; bifurcation of the highway Ioánina–Árta to Térovo , on limestone rocks, 285 m alt., UTM DJ 86, 39.4161°N 20.8459°E, leg. Fauer & Subai 28.09.1989, S 15863 View Materials /1   ; ruins of the antic amphitheater near Dodoni (= SSW of lake Ioánina), 640 m alt., UTM DJ 87, 39.5461°N 20.7879°E, leg. E. & P. Subai 30.07.1975, S 3832/9   ; leg. Neuteboom 26.7.1979, NNM/16 View Materials + 1 (juv.)   ; Ioánina, UTM DJ 89, 39.66°N 20.84°E, HNHM 40244 View Materials /1   ; Kria (= N of Pérama near Ioánina ), lime rocks above the upper (= southeastern) cemetery, 610 m alt., UTM DJ 89, 39.7226°N 20.8409°E, leg. Fauer & Subai 27.09.1989, S 15857 View Materials /49   ; E border of Amfithea (= Stroúni , E of Pérama), on limestone rocks, 480 m alt., UTM DJ 89, 39.6860°N 20.8727°E, leg. E. & P. Subai 24.07.1975, S 6676/27 + 1 (alk.)   ; leg. Pintér & Subai 30.7.1976, S 3833/2   ; 2 km E of Amfithea (= Stroúni, E of Pérama ), on limestone rocks, 480 m alt., UTM DJ 89, 39.6802°N 20.8871°E, leg. E. & P. Subai 24.07.1975, S 3834/2   ; NE border of Pérama , on limestone rocks behind the houses, 490 m alt., UTM DJ 89, 39.6986°N 20.8482°E, leg. Fauer & Subai 28.09.1989, S 15862 View Materials /39   ; leg. Subai 12.5.1995, S 15845 View Materials /34   ; Pérama , on limestone rocks at exit of the stalactite cave, 500 m alt., UTM DJ 89, 39.6949°N 20.8457°E, leg. E. & P. Subai 29.07.1975, S 3823/7, 3835/6   ; leg. Pintér & Subai 30.7.1976, HNHM 41385 View Materials /15, 49537/4, S 3825/36   ; leg. Neuteboom 27.7.1979, NNM/4 View Materials + 2 (juv.)   ; leg. Gittenberger & Uit de Weerd 14.5.2000, NNM/3 View Materials   ; road between Amótopos and Nisista , UTM DJ 94, 39.27°N 20.97°E, leg. Neuteboom 26.7.1979, NNM/6 View Materials   ; Velás monastery (= approx. 2 km S of Kalpáki), on limestone rocks above the spring and in a small cave, 450 m alt., UTM DK 61, 39.8705°N 20.6253°E, leg. Fauer & Subai 27.09.1989, S 15858 View Materials /2   ; leg. Subai 16.7.1990, S 15871 View Materials /4   ; Kalpáki   GoogleMaps , UTM DK 61, 39.88°N 20.62°E, leg. Neuteboom 27.7.1979, NNM/1 View Materials   ; 1 km NW of Asfáka , UTM DK 70, 39.78°N 20.72°E, leg. Kiss & Pintér 25.7.1986, S 29880 View Materials /11 + 5 (juv.)   ; Kalpáki–Monodéndri highway, dry lime rocks at road junction to Elafótopos , 860 m alt., UTM DK 71, 39.8753°N 20.6951°E, leg. E. & P. Subai 30.07.1975, S 3826/1   ; Arísti , UTM DK 72, 39.93°N 20.67°E, leg. Neuteboom 27.7.1979, NNM/1 View Materials   ; gorge entrance at bridge across Sarantáporos River between Exohí and Amárantos, dry rock slope, 445 m alt., UTM DK 74, 40.1114°N 20.7218°E, leg. Subai 16.07.1990, S 15869 View Materials /24   ; leg. Subai & Szekeres 16.5.1997, S 15846 View Materials /1   ; Loutrá (= 5 km N of Amárantos), W slope of limestone mountain, 1270 m alt., UTM DK 74, 40.1745°N 20.7297°E, leg. Riedel & Subai 12.04.1988, S 15821 View Materials /1   ; Loutrá (= N of Amárantos), limestone mountain SW side, 1120 m alt., UTM DK 74, 40.1676°N 20.7321°E, leg. Fauer & Subai 25.09.1989, S 15860 View Materials /2   ; mountain E of Loutrá near Amárantos , on N exposed limestone rocks and in forest approx. 500 m SE of Loutra, 1250 m alt., UTM DK 74, 40.1745°N 20.7297°E, leg. Subai & Szekeres 16.05.1997, S 15847 View Materials /3   ; Tímfi Mountains , limestone grove after 3h 30' walking E of Pápigo village (= approx. 1–2 km NE of refuge hut), on limestone boulders and under stones, 1650–1700 m alt., UTM DK 82, 39.9961°N 20.7709°E, leg. Subai 24.05.1991, S 15819 View Materials /5   ; Tímfi Mountains, near Drakólimni Lake , 2215 m alt., UTM DK 82, 39.98°N 20.77°E, leg. Gittenberger 26.7.1997, NNM/1 View Materials   ; 6.6 km from Kónitsa direction Pádes , on limestone rocks in dry mixed forest, 1000 m alt., UTM DK 83, 40.0547°N 20.7898°E, leg. Subai 16.07.1990, S 15870 View Materials /2   ; Pádes (= approx. 28 km E of Kónitsa), on limestone rocks at W and N border of the village, 1150–1250 m alt., UTM DK 93, 40.0408°N 20.9054°E, leg. Subai 16.07.1990, S 15868 View Materials /2   ; rocks at S border of Agía Paraskeví Village , 995 m alt., UTM DK 94, 40.1438°N 20.8965°E, leg. Fauer & Subai 25.09.1989, S 15859 View Materials /1   ; approx. 1 km E of Agía Paraskeví (= in N part of Smólikas Mountains ), on rocks beside the highway, 1060 m alt., UTM DK 94, 40.1411°N 20.9069°E, leg. Fauer & Subai 25.09.1989, S 15861 View Materials /2   ; 3 km N of Drossopigí, meadow and rocks (no lime) approx. 50 m E of the old arched bridge above the Sarantáporos river , 705 m alt., UTM DK 95, 40.2281°N 20.9133°E, leg. Subai 15.07.1990, S 15873 View Materials /1 + 1 (juv.)   ; 17.5 km NE von Árta, an Felsen neben der Landstrasse, 180 m alt., UTM EJ 14, leg. Gittenberger & Uit de Weerd 14.5.2000, NNM/2 View Materials . —   Evritanía, in spruce forest 6.8 km S of Proussós (= direction Agrínio ), under stone plates, 1300 m alt., UTM EH 58, 38.7164°N 21.6533°E, leg. Subai 17.05.1991, S 15818 View Materials /13   ; bridge at Mégdova river (= Tavropós , 26 km W of Karpeníssi), on limestone rocks, 340 m alt., UTM EJ 51, 38.9415°N 21.6809°E, leg. Fauer & Subai 30.09.1989, S 15854 View Materials /3   ; small valley at mountain slope E of Kalesméno (= 16 km W of Karpeníssi), on limestone rocks, 880 m alt., UTM EJ 60, 38.9212°N 21.7281°E, leg. Fauer & Subai 30.09.1989, S 15856 View Materials /1   ; 1.5 km N of Vráha, approx. 500 m from Moni Vráha , mountain N and S slope, on limestone rocks, approx. 1100 m alt., UTM EJ 62, 39.0776°N 21.7930°E, leg. Subai & Szekeres 12.05.1997, S 15815 View Materials /2 + 1(alk.)   ; Kaliakoúda Mountains, Timfristós Mountain (= " Kaljakuda Berge, Veluchi "), UTM n. det., leg. Krüper, 1890, SMF 101913 View Materials /4, 101914/17, 284854/3. — Etoloakarnanía, UTM n. det., SMF 101936/3   ; Etoloakarnanía, on the narrow land strip leading to Lefkáda Island, on limestone rocks and on ruins of a small castle, UTM DH 79, 38.8434°N 20.7213°E, leg. E. & P. Subai 06.08.1975, S 3816/4   ; Vónitsa, UTM DJ 90, 38.92°N 20.88°E, leg. Conéménos, 1890, SMF 284856 View Materials /2   ; Veloútsa Mountain SE slope above Astakós , on limestone rocks, approx., 185 m alt., UTM EH 06, 38.5370°N 21.0692°E, leg. Subai, 19.05.1991, S 15817 View Materials /19   ; small valley 1 km E of Astakós , on limestone rocks, UTM EH 06, 38.5323°N 21.0998°E, leg. Subai, 19.05.1991, S 15816 View Materials /10   ; Lisimáhia (= Murstiana, Mushtiona), UTM EH 26, 38.54°N 21.33°E, leg. Krüper, 1928, SMF 101916 View Materials /32   ; Messolóngi, UTM EH 34, 38.36°N 21.42°E, SMF 285671 View Materials /2   ; leg. Krüper, 1899, SMF 101915 View Materials /7, above Kefalóvrisso , on limestone rocks, 30 m alt., UTM EH 35, 38.4598°N 21.3670°E, leg. Fauer & Subai 30.09.1989, S 15855 View Materials /6   ; Klisura gorge between Agrínio and Etolikó, on limestone rocks above Ag. Eleusis monastery, 290 m alt., UTM EH 36, 38.5006°N 21.3674°E, leg. Pintér, E. & P. Subai 31.07.1976, S 3815/3   ; Berg Varássova (= Karvassara), UTM EH 54, 38.36°N 21.59°E, leg. Conéménos, 1884, SMF 284849 View Materials /2   ; Gurica (= Goritza, between Mirtéa and Ágios Nikólaos monastery) at Trihonída Lake (= Lake of Vrachori ), UTM EH 57, 38.58°N 21.61°E, leg. Krüper, 1891, SMF 101938 View Materials /5, 101940/17, 207802/1, 284831/4, 284843/6   ; N of Amfilohía , on limestone rocks 100–200 m from the bifurcation direction Sardínia, 60 m alt., UTM EJ 10, 38.8833°N 21.1740°E, leg. Fauer & Subai 29.09.1989, S 15865 View Materials /16   ; 5.9 km from Empessós direction Perdikáki , on limestone rocks in valley of two lime mountains, approx. 600 m alt., UTM EJ 31, 39.0058°N 21.3606°E, leg. Fauer & Subai 29.09.1989, S 15866 View Materials /2   ; 2.5 km E of Halkiópouli , on limestone rocks, 600 m alt., UTM EJ 31, 38.9829°N 21.3700°E, leg. Subai 17.05.1995, S 15839 View Materials /2   ; 9.5 km from Halkiópouli direction Tríklino , on limestone rocks, 950 m alt., UTM EJ 31, 38.9666°N 21.3964°E, leg. Fauer & Subai 29.09.1989, S 15864 View Materials /1   ; 11.5 km E of Halkiópouli , on small limestone rocks, 750 m alt., UTM EJ 31, 38.9702°N 21.4000°E, leg. Subai 17.05.1995, S 15838 View Materials /2.   Fokída, Strómi Village (= 13.5 km SW of Paúliani), on limestone rocks, 850 m alt., UTM FH 08, 38.7089°N 22.2568°E, leg. Subai 12.04.1986, S 15822 View Materials /1. —   Ahaïa , Pátra (= Patras), UTM EH 63, 38.23°N 21.73°E, SMF 101942 View Materials /1.  

Remarks: According to Cernohorsky (1978: 299), the publication of Anton was issued "around the middle of the year, 1838", but therein, the name Helix corcyrensis   remained a nomen nudum. Thus, Helix canalifera ANTON, 1838   is the first correct and valid description of the Lindholmiola species   from Corfu. This rarely used name (last mention by Polinski (1924: 130)) is here considered a nomen oblitum. However, Rossmässler (1838: 40, Taf. 39, Fig. 538) validated the name H. corcyrensis   ( ICZN 23.9 cannot be applied), and it was used under his authority in the subsequent literature. The type specimen of Rossmässler is missing in his collection (note of the late Dr Zilch).

In its shell form, L. corcyrensis   is quite stable, but it varies considerably in size. The populations on the Ionian Islands and the western Epirus are small to medium-sized with a narrower umbilicus and a flat spire (cefalonica sensu auct.). In the central parts of Epirus, shells with a broader umbilicus occur. Similarly sized shells, but with an even larger umbilicus and flatter spire are recorded from the Tímfi and Grammos Mountains , where they live at altitudes between 1200–2000 m.

The eastern part of the distribution area of L. corcyrensis   is inhabited by populations with a strikingly different shell. Here, shells grow larger, with a more elevated spire and a large umbilicus (octogyrata sensu auct.). However, in the same area, shells have been recorded, which do not differ from those in the Epirus, and thus, octogyrata cannot be considered a separate taxon.

Very small specimens can be found on the island of Lefkáda in Greece and in some places on the eastern shore of Lake Ohrid.

Distribution ( Fig. 29 View Fig ): Lindholmiola corcyrensis   inhabits the south-western Balkan Peninsula. There are two isolated localities known, the Pelješac Peninsula and the Island Badija in Croatia, and the Monte Gargano in Italy. South of the Croatian locality, there is an area of almost 200 km length without any Lindholmia species at all. The main distribution area of L. corcyrensis   starts in south-eastern Montenegro and covers the whole of Albania and western Greece. In the south, it reaches the island of Kefallinia and the northern shores of the Bay of Korinthos. The southernmost record from close to Pátra in Ahaïa could not be verified by recent collecting efforts. The eastern boundary of its distribution area follows the eastern rim of the Pindos Mountains.

R

Departamento de Geologia, Universidad de Chile

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Stylommatophora

Family

Helicodontidae

Genus

Lindholmiola

Loc

Lindholmiola corcyrensis ( ROSSMÄSSLER, 1838 )

Subai, Peter & Neubert, Eike 2014
2014
Loc

Helix (Helix) canalifera

ANTON, Verzeichniss der Conchylien 1838
1838
Loc

Helix canalifera

ANTON, Verzeichniss der Conchylien 1838
1838
Loc

Helix canalifera

ANTON, Verzeichniss der Conchylien 1838
1838
Loc

Helix canalifera

ANTON, Verzeichniss der Conchylien 1838
1838
Loc

girva

FRIVALDSZKY 1835
1835
Loc

girva

FRIVALDSZKY 1835
1835
Loc

girva

FRIVALDSZKY 1835
1835
Loc

girva

FRIVALDSZKY 1835
1835
Loc

Helix contorta

LINNAEUS 1758
1758
Loc

Helix contorta

LINNAEUS 1758
1758
Loc

Helix contorta

LINNAEUS 1758
1758
Loc

Helix contorta

LINNAEUS 1758
1758