Florissantoraphidia mortua ( Rohwer, 1909 ) Rohwer, 1909

Makarkin, Vladimir N. & Archibald, S. Bruce, 2014, A revision of the late Eocene snakeflies (Raphidioptera) of the Florissant Formation, Colorado, with special reference to the wing venation of the Raphidiomorpha, Zootaxa 3784 (4), pp. 401-444 : 432-434

Florissantoraphidia mortua ( Rohwer, 1909) , comb. nov.

( Figs 19 , 20 )

Raphidia mortua Rohwer, 1909: 533 , 534; Cockerell 1912: 215, 216 [as a probable variety of Raphidia exhumata ]; Cockerell 1914: 715; Martynov 1925a: 245; Carpenter 1936: 143, 146, 148, Fig. 10 ; Rodeck 1938: 290; Oswald 1990: 160; Engel 2003: 124, 125; Engel 2009: 211, 213 (Tabl. 1); Engel 2011: 79 (Tabl. 2), 80.

Raphidia moriua [sic]: Carpenter 1953: Fig. 6 .

“ Raphidia ” mortua : Aspöck et al. 1991: 540, 665, Fig. 3053; Engel 2002: 22.

Diagnosis. Distinguished from F. funerata by distal position of 2ra-rp in forewing, absence of branch of RA distad pterostigma in both wings; by forked branches of MP in hind wing.

Redescription. Meso-, metathorax preserved; abdomen possibly present, region poorly preserved. Details not visible.

Forewing narrow, ca. 11 mm long, 2.9 mm wide. Costal space relatively narrow, strongly dilated at proximal 1/ 3. Length from termination of ScP to pterostigma slightly less than length of pterostigma. Subcostal veinlets simple, widely spaced. One crossvein between ScP, RA; other crossvein closes pterostigma proximally. Pterostigma long, rather pale. RA with two branches: one long oblique within pterostigma, another closing pterostigma distally; portion of RA distad pterostigma rather long, lacks branches. RP with two branches proximad 3ra-rp, probably two distad; RP 1 deeply forked, RP 2 simple. Two crossveins between RA, RP; 2ra-rp somewhat distad termination of ScP; 3ra-rp located within pterostigma, near its ending. Two radial cells, nearly equal in length. One intraradial crossvein, between RP 1, RP 2. Three crossveins between RP, MA; proximal crossvein 1rp-m rather far from origin of RP. MA dichotomously branched; MP weakly zigzagged, pectinately branched; anterior trace, two branches all simple. Two long intramedian crossveins. M, CuA diverge at apparent R ( free stem of M+CuA absent or very short). Anterior trace of CuA, one branch simple. CuP simple. Intracubital crossvein long. Crossvein between CuP, AA1 not detected. AA1 simple; AA2 incompletely preserved; crossvein between (iaa) these rather short.

Hind wing approximately 9–9.5 mm long, 2.7 mm wide (estimated). Costal space, ScP not preserved. In subcostal space, one crossvein detected, closed pterostigma proximally. Pterostigma long. RA with one preserved rather short branch, which closes pterostigma distally; incorporated branch not detected. Three crossvein between RA, RP; 3ra-rp located within pterostigma, in distal portion. RP with two branches proximad 3ra-rp. RP 1 deeply forked. Two intraradial crossvein detected, both between RP 1, RP 2. Three crossveins between RP, M/MA detected; 1r-m long, running parallel to R; connecting RP, M relatively far distad wing base. M forked distad origin of RP. MA deeply forked, each branch forked. MP pectinately branched, anterior trace shallowly forked with one deeply forked, one simple branch (right wing); simple with two deeply forked and one simple branch (left wing). Two intramedian crossveins. CuA pectinate, with three simple branches (right wing), probably with two branches (left wing). Long intracubital crossvein (icu) partially preserved. CuP, anal veins fragmentarily preserved; their pattern implies that CuP simple.

Type material. Holotype: No. 4512 (collected by G.N. Rohwer in 1908), deposited in UCM. An incomplete and crumpled specimen. Verbatim label data [typed and hand written]: “F / UCM Paleontology TYPE Collection / Neuroptera : Raphidiidae / UCM #: 4512 Acc#: / Locality #: 83014 Description: lateral / ID: Raphidia mortua Rohwer / Age: Late Eocene Collector: GN Rohwer 1908 / Fm: Florissant Citation: Toohey #52984”. The specimen has black ink writings on the rock that say “ Raphidia / motrua / Roh. / TYPE ” and “N 14”; red ink writing that says “ TYPE ”; and red / black ink writing on a circular paper glued to the rock that says “ TYPE / 4512”.

Type locality. Florissant: Station 14 ( UCM locality No. 83014).

Figures

FIGURE 6. Megaraphidia exhumata (Cockerell, 1909). Holotype UCM 4513. A, the specimen as preserved. B, forewing venation. Scale bar = 4 mm (both to scale).

FIGURE 10. Archiraphidia tumulata (Scudder, 1890). Holotype MCZ 244. A, counterpart. B, part (converted to right). Scale bar = 4 mm (both to scale).

FIGURE 19. Florissantoraphidia mortua (Rohwer, 1909). Holotype UCM 4512 as preserved (covered by Canada balsam). Scale bar = 4 mm.

FIGURE 20. Florissantoraphidia mortua (Rohwer, 1909). Wing venation of the holotype UCM 4512. A, right forewing. B, right hind wing. C, left forewing. D, left hind wing. Scale bar = 4 mm (all to scale).