Diplomaragna terricolor ( Attems, 1899 )

Mikhaljova, Elena V., 2021, The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida), Zootaxa 5060 (1), pp. 1-44 : 16

Diplomaragna terricolor ( Attems, 1899)

Placodes terricolor Attems 1899: 320 , figs.

Placodes terricolor — Attems 1904: 49.

Diplomaragna terricolor — Attems 1907: 123; Lokšina & Golovatch 1979: 383; Golovatch 1979a: 88; Kurcheva, Mikhaljova 1980: 121; Mikhaljova 1981: 87; 1983: 84; 1988: 70; 1993: 29; 1997: 130; 1998: 26, figs, map; 2000: 173, figs; 2002a: 150; 2004: 137, figs, map; 2005: 4; 2009a: 605; 2009c: 4; 2009d: 394; 2016: 20; 2017: 166, figs, map; Mikhaljova, Petukhova 1983: 52; Mikhaljova, Bakurov 1989: 40; Shear 1990: 28, figs; Ganin 1997: 121; 2006: 616; 2009: 153; 2011: 336; Mikhaljova & Marusik 2004: 6; Mikhaljova & Sergeev 2021: 171 , figs.

Material examined. 1 female ( FSCB), Russia, Vladivostok, Botanical garden, 22. V.–1. VI.2010, leg. S.A. Shabalin; 1 male ( FSCB), Russia, Primorsky Krai, near Vladivostok, Peschaniy Peninsula , broadleaved forest, 24.IX.2020, leg. L.A. Prozorova.

Distribution. Russia: Far East (Primorsky Krai, southern part of Khabarovsky Krai).

Remarks. Originally described as Placodes terricolor from the environs of Vladivostok ( Attems 1899), later this species was replaced to the new genus Diplomaragna and the subfamily Diplomaragninae Attems, 1907 (see Attems 1907) and was re-described using topotype material ( Shear 1990).

Morphological variability of this species was recently revealed ( Mikhaljova & Sergeev 2021). There are two distinct extreme forms of the gonopods and of the male coxae 11. The differences in the structure of these extreme forms are so strong that their males would even claim the status of independent species. However, the forms are linked by a complete series of transitions between gonopods of the males from different distant districts of Primorsky Krai and Khabarovsky Krai. It is assumed that this morphological variability is associated with different environmental conditions of the habitats of this species.