QUINQUECAPSULARIIDAE Dumitrica, 1995

Suzuki, Noritoshi, Caulet, Jean-Pierre & Dumitrica, Paulian, 2021, A new integrated morpho- and molecular systematic classification of Cenozoic radiolarians (Class Polycystinea) - suprageneric taxonomy and logical nomenclatorial acts, Geodiversitas 43 (15), pp. 405-573 : 464

Family QUINQUECAPSULARIIDAE Dumitrica, 1995

Quinquecapsulariidae Dumitrica, 1995: 21 . — De Wever et al. 2001: 200-201 [in Entactinaria ]. — Afanasieva et al. 2005: S278 [in Order Capsulata ]. — Afanasieva & Amon 2006: 117.

Quinquecapsulariidae O’Dogherty, 1994: 268 [nomen nudum].

TYPE GENUS. — Quinquecapsularia Pessagno, 1971b: 362 [type species by objective designation: Quinquecapsularia spinosa Pessagno, 1971b: 364 ].

INCLUDED GENERA (CENOZOIC ONLY). — Joergensenium BjØrklund, Dumitrica, Dolven & Swanberg, 2008: 460 .

DIAGNOSIS. — The central structure is very small with twin pentagonal frames located parallel to each other. Three to five connecting bars between these two pentagonal frames are present. This central structure comprises two acute corners and one straight beam is arising from each corner. A Cenozoic member of this family has three concentric shells: the innermost shell is the central structure, the second internal shell is a spherical outer medullary shell with a patterned, indented surface, and the outermost is large latticed cortical shell. The shape of the outer medullary shell is closely related by radial beams. The endoplasm completely surrounds the outer medullary shell.

STRATIGRAPHIC OCCURRENCE. — Late Miocene-Living.

REMARKS

The family Quinquecapsulariidae was initially proposed for a Cretaceous spherical polycystine Quinquecapsularia spinosa Pessagno, 1972 . Joergensenium is the only known Cenozoic genus. The oldest report of this family dates back to the Early Jurassic ( Empirea Whalen & Carter in Carter et al. 1998). BjØrklund et al. (2008) insisted on Joergensenium being an endemic Norwegian genus but the Joergensenium -species was already identified everywhere in the Neogene.The Internal skeletal structure for Joergensenium was illustrated ( Ikenoue et al. 2016; pl. 6). Based on molecular data, Joergensenium is infected with Marine Alveolata Groups I and II ( Ikenoue et al. 2016).