Antocha (Antocha) fortidens Alexander, 1933b

Markevičiūtė, Radvilė, Podenas, Sigitas, Saldaitis, Aidas & Bernotienė, Rasa, 2019, A new species of Antocha Osten Sacken, 1860 (Diptera: Limoniidae) from Sichuan China, Zootaxa 4661 (1), pp. 118-132 : 125-127

Antocha (Antocha) fortidens Alexander, 1933b

( Figs 6, 7 , 31–36 )

Antocha (Antocha) fortidens Alexander, 1933b: 537 (description); Podenas, 2016: 52.

Diagnosis. Medium-sized, grayish-brown species with light brownish-white wing, wing veins margined with cloud- ed gray pattern, pterostigma brownish-gray, caudal margin of tergite 9 with two rounded projections, gonocoxite oval with elongate apex, outer gonostylus blackened with subapical tooth on outer margin.

Type material examined. HOLOTYPE, male (slide mounted): China, Tibet Border near Tang-Gu, 14,000 ft., August 3–6, 1930, D.C. Graham ( USNM).

Additional material examined. 1 male (in ethanol): China, W. Sichuan, near Barkam, Zhe Gu Shan pass, alti- tude 4100 m, N31°51.528′, E102°40.312′, 2011.VII.28–29, coll. Floriani & Saldaitis ( NRC) .

Redescription. Adult, male (N= 2). General body colour brownish-gray. Body length 5.6 mm. Wing length 8.7 mm.

Head. Dark gray, covered with grayish pruinosity. Antenna ( Fig. 31 ) 1.5 mm long; scape elongate, nearly cy- lindrical, light yellowish-brown; pedicel elongate light yellowish-brown; flagellum 14-segmented, generally light yellowish-brown; flagellomeres elongate, narrower at distal end; apical segment elongate approximately 1.2 times as long as penultimate. Rostrum and mouthparts light brownish-yellow, palpus light brown.

Thorax. Generally light brown. Pronotum light brown. Main colour of mesonotal prescutum light brown. Three conspicuous brown stripes. Scutum brown. Scutellum light brown. Mediotergite brown. Pleuron generally light brown, only anepimeron brownish yellow at wing base, turning whitish yellow towards posterior spiracle. Wing ( Fig. 32 ) light brownish-white; pterostigma elongate, brownish-gray; wing membrane near veins with clouded gray pattern, including areas near midlength of Rs, along cord and outer end of cell dm and as marginal seams along longitudinal veins. Venation: vein Sc long, reaching slightly before branching point of Rs; radial sector long and straight; cell dm comparatively small, approximately twice as long as wide; distal section of vein M 1+2 slightly more than twice as long as anterior margin of cell dm. Veins A 1 and CuP strongly divergent; tip of A 1 reaches wing margin beyond level of Rs origin; anal angle large, nearly right-angled. Male halter 1.05 mm long, base of stem pale brownish-gray, remaining part of stem and knob pale brownish. Legs: coxae light brownish-yellow, darkening more intense on fore and less distinct on posterior coxae; trochanters light brown; femora and tibiae brown with narrowly darkened distal end, three basal tarsomeres brown all with darkened distal end, remaining tarsomeres dark brown; legs covered with short, semi-erect, brown setae. Length of femora: I: 6.2 mm, II: 6.9 mm, III: 7.4 mm, tibiae: I: 7.2 mm, II: 7.1 mm, III: 8.1 mm; tarsi I: 6.6 mm, II: 6.1 mm, III: 6.4 mm.

Abdomen. Generally brown, pruinose. Tergites brown, gradually darkened posteriorly. Sternites brown, pru- inose, lighter than tergites. Abdominal segments with distinct narrow dark brown ring at posterior margin. Male hypopygium ( Fig. 33 ) light brown, a little lighter than abdomen. Tergite 9 ( Fig. 35 ) transverse, conspicuously bilobed, caudal margin with two rounded projections. Sternite 9 ( Fig. 34 ) rounded. Gonocoxite oval, with elongate apex. Outer gonostylus blackened, with subapical tooth on outer margin. Inner gonostylus arcuated, weakly spatulate distally, set with short curved spinous setae. One pair of parameres simple, narrow, sabre-shaped, tip below tip of aedeagus. Aedeagus ( Fig. 36 ) narrow with lateral flanges.

Elevation range in China. Specimens were collected at 4100–4267 m.

Period of activity in China. Adults are flying from July through August.

Habitat. Small streams surrounded by mountainous mixed forest with swampy, mossy meadows. The habitat is dominated by various species of Alnus , Prunus , Quercus , Rhododendron , Abies , different species of other smaller shrubs and ferns.

Distribution. China (Sichuan, Xizang, Tibet).

Remarks. A redescription of this species is provided to add several features not mentioned in the original de- scription; i.e., characters of head, pedicel, rostrum, palpus, sternite 9 and aedeagus. These features are important in identifying A. (A.) fortidens , which is similar to A. (A.) nebulipennis immaculata and A. (A.) nebulipennis nebulipennis .

Figures

FIGURES 1–12. Distinctive features of genus Antocha in China, Sichuan. 1. wing (A. bella sp. nov., holotype); 2. wing (A. bidens, holotype); 3. male antenna of genus Antocha; 4. male antenna of genus Antocha; 5. male antenna of genus Antocha; 6. male genitalia, dorsal view (A. fortidens); 7. genital structures (A. fortidens, holotype); 8. male genitalia, dorsal view (A. nebulipennis nebulipennis, metatype); 9. outer gonostylus and male tergite 9 (A. emarginata, holotype); 10. outer gonostylus (A. bidens, holotype); 11. wing (A. nigribasis, paratype); 12. male genitalia, dorsal view (A. bella sp. nov., holotype).

FIGURES 31–36. Antocha (Antocha) fortidens male. 31. antenna; 32. wing; 33. genitalia, dorsal view; 34. sternite 9; 35. tergite 9; 36. genitalia structures and aedeagus, dorsal view.