Adelomomphinae Minet & Park

Park, Kyu-Tek, Koo, Jun-Mo & Minet, Joël, 2020, Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea), Zootaxa 4845 (2), pp. 151-190 : 181

Subfamily Adelomomphinae Minet & Park , subfam. nov.

LSID: urn:lsid:zoobank.org:act:17F2C930-B395-4DF7-889D-F613E0A03826

Type-genus: Adelomompha Minet & Park , gen. nov. by present designation.

Diagnosis. This group share with the other Momphidae the following traits: (1) the compound eye is notched in the vicinity of the antennal foramen (as in Fig. II-18.4Q of Komai et al. 2011), (2) the head is smooth-scaled, (3) the ocelli are absent, (4) in the hind wing, the anterior cubital vein (CuA) is somewhat arched and has its distal section (proximad of CuA2) fairly close and parallel to the anal margin (as in, e.g., Figs. 2A , 5A and 8A of Luz et al. 2014), (5) the abdominal terga II–VII have large areas of close-set spiniform scales, (6) the apodemes of abdominal sternum II are practically absent in the male, (7) the aedeagus has largely desclerotized walls, (8) the vinculum has narrow arms, (9) when observed in lateral view, the uncus is long, slender, pointed apically, curved anteriorly, and separated from the tegumen by a well indicated furrow, (10) the gnathos is absent. Nevertheless the subfamily Adelomomphinae differs from the other Momphidae (here placed in subfamily Momphinae , revised status) by the absence in male and female genitalia of, respectively, a modified transtilla (typically bearing, in Momphinae , a central, scobinate plate, which is sometimes erroneously regarded as part of a gnathos: Koster & Sinev 2003) and a pair of basically sickle-shaped signa (secondarily lost in two species of Palaeomystella Fletcher, 1940 : see Luz et al. 2014, and Basilio et al. 2015). These last two genitalia traits had been regarded as momphid autapomorphies by Heikkilä et al. (2014: 582).

Description. Since the subfamily Adelomomphinae is monotypic, its description does not differ from that of the type-genus (see hereafter). We just summarize here its main traits. Head smooth-scaled. Compound eye notched dorsally. Antenna longer than forewing in both sexes; scape without a pecten. Labial palpus sickle-shaped. Foreleg well developed, with an epiphysis; hindtibia of male dorsally fringed with long piliform scales; spur formula: 0-2-4. Forewing venation: R 4 stalked with R 5, the latter running to the costa; only two M veins. Hind wing venation complete; Rs to costa; M 1 and M 2 short-stalked. Abdominal terga II to VII with wide spinose areas. Antero-abdominal sternum with venulae and sexually dimorphic apodemes (practically absent in the male). Male genitalia: aedeagus without an anterior coecum and sclerotized only dorsally; arms of vinculum narrow; saccus, transtilla and gnathos absent; valve broad, undivided, provided with a large, rounded central structure; uncus elongate. Female genitalia: corpus bursae without sickle-shaped structures but with a small, plate-like, toothed/spinose signum.

Figures

FIGURE 2. Parkiana janineae (Viette, 1954), comb. nov.: A, holotype; B, labels; C, female genitalia, prep. J. Minet no. 1666; D, ditto, close-up of segment VIII (ventral view); E, ditto, corpus bursae; F, ditto, ductus bursae (anterior region); G, ditto, abdomen. Scale bar for the genitalia: 0.5 mm.

FIGURE 5. Torodora ankasokella (Viette, 1968), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. J. Minet no. 1662; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm.

FIGURE 8. Torodora hiarakella (Viette, 1988), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. P. Viette no. 5755; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm.