Ctenognathus colensonis ( White, 1846 )

Larochelle & Larivière, Marie-Claude, 2021, Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2021 (864), pp. 1-96 : 53-54

Ctenognathus colensonis ( White, 1846)

Fig. 75 , 117 , 127

Anchomenus colensonis White, 1846: 3 . Lectotype (here designated): female (BMNH) labeled “[female symbol] (handwritten) / [?]5 [first number unreadable] (hand-written) / [NC/MC/SC] Canterbury (typed) / New Zeal. Broun Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / Platynus colensonis (hand-written).” LECTOTYPE [female symbol] Anchomenus colensonis White, 1846 designated by Larochelle & Larivière, 2021 (red label; typed).”

Platynus colensonis: Bates 1874: 239 .

Anchomenus (Platynus) colensonis: Broun 1880: 26 .

Anchomenus colensonis: Sharp 1884: 298 .

Anchomenus integratus Broun, 1908: 348 . Holotype: female (BMNH) labeled “Type (circular red-bordered label; typed) / 2667 (hand-written) / New Zeal. Broun. Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / [MC] Broken River (hand-written) / Anchomenus integratus . (hand-written).” New synonym

Agonum (Platynus) colensoni [sic]: Csiki 1931: 865.

Agonum (Anchomenus) integratum: Csiki 1931: 865 .

Anchomenus colensonis: Hudson 1934: 174 .

Agonum integratus [sic]: Johns 1980: 62.

Ctenognathus colensonis: Larochelle and Larivière 2007: 81 , 111.

Ctenognathus integratus: Larochelle and Larivière 2007: 81 , 111.

Description. Body length 10.8–11.2 mm. Head, pronotum, and elytra piceous black; lateral margins of elytra reddish; abdomen piceous black; antennae and palpi rufotestaceous; femora and tibiae pale yellow; tarsi reddish. Microsculpture isodiametric on head, pronotum, and elytra. Iridescence absent. Very shiny. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, deeply emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate apically; two setae. Submentum with ten setae. Palpi with terminal segment truncate apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc, wide, slightly cordate, widest before middle; apex slightly emarginate; anterolateral angles well developed, broadly rounded; anterior bead complete; sides moderately rounded, slightly or moderately sinuate posteriorly; lateral beads complete; lateral depressions narrow anteriorly, widened posteriorly; two setiferous punctures (anterior and posterior) on each side; posterolateral angles slightly obtuse, almost rectangular; laterobasal foveae shallow, moderately wide, prolonged forward; posterior bead obsolete; base subtruncate. Legs. Moderately long. Metafemora with two posteroventral setae. Meso- and metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Slightly convex, not sloping down toward apex, oblong, widest about middle. Shoulders moderately rounded. Sides moderately rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae shallow, finely punctate. Intervals depressed; interval 3 with three shallow, minute setiferous punctures. Umbilicate series with 15–22 setiferous punctures. Subapical sinuations feeble. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with four long apical ambulatory setae; female with eight or ten long apical ambulatory setae. Aedeagus. Lateral view ( Fig. 117 ): strongly arcuate; base strongly convex dorsally, with basal lobe moderately wide; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area very wide in apical half and moderately long; apex moderately convex dorsally, slightly concave ventrally, with extreme tip moderately wide and long. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area.

Material examined. 363 specimens ( AMNZ, BMNH, CMNH, JNNZ, LUNZ, NZAC).

Geographic distribution ( Fig. 127 ). South Island: BR, MC, MK, NC, NN, SC.

Ecology. Lowland, montane, subalpine. Epigean, arboreal. Dry forests (beech). Shaded ground; dry soil. Nocturnal; hides during the day under logs and fallen branches. Gregarious.

Biology. Seasonality: September–May, July. Tenerals: November–December, February–March, May. Predacious (based on mouthpart morphology).

Dispersal power. Subapterous (incapable of flight). Moderate runner. Regular climber (on tree trunks).

Collecting techniques. Pitfall trapping; turning logs and fallen branches; examining tree trunks at night.

References. Larochelle and Larivière 2001: 130 (as “ Anchomenus ” colensonis , “A.” integratus ; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus colensonis , C. integratus ; list), 2016: 36–37 (as C. colensonis , C. integratus ; list).

Remarks. White described Anchomenus colensonis from two specimens, one of which (a female) could be located (BMNH) and is here designated as lectotype in order to preserve stability of nomenclature in the future. Examination of the type of Anchomenus integratus revealed it to be conspecific with Ctenognathus colensonis .

Figures

Figures 73–76. Dorsal habitus. 73) Ctenognathus intermedius. 74) C. otagoensis. 75) C. colensonis. 76) C. earlyi new species. Scale line = 1 mm.

Figures 115–122. Aedeagus, lateral. 115) Ctenognathus xanthomelus. 116) C. intermedius. 117) C. colensonis. 118) C. otagoensis. 119) C. earlyi new species.120) C. davidsoni new species.121) C. oreobius. 122) C. hoarei new species.

Figures 124–129. Species distribution maps. 124) Ctenognathus adamsi. 125) C. arnaudensis. 126) C. cardiophorus. 127) C. colensonis. 128) C. davidsoni new species. 129) C. earlyi new species.