Titiotus heberti, Platnick & Ubick, 2008

Platnick, Norman I. & Ubick, Darrell, 2008, A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae), American Museum Novitates 3608 (1), pp. 1-36 : 20-24

Titiotus heberti , new species

Figures 47–51 ; map 1

TYPES: Male holotype and female allotype from Wilsonia, Kings Canyon National Park , Tulare Co. , California (Sept. 13, 1959; W. Gertsch, V. Roth) , deposited in AMNH.

ETYMOLOGY: The specific name is a patronym in honor of Mr. Blaine Hebert, assiduous collector of these and other Californian spiders.

DIAGNOSIS: Males resemble those of T. tulare in having a large, wide dorsal prong on the tibial apophysis, but differ in having a distally narrower middle prong (fig. 49); females differ from those of T. tulare in having a posteriorly wider epigynal septum (fig. 50).

MALE: Total length 12. Coloration as in T. californicus . Leg spination: femora: I p0-3-1, r1-2-1; II r1-2-1; IV p1-2-1; tibiae: I, II d1-0-1, r1-1-0; III r1-0-1; metatarsi: I p1-0-0, r1-1-0; III p1-2-2. Embolus twisted near base, distally wide; median apophysis with subdistal and retrolateral projections; middle prong of tibial apophysis bifid, with subequal branches, dorsal prong large, wide, flattened (figs. 47– 49).

FEMALE: Total length 17. Coloration as in T. californicus . Leg spination: femora: I p0-3- 1, r1-2-1; II r1-2-1; III r1-1-2; IV p1-2-1; patella III p0-0-0; tibiae I, II r1-1-0; metatarsi III, IV p1-2-2. Epigynal septum wide, with posteromedian bulge occupying most of septal width; posterior ducts oblique, arched (figs. 50, 51).

OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Fresno Co. : Cedar Grove, Kings River Canyon , July 16, 1952, elev. 4633 ft (W. Gertsch, AMNH), 8♀ , July 5, 1956, elev. 4633 ft (W., J. Gertsch, AMNH), 23, 6♀ ; Hume Lake , Dec. 17, 1984 (K. Walker, CAS), 1♀ ; Lily Pond , 2 mi W Wishon Lake, Oct. 31, 1982, in sandy soil (M. Marshall, CAS), 1♀ ; Ranger Station, Kings River, Kings Canyon National Park , June 17, 1969, in house (M. Zardus, CAS), 13; Shaver Lake , July 15, 1952 (W. Gertsch, AMNH), 5♀ , Sept. 12, 1959 (W. Gertsch, V. Roth, AMNH), 123, 3♀ ; vicinity Wishon Resv. , July 2–6, 1969 (D. Marqua, MET), 13, 2♀ , May 28–June 14, 1972 (M. Thompson, MET), 2♀ . Kern Co.: Cedar Creek Camp Ground, 0.25 mi N Highway 155, Greenhorn Mountains , 35 ° 44.44 9 N, 118 ° 35.54 9 W, Aug. 25, 2006, on wall of old mine shaft in steep, rocky canyon side near stream, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀ , Aug. 26, 2006 (matured Sept. 4, 2006), on ceiling at mouth of old mine shaft in steep, rocky canyon side stream nearby conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀ , Aug. 26, 2006 (matured Sept. 8, 2006), on base of tree near stream in steep-sided canyon, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀ , Aug. 26, 2006 (matured Dec. 9, 2006), on side of big rock near stream in steep-sided canyon, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 13. Madera Co. : Cascadel Woods, T 8S,R 23E,S16, July 1–3, 1983 (J. Aotok, CAS), 13; Coarsegold, no date (Oliva, UCR), 1♀ , Apr. 21, 2004 (B. Harrigom, UCR), 1♀ ; Wishon, July 5, 1971 (D. Marqua, MET), 13. Mariposa Co.: Foresta Road , Yosemite National Park , 37 ° 42.242 9 N, 119 ° 43.989 9 W, June 15–18, 2003, pitfalls, elev. 1392 m (A. Leache, C. Conroy, UCB), 63. Tulare Co. : Ash Mountain , Sequoia National Park , Apr. 26, 1951 (E. Schlinger, AMNH), 13; near Ash Mountain , Sequoia National Park , July 9, 1958 (W. Gertsch, V. Roth, AMNH), 13, 5♀ ; 1 mi below Belnap Springs, near Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 23, 2♀ ; Carmoe Crevice , Nov. 3, 1993 (J. Krejca, CAS), 13; NE Case Mountain Peak , 36.41790 ° N, 118.7933 ° E, July 14–18, 2003, dry pitfall, elev. 1691 m ( R. Fisher, AMNH), 13; NE Case Mountain Peak , 36.41870 ° N, 118.79264 ° E, July 14–18, 2003, dry pitfall, elev. 1672 m ( R. Fisher, AMNH), 13; NNE Case Mountain Peak, E of E end of Salt Creek , 36.42059 ° N, 118.79776 ° E, May 2003, dry pitfall, elev. 1749 m ( R. Fisher, AMNH), 13 , 36.42083 ° N, 118.79163 ° W, Sept. 2002, dry pitfall, elev. 1609 m ( R. Fisher, AMNH), 1♀ ; NW Case Mountain Peak, S of Salt Creek , 36.41871 ° N, 118.81930 ° W, May 2003, dry pitfall, elev. 1709 m ( R. Fisher, AMNH), 13; just S Case Mountain Peak , 36.407 ° N, 118.8016 ° E, July 14–18, 2003, dry pitfall, elev. 1998 m ( R. Fisher, AMNH), 13; SE Case Mountain Peak, Salt Creek Ridge , 36.39964 ° N, 118.77975 ° W, Sept. 2002, dry pitfall, elev. 1808 m ( R. Fisher, AMNH), 13; W Case Mountain Peak , 36.41120 ° N, 118.80909 ° W, July 2002, dry pitfall, elev. 1951 m ( R. Fisher, AMNH), 1♀ , 36.41139 ° N, 118.80843 ° W, July 2002, dry pitfall, elev. 1932 m ( R. Fisher, AMNH), 1♀ , July 14–18, 2003, dry pitfall, elev. 1951 m ( R. Fisher, AMNH), 1♀ ; just W Case Mountain Peak , 36.41139 ° N, 118.80843 ° W, July 14–18, 2003, dry pitfall, elev. 1932 m ( R. Fisher, AMNH), 13; Cedar Cave, Redwood Canyon, Kings Canyon National Park , Aug. 16, 1984, under rock outside cave entrance, elev. 1,600 m (D. Ubick, CDU), 1♀ ; Crystal Sequoia Cave, Sequoia National Park , Apr. 16, 1946 (F. Oberhansley, AMNH), 1♀ , July 26, 1991 (D. Ubick, CDU), 1♀ ; cave in bank on road to Crystal Cave, Sequoia National Park , Nov. 6, 1949, in total darkness (A. Lange, AMNH), 1♀ ; Harry’s Bend Cave, July 10, 2004 (J. Krejca, A. Gluesenkamp, C. Richard, CAS), 13; Kaweah Power Station #3, Ash Mountain , 40 mi NE Visalia, July 6, 1983 (D. Burdick, CAS, CDU), 23, May 9, 1993 (D. Burdick, CAS), 13, July 2, 1995 (D. Burdick, CAS), 13; Lindcove , July 13, 1976, dead on doorstep (D. Carroll, UCR), 1♀ ; McIntire Creek, near Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 13, 1♀ ; 5–10 mi on Mineking Road from Three Rivers , July 4, 1956, elev. 2000–4000 ft ( AMNH), 13, 4♀ ; Moorhouse Creek, near Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 13 ; 1 mi W Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 2♀ ; Roads End, Kern River, July 3, 1956 (W. Gertsch, AMNH), 13; Soda Creek , W Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 23, 2♀ ; Wilsonia, Kings Canyon National Park , Sept. 13, 1959 (W. Gertsch, V. Roth, AMNH), 23, 3♀ ; 9 mi N Woodlake, Mar. 22, 1941 (S., D. Mulaik, AMNH), 13. Tuolumne Co.: Tuolumne , May 2004, in house (M. Rauscher, CDU), 13 .

DISTRIBUTION: Known only from the Sierras, generally to the south of T. gertschi (map 1).

Figures

Figs. 47–51. Titiotus heberti, new species. 47. Left male palp, prolateral view. 48. Same, ventral view. 49. Same, retrolateral view. 50. Epigynum, ventral view. 51. Same, dorsal view.