Systellopini, Von Dalla Torre, 1912

Hutchinson, Paul M. & Allsopp, Peter G., 2022, Liomenochilus ongi (Coleoptera: Scarabaeidae: Melolonthinae: Systellopini), a new genus and new species from northern Western Australia, Zootaxa 5100 (1), pp. 119-128 : 120

Tribe Systellopini

Revised diagnosis ( Figs. 1–6 , 8–9 , 12–24 ). Modified from Allsopp (1990). Male. Form normal, not unusually depressed; surface of body and legs dull coloured, not metallic. Labrum large, extended in front of clypeus and in same general plane, either clearly differentiated by a complete suture dorsally or suture only evident on lateral declivity. Clypeus without a fringe of erect, stout-pointed spines along anterior margin, lateral edges usually deeply emarginate before ocular canthus. Antennal club with 3, 5, 6 or 7 lamellae. Pronotum with defined lateral margins and edges. Mesosternum and metasternum without a trace of a median process. Anterior leg with coxae projecting to a distance not greater than width of coxal cavity; tibiae each with 1 or 2 teeth on outer edge in addition to apical tooth. Metacoxae inflated, usually without defined lateral margins; metafemora inflated; metatibia very short and stout, ratio length:width 0.9–2.2:1, complete transverse, setose ridge on middle of outer side, 2 distal spurs, 1 above and 1 below articulation of tarsus. All claws simple and equal. Abdomen without longitudinal ridge along each side. Parameres long and slender, simple and symmetrical.

The tribe contains Chilodiplus Sharp, 1877 (4 species), Enamillus Sharp, 1877 (8 species), Prochelyna Erichson, 1847 (3 species), Sarothromerus Blackburn, 1907 (3 species), Sphyrocallus Sharp, 1877 (3 species), and Systellopus Sharp, 1877 (1 species) ( Allsopp 1990, 1993; Weir et al. 2019), and Liomenochilus Hutchinson & Allsopp new genus (1 species). A further species, Metascelis flexilis Westwood, 1842 , was included in the Systellopini ( Dalla Torre 1912), but the only known specimen is from the India-Nepal border and is not a systellopine ( Allsopp 1990).

Figures

FIGURES 1–7. Liomenochilus ongi Hutchinson & Allsopp, new species, holotype male. 1, dorsal view; 2, ventral view; 3, lateral view; 4, head, dorsal view; 5, head and pronotum, oblique view; 6, pygidium; 7, label.

FIGURES 8–9. Liomenochilus ongi Hutchinson & Allsopp, new species, holotype male, aedeagus. 8, dorsal view; 9, lateral view.

FIGURES 12–18. 12–17. Representative males of each of six systellopine genera, dorsal view. 12, Chilodiplus weiri Allsopp, 1993 (photograph by CSIRO). 13, Enamillus pilicollis (Sharp, 1877). 14, Prochelyna heterodoxa Burmeister, 1855. 15, Sarothromerus sharpi (Blackburn, 1905). 16, Sphyrocallus brunneus Sharp, 1877. 17, Systellopus obtusus Sharp, 1877. 18, Antennal club of male Sarothromerus sharpi (Blackburn, 1905).

FIGURES 19–24. Representative males of each of six systellopine genera, head and pronotum, oblique view. 19 Chilodiplus weiri Allsopp, 1993 (photograph by CSIRO). 20, Enamillus pilicollis (Sharp, 1877). 21 Prochelyna heterodoxa Burmeister, 1855. 22, Sarothromerus sharpi (Blackburn, 1905). 23, Sphyrocallus brunneus Sharp, 1877. 24, Systellopus obtusus Sharp, 1877.