Coptocephala arcasi Baguena, 1960

Bezdċk, Jan & Regalin, Renato, 2015, Identity of species-group taxa of the Western Palaearctic Clytrini (Coleoptera: Chrysomelidae) described by Maurice Pic and Louis Kocher, Acta Entomologica Musei Nationalis Pragae (suppl.) 55, pp. 1-114 : 12-13

Coptocephala arcasi Baguena, 1960

( Figs 3 , 73–77 )

Coptocephala rubicunda var. dalmatina Pic, 1918c: 17 (unavailable infrasubspeciħc name).

Coptocephala (Coptocephala) arcasi Baguena, 1960: 25 (original description).

Type localities. Coptocephala rubicunda var. dalmatina : ‘Dalmatie’. Coptocephala arcasi : ‘Andalucía’ (suggested as patria falsa by REGALIN & MEDVEDEV 2010a).

Type material examined. Coptocephala arcasi : HOLOTYPE: 1 ♂, ‘ Andalucía [w, h] // Schauffuss / … [partly illegible, reverse side of previous label, w, h] // Col. del Sr. / Perez Arcas [w, p] // Holotipo [r, p] // COPTOCEPH. ♂ / ARCASI m. [h] / L. Báguena det. [w, p]’ ( MNCN).

Original material of infrasubspeciħc entity. Coptocephala rubicunda var. dalmatina : 13, ‘type [w, h] // v. dalmatina / Pic [w, h] // Dalmatie [w, h] // Ragusa [w, h] // TYPE [r, p] // Museum Paris [p] / Coll.Pic [w, h]’ ( MNHN – coll. Pic).

Additional material examined. BOSNIA HERZEGOVINA: Herzegovina, Velez-Planina, 1900, 1 ♂ ( NHMB). CROATIA: Dalmatia, 1 ♂ ( LMCM); Dalmatia, 1 ♂, Apfelbek leg. ( LMCM) ; Beccagnazzo , vii.1937, 1 ♀, Müller leg. ( MSNT – coll. Müller) ; Bella Vista , Ragusa [= Dubrovnik], 4.vii.1909, 1 ♂ ( LMCM) ; Cerovlje , Carso Trieste, 14.viii.1954, 1 ♂ 1♀, Springer leg. ( MSNM) ; Cibaca [= Ć iba č a], Ragusa D., vii.1914, 3 ♂♂♂♂ ♀♀, Mussapp leg. ( MSNT – coll. Müller) ; Gravosa D. [= Gruž ], vii.1914, 1 ♀, Mussapp leg. ( MSNT – coll. Müller) ; Gravosa [= Gruž], 1 ♂ ( RRCM) ; Hvar Isl. , 8.–16.viii. 1990, 1 ♂ 1 ♀, N. Pilon leg. ( RRCM) ; Hvar Isl. , vii. 1979, 1 ♂ ( RRCM) ; Krk Isl., viii.1931, 1 ♂, H. Hopp leg. ( NHMB); Dalmatia , Krk Isl., viii.1931, 1 ♂ 1 ♀ ( NHMB); Krk Isl. , 1 ♂ 1 ♀, Mader leg. ( NHMW) ; Lussin [= Lošinj], Cigale, viii. 1923, 1 ♂, Boehm leg. ( MSNT – coll. Müller) ; Is. Lussin [= Lošinj Isl. ], 1924, 1 ♀, Boehm leg. ( MSNT – coll. Müller) ; Mali Lošinj , 10.viii.1982, 1 ♂ ( DSCC) ; Split, 1 ♂, Obenberger leg. ( LMCM) ; Spalato [= Split], 4 ♂♂♂♂ ♀, Karaman leg. ( LMCM) ; Spalato [= Split], 1914, 1♂ ( LMCM) ; Trau [= Trogir], 1 ♂ ( MSNT – coll. Müller) ; Is. Uglian [= Ugljan Isl.], Lukoran , vii. 1917, 1 ♂, Müller leg. ( MSNT – coll. Müller) ; Unije , 1 ♀ ( MSNM) ; Unije , vii.1922, 3 ♂♂♂♂ ♀♀, Müller leg. ( MSNT – coll. Müller, 1 ♂ MSNV) ; Zara [= Zadar], 1 ♂ 1 ♀ ( MSNM) ; Zara [= Zadar], 15.vii.1937, 1♂, Müller leg. ( MSNT – coll. Müller) ; Zara [= Zadar], 2 ♂♂( MSNV) ; Zelenika , viii.1906, 1 ♀ ( LMCM) ; Zlatni rt, Rovinj, 10.–12.vii.1998, 17 ♂♂♂♂ ♀♀, R. Regalin leg.( RRCM). ITALY: Sistiana, 24.viii.1930, 1 ♂, De Medici leg. ( MSNM) ; Sistiana , 3 ♂♂♂♂ ♀♀, De Medici leg. ( MSNT – coll. Müller) ; Duino , 17.viii.1954, 1 ♀, Springer leg. ( MSNM) ; Trieste , vii. 1902, 1 ♂ ( MSNV) ; Triest, M. Cal , viii.1925, 2 ♂♂♂♂ ♀ ( MSNM) ; Trieste, M. Cal , viii.1925, Sesleri elatum , 1 ♂, Carrara leg. ( MSNT – coll. Müller) ; Trieste, M. Cal , 6 ♂♂ 3 ♀♀ ( MSNT – coll. Müller) ; Trieste, M.te Kal, 1 ♂, Müller leg. ( MSNT – coll. Müller) ; Trieste , Monte Spaccato, Gabroscek, 5.viii.1910, 1♀ ( MSNT – coll. Müller) ; Trieste , Monte Spaccato, 5.viii.1910, 13, Ciana leg.( MSNT – coll. Müller) ; Zaule , Rosandra , ix.1955, 1♀, Sauli leg. ( MSNM). MONTENEGRO: Bocche di Cattaro [= Boka Kotorska ], 3 ♂♂♂♂ ♀ ( NHMB). SLOVENIA: Küstl. Orlek, 20.ix.1922, 2 ♀♀, Springer leg. ( MSNM).

Distribution. Bosnia and Herzegovina, Croatia, Italy, Montenegro, Slovenia (WARCHA Ł OWSKI 1991, REGALIN & MEDVEDEV 2010b, present paper).

Comments. Coptocephala arcasi was described from Spain. Already WARCHA Ł OWSKI (1991) doubted its Spanish origin, discussed the possible misidentiħcation as C. cyanocephala ( Lacordaire, 1848) or C. fossulata Lefèvre, 1872 and treated it as a nomen dubium. Recently, REGALIN & MEDVEDEV (2010a), based on the study of holotype of C. arcasi , stated it was mislabelled and found that the species frequently occurs in former Yugoslavia and northern Italy. Speciħc chromatic pattern on elytra suggests that the holotype of C. arcasi could originate from Montenegro, as the specimens with the same pattern are known from Kotor Bay.

We also had the opportunity to examine the original specimen of Coptocephala rubicunda var. dalmatina Pic and based on the examination of its aedeagus we consider it conspeciħc with C. arcasi . However, the variety dalmatina is an unavailable infrasubspeciħc name as it was described in a publication containing also a description of a subspecies ( PIC 1918c), thus it became infrasubspeciħc according to Article 45.6.4 of the Code ( ICZN 1999). The only name that can be used for the species is C. arcasi . Its distribution was indicated only by country acronyms in the Palaearctic catalogue ( REGALIN & MEDVEDEV 2010b). Here we present speciħc data for each country.

Coptocephala arcasi shows a considerable chromatic variability of elytral pattern. Northern population from Italy (Trieste) has increased black spots on elytra. Towards the south the spots are gradually reduced to completely vanished. Similarly, labrum which is usually black with brown margins tends to be reddish-brown or reddish towards the south. Tibiae and tarsi are also more or less reddish-brown in southern populations.

Figures

Figs 1–3. 1–2 – Coptocephala aeneopicta (Fairmaire, 1864): 1 – aedeagus (dorsal, lateral and ventral views); 2 – spermatheca. 3 – aedeagus of Coptocephala arcasi Báguena, 1960 (dorsal, lateral and ventral views). Scale bars: 0.5 mm for Figs 1 and 3, 0.25 mm for Fig. 2.

Figs 71–77. 71 – Maurice Pic in 1895 inAlgeria (downloaded from http://www.zin.ru); 72 – Maurice Pic in ħnal stage of his life (from VILLIERS 1958). 73–77 – Coptocephala arcasi Baguena, 1960 (original specimen of Coptocephala rubicunda var. dalmatina Pic, 1918, male, 5.4 mm): 73 – habitus; 74 – head; 75 – labels; 76 – aedeagus in ventral view; 77 – aedeagus in lateral view.