Tetraneuromyia brevipalpis, Jaschhof & Jaschhof, 2019

Jaschhof, Mathias & Jaschhof, Catrin, 2019, New Dicerurini from Europe, mostly Sweden (Diptera: Cecidomyiidae, Porricondylinae), Zootaxa 4559 (2), pp. 245-264 : 258-259

Tetraneuromyia brevipalpis sp. nov.

Figs 32–34

Diagnosis. This new Tetraneuromyia is based on characters of males; females and larvae are not known. The palpus of T. brevipalpis is shorter than in any of the congeneric species; although consisting of 4 segments, as in other Tetraneuromyia , it is usually only half, occasionally up to three fifths as long as the head height. Normally in Tetraneuromyia , palpus length approximates head height. Apart from that, there are certain genitalic structures rendering T. brevipalpis unmistakable, with particulars as follows. An obvious peculiarity is the ventral gonocoxal emargination, which is larger than in any other Tetraneuromyia and strongly sclerotized all round ( Fig. 32 , ↓ 1). Another character specific to T. brevipalpis is that the anterior processes of the gonocoxal apodemes are extremely short ( Fig. 32 , ↓ 2); normally in Tetraneuromyia they are much longer, protruding far beyond the anterior gonocoxal edge (see, for example, Fig. 38 ). Of the two pairs of tegminal processes, the posterior processes are directed ventrally, by that resembling cowhorns ( Fig. 32 , ↓ 3), while the dagger-shaped lateral processes are directed laterally ( Fig. 33 , ↓ 4).

Other male characters. Body size 1.7–1.9 mm. Head. Eye bridge 0–1 ommatidium long dorsally. Scape and pedicel concolorous with flagellum. Neck of fourth flagellomere 2.0–2.1 times as long as node ( Fig. 34 ). Circumfila with 1–2 short posterior extensions. Thorax. Anepisternum asetose. Wing 2.8 times longer than wide. M 1+2 usually completely faded. Genitalia ( Fig. 32 ). Posterior edge of ninth tergite slightly concave. Gonocoxae with narrow asetose portion ventrally. Gonostylus robust, slightly bent inwards, broadly rounded apically, with inconspicuous tuft of large microtrichia but no tooth apically. Ejaculatory apodeme thick, with sclerotized broadening apically.

Etymology. The species epithet, brevipalpis , is a Latin adjective referring to the short palpus.

Type material. Holotype. Male , Sweden, Öland , Mörbylånga , Skogsby, Station Linné, swampy meadow bordered by willow bushes, 6–27 September 2016, Malaise trap, M. & C. Jaschhof & E. Gustavsson (specimen no. CEC 1492 in NHRS). Paratypes. 1 male, Mörbylånga, Frösslunda alvar, pasture, 1 September–5 October 2003, MT, SMTP (trap 20, collecting event 236) (spm. no. CEC 1493 in NHRS) ; 1 male, Mörbylånga, Gamla Skogsby, “diversity meadow”, bushy meadow at forest edge, 15 August–6 September 2006, MT, SMTP (trap 22, collecting event 1829) (spm. no. CEC 1494 in NHRS) ; 3 males, same locality, 19 September–10 November 2015, MT, MCJ (spms nos CEC1495 – CEC 1497 in SDEI) ; 2 males, Mörbylånga, Västerstad elm forest NR, old-growth broadleaf forest, MT, SMTP (trap 3001, collecting event 3058) (spms nos CEC1498 – CEC 1499 in NHRS) ; 2 males, same data as the holotype, (spms nos CEC1503 – CEC 1504 in SDEI) ; 1 male, same data but 5 August–5 September 2016 (spm. no. CEC 1502 in SDEI) ; 1 male, same data but 8 October–9 November 2016 (spm. no. CEC 1505 in SDEI) ; 1 male, Mörbylånga, Gamla Skogsby, mixed broadleaf forest, 5 August–5 September 2016, MT, MCJ & E. Gustavsson (spm. no. CEC 1506 in SDEI) ; 3 males, same data but 6 September–7 October 2016 (spms nos CEC1507 – CEC 1509 in NHRS) ; 1 male, same data but 30 August–19 September 2017, MCJ (spm. no. CEC 1511 in NHRS) ; 1 male, same data but 20 September–19 October 2017 (spm. no. CEC 1510 in NHRS) ; 1 male, Öland, Borgholm, Rönnerum-Abbantorp NR, swampy mixed broadleaf forest, 22 August–30 September 2015, MT, MCJ (spm. no. CEC 1500 in SDEI) ; 1 male, Borgholm, Skepparsäng NR, dry pine forest, 24 August–1 October 2015, MT, MCJ (spm. no. CEC 1501 in SDEI) .

Other material examined. Slovak Republic: 1 male, Muránská planina NP, Hrdzavá dolina, 28 June–26 July 2010, MT , J. Roháček and J. Ševčík (spm. no. A 7788 in NMPC) .

Figures

FIGURES 32–40. Tetraneuromyia brevipalpis, male, holotype. 32: Genitalia, ventral. 33: Tegmen, ventral. 34: Fourth flagellomere, lateral. Tetraneuromyia discrepans, male, holotype. 35: Fourth flagellomere, lateral. 39: Genitalia, ventral. Tetraneuromyia lapponica, males. 36: Fourth flagellomere, lateral, holotype. 37: Gonostylus, dorsal, paratype. 38: Genitalia, ventral, holotype. 40: Tegmen, ventral, holotype. Scales 0.05 mm. Arrows refer to characters described in the diagnosis.