Anaulacaspis nigerrima ( BERNHAUER, 1908 )

Assing, Volker, 2016, Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae), Beiträge Zur Entomologie = Contributions to Entomology 66 (2), pp. 201-255 : 229-230

Anaulacaspis nigerrima ( BERNHAUER, 1908)

( Figs 57–61 , Map 3 )

Falagria nigerrima BERNHAUER, 1908: 38 f.

Type material examined: Lectotype ♂, present designation [dissected prior to present study]: “♂ / Buchara / nigerrima Bh. Cotypus / Chicago NHMus, M. Bernhauer Collection / Syntypus Falagria nigerrima Bernhauer, V.I. Gusarov rev. 2015 / Anaulacaspis nigerrima (Bh) , V.I. Gusarov det. 2015 / Lectotypus ♂ Falagria nigerrima Bernhauer , desig. V. Assing 2016 / Anaulacaspis nigerrima (Bernhauer) , det. V. Assing 2016” ( FMNH). Paralectotypes: 1 ♂: “♂ / 124. / Buchara / laevigata Epp., Knauth / nigerrima Bh. / nigerrima Brh. Typus” ( FMNH); 1 ♂: “♂ / Margelan / nigerrima Bh. Cotypus ” ( FMNH); 1 ♂: “♂ / Sibiria or., Reitter / nigerrima Bh. , det. Bernh.” ( FMNH); 2 ♀♀: “Turcmenien. Reitter. Leder. / nigerrima Brh. Cotypus ” ( FMNH). All paralectotypes dissected prior to present study and additionally with the following labels: “Chicago NHMus, M.Bernhauer Collection / Syntypus Falagria nigerrima Bernhauer, V.I. Gusarov rev. 2015 / Anaulacaspis nigerrima (Bh) , V.I. Gusarov det. 2015 / Paralectotypus Falagria nigerrima Bernhauer , desig. V. Assing 2016 / Anaulacaspis nigerrima (Bernhauer) , det. V. Assing 2016”.

Comment: The original description is based on an unspecified number of syntypes from “Buchara, Margelan, Turkmenien, Turkestan (Mts. Ghissar) und Ostsibirien” ( BERNHAUER 1908). Six syntypes, four males and two females are deposited in the Bernhauer collection at the FMNH. One of the males from “Buchara” is designated as the lectotype.

Additional material examined: Iran: Razavi Khorasan: 4 ♂♂, 2 ♀♀, 20 km NW Torbat-e, Heydariyeh, Senobar , 35°26'N, 59°06'E, 1730 m, 28.V.2006, leg. Frisch & Serri ( MNB, cAss) .

Afghanistan: 1 ♀, N Herat, SW Khouchk, Tchaouni Khoch Robat , 29.VI.1959, leg. Lindberg ( NHMW) .

Turkmenistan: 10 exs., locality not specified [“Turcmenien”, “ Turkestan ”], leg. Reitter ( MHNG, MNB, NHMW, cAss) .

Kyrgyzstan: 2 ♂♂, 2 ♀♀, Batken , S Ay-Kol, Teo-Jailoo Valley , Tegir-Malik River , 39°43'N, 69°42'E, 2100 m, 28.VI.2012, leg. Frisch ( MNB, cAss); 2 ♂♂, 1 ♀, Batken, Isfana – Isfara, WSW Ozgorush, Turkestan range, Lyailek River, 39°44'N, 70°00'E, 1520 m, 22.VI.2012, leg. Frisch ( MNB, cAss); 2 ♂♂, 1 ♀, Batken, Isfana – Isfara, WSW Ozgorush, tributary of Lyailek River, 39°44'N, 70°00'E, 1560 m, 23.VI.2012, leg. Frisch ( MNB); 1 ex., Ferghansky Alatau, Toskool-Ata, 1.VIII.2002 ( MNB); 1 ex., Kyrghyzky Alatau, foothills, O.to-Say, 1000 m ( MNB) .

Uzbekistan: 11 exs., Taškent, leg. Leder, Reiter, Faust, Staudinger ( MNB, NHMW, cAss) ; 3 exs., Buchara ( MHNG, MNB) .

Tajikistan: 1 ♂, Varzob district, Varzob, Campus ‘ Chaika’ , 38°46'N, 68°49'E, 1110 m, 7.–8.VII.2012, leg. Barševskis (cAss) ; 1 ex., Varzob district, 48 km N Dushanbe, Khodja Obi Gharm , 1900 m, 3.–4.VII.2012, leg. Barševskis (cAss) ; 37 exs., Hissar-Alai, Adjuk cleft near Varzob , 1200 m, stream bank, 1.–3.VII.1990, leg. Schülke, Wrase ( MNB, cAss, cWun) .

Country not specified: 6 exs., “ Transkaspien ”, leg. Reitter ( NHMW) .

Redescription: External characters as in A. nigra , except as follows:

Whole body black including antennae black, legs blackish brown (old material usually paler, with the pronotum and elytra brown and the legs pale-brown).

Head and pronotum with very fine and rather sparse punctation, without distinct sexual dimorphism. Eyes of somewhat variable length, 0.9–1.1 times as long as pronotum. Posterior margin of tergite VIII broadly concave ( Fig. 57 ).

♂: sternite VIII ( Fig. 58 ) obtusely pointed posteriorly; median lobe of aedeagus ( Figs 59–61 ) 0.27–0.30 mm long, ventral process apically acute (both in ventral and in lateral view), distinctly sinuate, and without membranous median projection at base (lateral view).

Distribution and natural history: Anaulacaspis nigerrima is widespread and apparently not uncommon in Middle Asia (Northeast Iran, Turkmenistan, Kyrgyzstan, Uzbekistan, Tajikistan, Afghanistan) ( Map 3 ). One paralectotype was collected in an unspecified locality in Siberia (“Siberia or.”). The altitudes indicated on the labels range from 1000 to 2100 m. At least some of the specimens were collected on the banks of rivers and streams.

Figures

Map 3: Distributions of Anaulacaspis formosa (black circles), A. elegans (white circles), A. nigrina (black diamonds), A. eminens (white diamonds), and A. nigerrima (stars) in the West Palaearctic and Middle Asia, based on revised records.

Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm.