Borralinae, Gray & Smith, 2008

Gray, Michael R. & Smith, Helen M., 2008, A New Subfamily of Spiders with Grate-shaped Tapeta from Australia and Papua New Guinea (Araneae: Stiphidiidae: Borralinae), Records of the Australian Museum 60 (1), pp. 13-44 : 20

Key to genera of Borralinae

1 Carapace without obvious patterning (with uniform light grey to fawn hair cover in life), profile arched, usually highest midcaput; eyes relatively small; MA unipartite; epigynum often with a large scape projecting over the epigastric groove (sometimes reduced or absent) (Gray & Smith, 2002, figs 1, 4) ............................................................................ Therlinya

—— Carapace patterned with a pair of longitudinal grey-brown stripes, profile in males typically slopes up to fovea, weakly arched to flattish in female; eyes relatively larger; MA bi- or unipartite; epigynum without scape, fossa divided or undivided (the “striped group” genera) .................................................................................................................. 2

2 Males .............................................................................................................................................. 3

—— Females .......................................................................................................................................... 9

3 RVTA with a long, thick stalk and “crochet-hook” like head; epigynal fossa undivided .................................................................................................................... 4

—— RVTA with stalk short or absent; epigynal fossa divided or undivided ........................................................................................................................................... 5

4 RTA large, deeply bifurcate (Gray & Smith, 2004, fig. 11a) ............................................... Pillara

—— RTA large, not bifurcate (Gray & Smith, 2004, fig. 4a) ..................................................... Borrala

5 Conductor with a ventral process (VCP). MA unipartite ( Figs 16a , 17a ) ............................................................................................................................. Elleguna

—— Conductor without a ventral process, MA bi- or unipartite ........................................................... 6

6 Conductor with a smoothly curved anterior margin, lacking processes ............................................................................................................................................. 7

—— Conductor with anterior margin modified by flange-like processes ............................................... 8

7 Lateral MA process weakly sclerotized and entire (not bifurcate); RDTA long, keel-like (“keel” shorter and notched in A. hayllari ) ( Figs 18a,b – 21a,b ) ................................................................................................................ Asmea

—— Lateral MA process strongly sclerotized and bifurcate spiniform; RDTA not keel-like ( Figs 9a,b – 12a,b ) ............................................................................ Jamberoo

8 TL area indistinct; MA lateral process large, sclerotized and spiniform, with a flap-like basal spine ( Figs 13a,b , 14a,b ) ....................................................... Karriella

—— TL large; MA lateral process not as above ( Figs 6a,b , 7a,b ) ........................................... Couranga

9 Epigynum with a large, median, knob-like process placed anteriorly; fossa absent. (Gray & Smith, 2004, fig. 13c–e) ............................................. Pillara (part)

—— Epigynum not as above; knob or knobs, if present, smaller and posteriorly placed; fossa present ........................................................................................................ 10

10 Fossa not divided (an indistinct median line may be present) ..................................................... 11

—— Fossa divided by a distinct median septum ................................................................................. 15

11 Fossa a small, in-sloped pit sometimes partially hooded by the anterior fossa margin; spermathecae posterior to fossa (overlap- ping in P. karuah ) (Gray & Smith, 2004, figs 10c, 11c, 12c) ..................................... Pillara (part)

—— Fossa larger, not as above; spermathecae posterior to or level with fossa ............................................................................................................................................. 12

12 Fossa elongate, open posteriorly; a pair of rounded knobs on the posterolateral epigynal plate (knobs may be damaged) (Figs 13d–f, 14d) ........................................................................................................................ Karriella

—— Fossa margin entire, knob-like processes absent ......................................................................... 13

13 Fossa large, floor with conjoined socket-like recesses posteriorly; lateral fossa margins flanged inwards over copulatory openings

Figures

Fig. 16. Elleguna major. (a,b), male palp (holotype): a, ventral; b, retrolateral. (c–e), epigynum: c, ventral, fl = flange (KS57656); d, ventral, flanges broken (KS83624); e, dorsal—internal genitalia (KS83624). Scale lines 0.5 mm: upper, a,b; lower, c–e.

Fig. 17. Elleguna minor. (a,b), male palp (KS57652): a, ventral; b, retrolateral. (c–e), epigynum (KS79572): c, ventral; d, lateral; e, dorsal—internal genitalia. Scale lines 0.5 mm: upper, a,b; lower, c–e.

Fig. 18. Asmea akrikensis. (a,b), male palp (holotype): a, ventral; b, retrolateral. Scale line 0.5 mm.

Fig. 21 Asmea mullerensis. (a,b), male palp (holotype):

Fig. 9. Jamberoo johnnoblei. (a,b), male palp: a, ventral (holotype); b, retrolateral (KS46335). (c–g), epigynum: c, ventral, d, lateral (KS84010); e,f, plugged and damaged (KS69656); e, ventral; f, lateral; g, dorsal—internal genitalia (KS69656). Scale lines 0.5 mm: upper lines, a,b; lower line, c–g.

Fig. 12. Jamberoo australis. (a,b), male palp (holotype): a, ventral; b, retrolateral (arrow to spicule on lateral process of MA). (c–e), epigynum: c, ventral, d, lateral (KS77020); e, dorsal—internal genitalia (KS46081). Scale lines 0.5 mm: upper, a,b; lower, c–e.

Fig. 13. Karriella treenensis. (a–c), male palp (holotype): a, ventral; b, retrolateral; c, tibia, dorsal.(d–g), epigynum: d, ventral (KS14775); e, lateral (WAM T78285); f, plugged and damaged epigynum, ventral (KS14862); g, dorsal—internal genitalia (KS14690). Scale line 0.5 mm: upper, a–c; lower, d–g.

Fig. 14. Karriella walpolensis.(a–c), male palp (holotype):a, ventral; b, retrolateral; c, tibia, dorsal.(d,e), epigynum:d, ventral (KS7240); e, dorsal—internal genitalia (KS14642). Scale line 0.5 mm: upper, a–c; lower, d,e.

Fig. 6. Couranga kioloa. (a,b), male palp (holotype): a, ventral; b, retrolateral. (c,d), epigynum: c, ventral (KS74064); d, dorsal—internal genitalia (KS69664). Scale lines 0.5 mm: upper, a,b; lower, c,d.

Fig. 7. Couranga diehappy. (a,b), male palp (holotype): a, ventral; b, retrolateral. (c–e), epigynum: c, ventral (KS75360); d, ventral, with plug (KS58105); e, dorsal—internal genitalia (KS75366). Scale line 0.5 mm.