Shearia calycina Mikhaljova, 2000

Mikhaljova, Elena V., 2021, The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida), Zootaxa 5060 (1), pp. 1-44 : 35

Shearia calycina Mikhaljova, 2000

Shearia calycina Mikhaljova 2000: 166 , figs.

Shearia calycina —Mikhalojva & Golovatch 2001: 110; Mikhaljova 2004: 163, figs, map; 2017: 200, figs, map; Nefediev & Nefedieva 2007b: 162; Mikhaljova et al. 2008: 53; Nefediev 2019b: 220 .

Distribution. Russia: Siberia (Republic of Altai).

Remarks. This species has been described from one male and 5 females from the South Altai Mts, at the northern extremity of the Yuzhno-Chuisky Mt. Range, the watershed of the Sebystei and Kokozek rivers, in stones at snow line, ca 2,800 m. a.s.l. According to the original description ( Mikhaljova 2000) the posterior gonopod angiocoxite is provided with an anterior short lateral process (ap, see Mikhaljova 2000, page 167, Fig. 38 ). Nefediev (2019b) has revealed a second bifurcated anterior angiocoxal process in gonopods of the males from the Republic of Altai, Kosh-Agach District, floodplain of the Karagem River, ca 1355 m a.s.l. He also named this process ap. However, he did not note that gonopods of the males from floodplain of Karagem River have the same short anterior angiocoxal processes as the ones of the holotype (see Nefediev 2019b, Figs 2 , 4 ). Thus, the species is rather variable. This phenomenon requires special research. Also, additional examination of the affiliation and nature of the second bifurcated process ap is needed (does it belongs to the angiocoxite?).

Figures

FIGURES 37–39. Pacifiosoma acutum sp. nov., male holotype. 37, 39, gonopods, front view, left colpocoxite broken; 38, papillae of anterior colpocoxite part; n, external edge of colpocoxite; ap, anterior angiocoxal process. Scales: 20 μm (Figs 38–39), 100 μm (Fig. 37).

FIGURES 1–2. Alineuma rodionovi sp. nov., male holotype. 1, gonopods, front view; 2, gonopods, caudal view, right femur of posterior gonopod telopodite is twisted; ms, trapezoidal structure; b, hyaline fimbriate branch; ap, anterior angiocoxal process; pp, posterior angiocoxal process; at, anterior gonopod telopodite; sg, subglobule of posterior angiocoxite. Scale in mm.

FIGURES 3–7. Diplomaragna budilovi sp. nov., male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view; c, horn; pp, posterior angicoxal process; ap, anterior angiocoxal process; cx, coxosternum; at, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7).