Ctenognathus sandageri ( Broun, 1882 )

Larochelle & Larivière, Marie-Claude, 2021, Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2021 (864), pp. 1-96 : 49-50

Ctenognathus sandageri ( Broun, 1882)

Fig. 70 , 113 , 143

Anchomenus sandageri Broun, 1882: 218 (redescribed in 1883: 218 and 1886: 748). Holotype: male (BMNH) labeled

“[ male symbol] (hand-written) / Type (circular red-bordered label; typed) / 46. (hand-written) / New Zealand Broun .

Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / [WN] Wellington (typed) / Anchomenus san-

dageri. (hand-written).” Agonum (Platynus) sandageri: Csiki 1931: 853 . Anchomenus sandageri: Hudson 1934: 174 . Ctenognathus sandageri: Larochelle and Larivière 2007: 81 , 111.

Description. Body length 10.2–11.1 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen piceous black, yellowish lateroapically; antennae and palpi rufotestaceous; femora and tibiae pale yellow; tarsi rufotestaceous. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Iridescence absent. Very shiny. Forebody narrow in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate or subrounded apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum moderately convex, impunctate, obsoletely wrinkled on disc, narrow, strongly cordate, widest about middle; apex strongly emarginate; anterolateral angles poorly developed, rounded; anterior bead complete; sides strongly rounded, strongly sinuate posteriorly; lateral beads complete; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles acute, slightly projected laterally; laterobasal foveae moderately deep and wide, prolonged forward; posterior bead incomplete, obsolete medially; base emarginate. Legs. Moderately long. Metafemora with two posteroventral setae. Meso- and metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Very convex, sloping down toward apex, subovate, widest about middle. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae deep, impunctate. Intervals subdepressed; interval 3 with three moderately deep, small setiferous punctures. Umbilicate series with 15–16 setiferous punctures. Subapical sinuations moderate. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view ( Fig. 113 ): moderately arcuate, strongly widened in apical half; base moderately convex dorsally, with basal lobe narrow; middle moderately convex dorsally, strongly concave ventrally, with dorsal membranous area strongly widened in apical half and moderately long; apex subtriangular, moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area.

Material examined. 201 specimens ( AMNZ, BMNH, CMNZ, JNNZ, MONZ, NZAC).

Geographic distribution ( Fig. 143 ). North Island: WA, WN.

Ecology. Lowland, montane. Epigean. Wet forests (broadleaf, beech, podocarp). Associated with streams. Shaded ground; wet soil. Nocturnal; hides during the day under stones and logs. Gregarious.

Biology. Seasonality: October–March. Tenerals: November, March, May. Predacious (based on mouthpart morphology).

Dispersal power. Subapterous (incapable of flight). Moderate runner.

Collecting techniques. Pitfall trapping; turning stones and logs.

References. Larochelle and Larivière 2001: 131 (as “ Anchomenus ” sandageri ; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus sandageri ), 2016: 37 (list).

Figures

Figures 69–72. Dorsal habitus. 69) Ctenognathus kaikoura new species. 70) C. sandageri. 71) C. helmsi. 72) C. xanthomelus. Scale line = 1 mm.

Figures 108–114. Aedeagus, lateral. 108) Ctenognathus takahe new species. 109) C. arnaudensis. 110) C. marieclaudiae new species. 111) C. edwardsii. 112) C. kaikoura new species. 113) C. sandageri. 114) C. helmsi.

Figures 142–147. Species distribution maps. 142) Ctenognathus pictonensis. 143) C. sandageri. 144) C. takahe new species. 145) C. tawanui new species. 146) C. tepaki new species. 147) C. urewera new species.