Kupeplatynus lucifugus ( Broun, 1886 ) Larochelle & Larivière, 2021

Larochelle & Larivière, Marie-Claude, 2021, Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2021 (864), pp. 1-96 : 25-26

Kupeplatynus lucifugus ( Broun, 1886) , new combination

Fig. 52 , 95 , 152

Anchomenus lucifugus Broun, 1886: 819 . Holotype: male (BMNH) labeled “[AK] Waitakerei [= Waitakere Railway-station] (hand-written) / [male symbol] (hand-written) / 45. [male symbol] (hand-written) / Type (circular red-bordered label; typed) / New Zeal. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; typed) / Ctenognathus lucifugus . (hand-written).”

Anchomenus munroi Broun, 1893: 984 . Lectotype (here designated): male (BMNH) labeled “Type (circular red-bordered label; typed) / 1771 (hand-written) / [AK] Hunua Clevedon. (hand-written) / New Zeal. Broun. Coll. Brit. Mus. 1922– 482. (white label with red horizontal line; typed) / Ctenognathus munroi . (hand-written). / LECTOTYPE [male symbol] Anchomenus munroi Broun, 1893 designated by Larochelle & Larivière, 2021 (red label; typed).” New synonym

Ctenognathus lucifugus: Broun 1893: 986 .

Ctenognathus munroi: Larochelle and Larivière 2007: 81 , 111.

Description. Body length 15.3–18.4 mm. Head and pronotum black; elytra smoky black; abdomen black; antennal segments 1–3 rufopiceous, segments 4–11 reddish; palpi reddish; legs black piceous. Microsculpture isodiametric on head, very transverse (with microlines) on pronotum, and granulate on elytra. Iridescence present on pronotum. Moderately shiny on head, very shiny on pronotum, dull on elytra. Head. Very wide. Mandibles moderately long, slightly curved anteriorly. Labrum slightly transverse, subtruncate anteriorly. Tempora not inflated. Submentum with four setae. Thorax. Pronotum strongly convex, impunctate, obsoletely wrinkled on disc, wide, suborbicular, widest about middle; apex slightly emarginate; anterolateral angles well developed, broadly rounded; anterior bead complete; sides strongly rounded, not sinuate posteriorly; lateral beads absent; lateral depressions narrow anteriorly, wide posteriorly; posterolateral angles broadly rounded; laterobasal foveae very deep and wide, prolonged forward; posterior bead complete; base truncate. Apex of prosternum glabrous. Legs. Moderately long. Metacoxae with three or four posteroventral setae. Metafemora with four posteroventral setae. Meso- and metatarsomeres 1–4 poorly developed, not carinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 strongly asymmetrical apically. Elytra. Strongly convex, sloping down toward apex, ovate. Basal margin complete, reaching about scutellum. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole punctate. Striae deep, coarsely punctate. Intervals strongly convex. Umbilicate series with 20–25 setiferous punctures. Subapical sinuations very strong. Apices mucronate. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view ( Fig. 95 ): base slightly convex dorsally, with basal lobe very wide; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area narrow and moderately long; internal sac with a few scale-like sclerites; apex moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb close to membranous area.

Material examined. 303 specimens ( AMNZ, BMNH, CMNH, LUNZ, MONZ, NZAC).

Geographic distribution ( Fig. 152 ). North Island: AK, BP, CL, ND, TK, TO, WO.

Ecology. Lowland, montane. Epigean, arboreal. Wet forests (broadleaf, podocarp, tree ferns) and tree plantations (pine). Associated with streams and slopes. Shaded ground; wet soil. Nocturnal; hides during the day in and under rotten logs.

Biology. Seasonality: Throughout the year. Tenerals: December–January, March. Predacious (based on mouthpart morphology). Occasionally infested with mites.

Dispersal power. Subapterous (incapable of flight). Swift runner. Regular climber (on trees).

Collecting techniques. Pitfall trapping; breaking and turning logs; pan trapping (yellow pan traps); window trapping.

References. Larochelle and Larivière 2001: 135 (as Ctenognathus lucifugus , C. munroi ; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 111 (as C. lucifugus , C. munroi ; list), 2016: 37 (as C. lucifugus , C. munroi ; list).

Remarks. Broun described Anchomenus munroi from two specimens, one of which (a male) could be located (BMNH) and is here designated as lectotype. This type designation is made to preserve stability of nomenclature in the future. Examination of this specimen revealed it to be conspecific with Kupeplatynus lucifugus .

Figures

Figures 49–52. Dorsal habitus. 49) Kiwiplatynus taranaki new species. 50) Kupeplatynus sulcitarsis. 51) K. crenatus. 52) K. lucifugus. Scale line = 1 mm.

Figures 94–100. Aedeagus, lateral. 94) Kupeplatynus crenatus. 95) K. lucifugus. 96) Tuiplatynus sophronitis. 97) T. libitus. 98) Ctenognathus novaezelandiae. 99) C. elevatus. 100) C. garnerae new species.

Figures 148–153. Species distribution maps. 148) Ctenognathus xanthomelus. 149) Kiwiplatynus bidens. 150) K. taranaki new species. 151) Kupeplatynus crenatus. 152) K. lucifugus. 153) K. sulcitarsis.