Paramesius brachypterus Thomson, 1859

Chemyreva, Vasilisa G. & Kolyada, Victor A., 2018, Review of the genus Paramesius Westwood, 1832 (Hymenoptera: Diapriidae, Spilomicrini) from Russia, with description of four new species, Zootaxa 4524 (4), pp. 453-472 : 458

Paramesius brachypterus Thomson, 1859

(Figs 1, 2, 6, 33)

Paramesius brachypterus Thomson, 1859: 371 .

Paramesius angustipennis Kieffer, 1911: 762 . Synonymized by Macek (2001).

Paramesius cameroni Kieffer, 1911: 767 . Synonymized by Macek (2001).

Paramesius spinosus Kieffer, 1910: 753 . Preoccupied by P. spinosus ( Ashmead, 1893) .

Paramesius spiniger Kieffer, 1912: 22 . (Replacement name), syn. nov.

Paramesius spinosus atriventris Kieffer, 1910: 753 , syn. nov.

Taxonomic remark. In the original description of Paramesius angustipennis Kieffer, 1911 , the author used the spelling augustipennis, but in all of his later papers he changed the original name and spelt it as angustipennis ( Kieffer, 1912; 1916) without an obvious reason or explicit statement of intention. However, we follow the article 33.2. of the ICZN (1999) which states that “when an unjustified emendation is in prevailing usage and is attributed to the original author and date it is deemed to be a justified emendation” and use the spelling angustipennis with the original author and date here.

Diagnosis. Body length 2.4–3.6 mm; pronotal cervical area densely pubescent, with numerous short and long setae; neck pubescent (Fig. 2); pleurostoma with sharp blade throughout (Fig. 4); wings reaching to half of T2 length to apex of metasoma or weakly longer; petiole ventrally with dense woolly pubescence (Fig. 6); propodeum with posterior margin in dorsal view weakly arcuate (Fig. 1); notauli full; mesoscutal suprahumeral sulcus absent (Fig. 1); male keel on A4 extending from base to half of segment length.

Variation. Face sculptured with horizontal winkles or smooth. Occipital flange smooth or punctured, narrow to moderately broad. Neck densely to poorly pubescent. Female. Body length 2.5–3.6 mm. Wings 0.45–0.72 times as long as body. A10–A13 or A9– A13 dark brown. A10–A 12 in dorsal view subquadrate to distinctly elongate. Petiole 0.26–0.30 times as long as T2. Anterior scutellar pit with 2–4 carinae. Brachypterous specimens with diminished mesosoma and flattened mesoscutum. Male. Body length 2.4–3.6 mm. A1–A12 yellow to brown. Anterior scutellar pit with 4–6 carinae. Petiole 0.45–0.56 times as long as T2.

Material examined. Syntype: ♀ ( MHNH) labelled “ Paramesius / spinosus K.”, “Vaud” [Switzerland], “Suisse”, “Type”, “Muséum Paris/ Collection/ Ernest André/ 1914” . Syntype: ♀ ( MHNH) labelled “ Paramesius / spinosus/ var./ atriventris”, “ Holotype ”, “Muséum Paris/ 1957/ coll. Kieffer ”, “ Paramesius / spiniger var./ atriventris K./ Type!/ P. L. G. Benoit det. 1956” [England]. UKRAINE. Cherkasy Prov., Dnepr, vicinity of Kanev , 20– 27.VII.1966, VT , 1♂. RUSSIA. Kaliningrad Prov., vicinity of Svetlogorsk, 1–12.VIII.1994 and 30.VII.2000, VK , 2♀, 3♂. Smolensk Prov., NP ”Smolenskoye Poozerye”, 4 km S from Przhevalskoye, 10–15. VI.1991, DK , 1♂.

Belgorod Prov., near Borysovka , 19–20. VI.2012, VC, 2♂ . Moscow Prov. , 12 km SW of Moscow, 23.VII.1975, V. Kostukov, 3♂ ; Moscow City, Bittsevskiy Park , 28.VIII.1993, 1 6–30.IX.1993, 10.X.1993 and 15–30.V.1994, VK, 2♀, 4♂ ; Katuary , 16–18.VIII.1954, B. Rodendorf, 2♂ ; Krasnozavodsk , 25.VII.1981, V. Alekseev, 1♂ ; Sokolniki , 10.VIII.1993, VK, 1♂ ; Pavlovskaya Sloboda , 13.IX.1993, VK, 1♀ ; Moscow City , 1 4–15.IX.1963, K. Arnoldi, 3♀ ; 102 km NE from Moscow, near Yaroslavskoe highway, 26.VII.1981, V. Alekseev, 1♂ ; Podolsk , 18–23.V.2010, K. Tomkovich, 1♀ . Primorsky Terr., 15 km E from Chernigovka , 20.VII.1991, SB, 1♀ ; Novokachalinsk , 10. VI.1972, AP, 1♀ .

Distribution. England (Kieffer 1910 as P. spinosus var. atriventris ); Switzerland (Kieffer 1910, as P. spinosus ); Germany, Austria, Czech Republic, Hungary, Poland (Macek, 2001); Ukraine ( Tymochko 2017), Russia (European part ( Kozlov 1978), *Far East).

Remark. The syntype of Paramesius spinosus var. atriventris is in a bad condition ( Notton, 2004), but it is possible to see that pleurostoma with sharp blade throughout, wings brachypterous, petiole ventrally with woolly pubescence, posterior margin of propodeum weakly arcuate and the antennae similar with Paramesius brachypterus .