Titiotus costa, Platnick & Ubick, 2008

Platnick, Norman I. & Ubick, Darrell, 2008, A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae), American Museum Novitates 3608 (1), pp. 1-36 : 33-34

Titiotus costa , new species

Figures 82–86 ; map 3

TYPES: Male holotype and female allotype taken in pitfall traps at Canyon, Contra Costa Co. , California (Aug. 25, 1981; D. Denning), deposited in AMNH.

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Males resemble those of T. humboldt and T. marin but can be distinguished by the distally expanded middle prong of the tibial apophysis (fig. 84), females by the rounded epigynal septum, less protuberant posterolateral epigynal margins, and more massive posterior ducts (figs. 85, 86).

MALE: Total length 11. Coloration as in T. flavescens . Leg spination: femora: I, II r1-2-1; III p0-2-1, r1-1-1; IV r1-2-1; tibiae: I d1-0-1, p1-0-0, v4-4-4, r0-1-1; II d1-0-1, v4-4-4, r0-1-1; III r2-0-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Embolus prolaterally expanded near tip, distally arched; median apophysis with only tiny subdistal projection; most ventral prong of tibial apophysis triangular, middle prong greatly expanded distally incised, dorsal prong distally narrowed (figs. 82–84).

FEMALE: Total length 11. Coloration as in T. flavescens . Leg spination: femora: II p0-2-2, r1-2-1; III r1-1-2; IV p1-2-1; tibiae: I p0-0-0, v4-4-4; II p1-0-0, v4-4-4, r1-0-0; III r1-0-1; metatarsi III p1-2-2. Epigynal septum widest at about half its length, posterolateral epigynal margins only slightly protuberant; posterior ducts massive, triangular (figs. 85, 86).

OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Alameda Co.: Berkeley Hills , May 19, 1976, in house (C. Prince, UCB), 13. Contra Costa Co. : Canyon , June 21, 1979, redwood grove (D. Denning, AMNH), 13 , July 1–10, 1981 (D. Denning, AMNH), 13 , Aug. 25, 1981, pitfalls (D. Denning, AMNH), 13; Moraga , Sept. 14, 1973 (D. Denning, AMNH), 1♀ ; Tilden Regional Park , pitfalls (J. Fraser, CAS) : June 20, 1980, 73, 2♀, July 4, 1980, 33, July 16, 1980, 103, 1♀, Aug. 1, 1980, 53, 3♀, Aug. 15, 1980, 23, 4♀, Aug. 29, 1980, 6♀, Sept. 12, 1980, 4♀, Sept. 26, 1980, 1♀; Univ. of California Richmond Field Station , Richmond , July 11, 1961, ground, on canvas (P. Craig, CDU), 1♀ . Napa Co.: N side, Howell Mountain , 2 mi NNE Angwin, all elev. 1300 ft (H. Leech, CAS) : June 9, 1975, 13, May 26, 1977, 13, June 19, 1977, 23, 1♀, Aug. 16, 1977, 1♀, July 1, 1978, 13, June 27, 1980, 1♀, Aug. 18, 1980, 1♀; Napa City, May 27, 2000, in linen closet of vacation home, riparian (K. Stockwell, UCR), 13; St. Helena , May 22, 2004, in wine cellar (J. Martel, UCR), 13, 1♀ . Sonoma Co.: Boyes Springs , Sept. 19, 1993, in house (J. Dorard, S Meehl, CDU), 1♀ ; Sonoma , Sept. 12, 2002, in garage ( UCR), 1♀ ; Sonoma Mountain , just E Cotati , June 6, 1973 (F. DeOme, CAS), 13 .

DISTRIBUTION: Eastern San Francisco Bay region, generally to the east of T. marin (map 3).

Figures

Figs. 82–86. Titiotus costa, new species. 82. Left male palp, prolateral view. 83. Same, ventral view. 84. Same, retrolateral view. 85. Epigynum, ventral view. 86. Same, dorsal view.