Tuiplatynus sophronitis ( Broun, 1908 ) Larochelle & Larivière, 2021

Larochelle & Larivière, Marie-Claude, 2021, Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2021 (864), pp. 1-96 : 27-28

Tuiplatynus sophronitis ( Broun, 1908) , new combination

Fig. 53 , 96 , 166

Anchomenus sophronitis Broun, 1908: 349 . Lectotype (here designated): male (BMNH) labeled “Type (circular red-bordered label; typed) / 2668. [male symbol] (hand-written) / New Zealand. Broun. Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / West Plains Southland (hand-written) / Anchomenus sophronitis (hand-written) / LECTOTYPE [male symbol] Anchomenus sophronitis Broun, 1908 designated by Larochelle & Larivière, 2021 (red label; typed).” Paralectotype: one female (BMNH) from the same locality as the lectotype, bearing a blue paralectotype label.

Agonum (Anchomenus) sophronitis: Csiki 1931: 865 .

Anchomenus sophronitis: Hudson 1934: 174 .

Ctenognathus sophronitis: Larochelle and Larivière 2007: 81 , 111.

Description. Body length 10.0– 11.3 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen piceous black; antennae and palpi rufotestaceous; femora dark yellow; tibiae and tarsi reddish. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Iridescence absent. Very shiny. Head. Mandibles moderately curved anteriorly. Labrum strongly transverse, deeply emarginate anteriorly. Mentum: medial tooth subtruncate apically; ten or eleven setae. Palpi with terminal segment truncate apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc, deeply wrinkled across apex and base, narrow, moderately cordate, widest before middle; apex slightly emarginate; anterolateral angles poorly developed, obtuse; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral beads complete; lateral depressions narrow anteriorly, widened posteriorly, prolonged forward; posterolateral angles obtuse; laterobasal foveae very deep, moderately wide; posterior bead complete; base subtruncate. Elytra. Moderately convex, not sloping down toward apex. Shoulders moderately rounded. Sides moderately rounded. Scutellar striole impunctate. Striae shallow, finely punctate. Intervals depressed; interval 3 with three very deep, large (foveate) setiferous punctures. Umbilicate series with 16–19 setiferous punctures. Subapical sinuations moderate. Aedeagus. Lateral view ( Fig. 96 ): moderately arcuate; base strongly convex dorsally, with basal lobe moderately wide; middle moderately convex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and moderately long; apex moderately convex dorsally, moderately concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area.

Material examined. 127 specimens ( AMNZ, BMNH, JNNZ, LUNZ, MONZ, NZAC).

Geographic distribution ( Fig. 166 ). South Island: CO, DN, FD, OL, SL.

Ecology. Lowland, montane, subalpine. Epigean, arboreal. Wet forests (beech, broadleaf). Associated with streams and mud flats. Shaded ground; wet soil. Nocturnal; hides during the day under logs and stones, and in moss growing on tree trunks. Gregarious.

Biology. Seasonality: September–March, May, August. Tenerals: March, May. Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales).

Dispersal power. Subapterous (incapable of flight). Moderate runner.

Collecting techniques. Turning logs and stones; examining moss on tree trunks; pitfall trapping.

References. Larochelle and Larivière, 2001: 132 (as “ Anchomenus ” sophronitis ; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus sophronitis ; list), 2016: 37 (as C. sophronitis ; list).

Remark. Broun described Anchomenus sophronitis from two specimens, one of which (a male) could be located (BMNH) and is here designated as lectotype in order to preserve stability of nomenclature in the future.

Figures

Figures 53–56. Dorsal habitus. 53) Tuiplatynus sophronitis, teneral. 54) T. libitus, teneral. 55) Ctenognathus novaezelandiae. 56) C. elevatus. Scale line = 1 mm.

Figures 94–100. Aedeagus, lateral. 94) Kupeplatynus crenatus. 95) K. lucifugus. 96) Tuiplatynus sophronitis. 97) T. libitus. 98) Ctenognathus novaezelandiae. 99) C. elevatus. 100) C. garnerae new species.

Figures 165–166. Species distribution maps. 165) Tuiplatynus libitus. 166) T. sophronitis.