Celebargiolestes toli, Vincent J. Kalkman, 2016

Vincent J. Kalkman, 2016, Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae), Odonatologica 45 (3), pp. 235-269 : 251-256

publication ID	https://doi.org/ 10.5281/zenodo.163451
persistent identifier	https://treatment.plazi.org/id/03938796-FFCE-FFFB-268D-8C52FC3DFE4B
treatment provided by	Plazi
scientific name	Celebargiolestes toli
status	sp. nov.

Treatment

Celebargiolestes toli sp. nov.

( Figs 11–12 View Figures 7 – 14 , 19–20 View Figures 19 – 22 , 28–31 View Figures 28 – 33 , 40–42 View Figures 40 – 45 )

Material studied

Holotype ♂. Indonesia, Sulawesi Tenggara, south of Sanggona, Gunung Watuwila , Sungai Lalonduwasi near Centipede camp , small and shaded stream in very steep valley, clear running water over boulders, small pools, 1 050 m (ca 03°49’S, 121°40’E), sample 89JvT035 , 02− 04-xi-1989, J. van Tol leg. ( RMNH). GoogleMaps

Further material

Sulawesi Tenggara, mainland

4♂, S of Sanggona, Mokowu river near Mokowu camp, foot of Watuwila Mts , rivulet with ponded areas, clear water, bottom with boulders, some sand, largely shad- ed, 150 m a.s.l., 29−31-x-1989, J. van Tol leg.; 18♂ 3♀, S of Sanggona, Gunung Watuwila, Sg. Lalonduwasi near Centipede camp, small, shaded stream in very steep valley, clear running water over boulders, small pools, 1 0 50 m a.s.l., 02−04- -xi-1989, J. van Tol leg.; 2♂ 1♀, NE of Kolaka, near Gunung Watowila, Centipede Camp, undisturbed hilly rainforest/narrow stream, 1 100 m a.s.l., 02−05-v-1989, J.P. Duffels leg.

Sulawesi Tengah, Banggai Archipelago

4♂, Banggai Archipelago, Peleng [Peleng Island], 05-viii-1941, J.J. van der Starre leg.

Sulawesi Selatan

1♂, Mountain Tangke Salokko, 1 500 m a.s.l., 15-xi-1931−01-i-1932, G. Heinrich leg.; 1♂, ca 55km over the road N of Wotu, brook with rather quietly flowing water, nearly fully shaded in the forest, but also collected on open spots near the road, ponded sites, bottom with enormous boulders, with pebbles and coarse sand, 920m, 29-iv-1991, J. van Tol leg.; 4♂, ca 40 km over the road N of Wotu, brooklet on steep slope near crossing with road, some ponded sites, full shade by hardly disturbed primary rain forest, width 50−100 cm, clear water, 500 m a.s.l., 29-iv-1991, J. van Tol leg.; 7♂ 2♀, 30 km N of Wotu, Sg Anoa, river through primary forest, waterfall near bridge Wotu-Tentena, upstream ponded sites, small tributaries, boulders, sand, 650 m a.s.l., 23-x-1993, J. van Tol leg.

Etymology

The species is dedicated to Jan van Tol (masculine singular genitive noun).

Male (holotype)

Head − Labium completely yellow. Rear of head of normal shape. Front of head, including labrum, mandibles, clypeus and sockets of antennae yelloworange ( Fig. 11 View Figures 7 – 14 ). Basal two segments of antennae yellow-orange, remainder brown. Top of head black.

Thorax − Sides of prothorax dull orange mottled with black; pronotum dull orange areas with median parts of anterior, middle and posterior lobe blackish. Posterior lobe of pronotum broad, seen from lateral gently sloping and with the outer border steeply raised, lateral corners rounded. Pattern of synthorax as in Figure 19 View Figures 19 – 22 ; darker areas black, pale areas yellowbrown.

Legs − Coxae and trochanters dull orange-yellow. Femora yellow with two broad but faint black rings, knees black, tibiae and tarsi yellow-brown. Spines pale orange-brown; femora with respectively 9–10, 9–10, 14–16 spines laterally; tibiae of fore legs with 11–12 smaller and 6 larger spines laterally; tibiae of mid and hind legs with 12–13 and 13 spines respectively laterally.

Wings − Venation black. Membrane hyaline with a smoky outer border. Fw and Hw with 2 Ax; Fw with 34 Px and Hw with 32 Px. Ac at about ¼ the distance between Ax1 and Ax2. Brace vein absent. Pterostigma dark brown. Up to three rows of cells between anal vein and border of Fw and up to four in Hw. Fields between Ir2 and R3, between R3 and IR3, and between IR3 and R4 at distal end containing 3 or more rows of cells.

Abdomen − Pattern on abdomen as in Figure 19 View Figures 19 – 22 ; dark pattern elements dark brown to black, pale pattern elements yellow-brown. Anterior half of S1 yellow, posterior half dark brown, S2 with dorsum black and sides pale brown, S3–7 dorsally largely dark, ventral and the lower lateral areas pale brown, anteriorly on each segment the pale brown reaches to the dorsum forming a ring covering approximately ¹ /₅ of the segment. S9–10 black, the latter strongly depressed in the middle, hind border of S10 without modifications. Appendages as shown in Figures 40–42 View Figures 40 – 45 ; brown with only base of lower appendages yellow. Lower appendages long, about half the length of upper appendages. Genital ligula as in Figures 28–31 View Figures 28 – 33 , with shallow and apically broad cleft. Lateral lobes of ligula large with rounded tip but not constricted in the middle.

Measurements [mm] − Total length 49, abdomen 39, Fw 36; Fw pterostigma 1.7 (costal length), 2.3 (longest length); Hw pterostigma 1.9 (costal length), 2.5 (longest length).

Variability

Head – Specimens from the vicinity of Wotu and from Peleng Island have the anterior third to half of labium black while specimens from the vicinity of Gn Watuwila (type area) have a completely yellow labium. Thorax – Fw and Hw with 2 Ax; Fw with 34–40 Px and Hw with 28–36 Px. In specimens from Peleng Island the yellow on the head is more extensive ( Figs 12 View Figures 7 – 14 , 20 View Figures 19 – 22 ).

Abdomen – Seen from ventral the lower appendages show differences in shape between the population from the Gn. Watuwila ( Fig. 44 View Figures 40 – 45 ) and those from the surroundings of Peleng Island ( Fig. 45 View Figures 40 – 45 ).

Measurements [mm] – Total length 44–45, abdomen 35, Fw 32–33, Hw 32– 33 (specimens from Peleng Island). Total length 46–52, abdomen 36–42, Fw 33–36, Hw 33–36 (specimens from the surroundings of Gn. Watuwila ). Total length 47–49, abdomen 37–39, Fw 32–34, Hw 32–34 (specimens from the surroundings of Wotu).

Female

Head − Pattern as in male. Thorax − Pattern on prothorax, synthorax and legs as in male. Fw and Hw with 2 Ax; Fw with 27–32 Px and Hw with 26– 27Px. Pt white with a slight yellow tinge, outer border in older specimens mottled greyish-black.

Abdomen − Stockier and shorter than in male. Pattern largely as in male; dark areas dark brown to black, pale parts are yellow-brown. S3–9 dorsally largely dark, ventral and lower lateral areas yellow, the yellow on the anterior of each segment reaches to the dorsum forming a yellow ring covering approximately ¹/₅ to ¹ /₁0 of the segment. S10 dorsally black but laterally largely pale brown, cerci brown, base of valvifer pale brown becoming brown to black towards the tip, shaped as in C. cinctus .

Measurements [mm] − Total length 41, abdomen 32, Fw 33; Hw 33 (one female from the surroundings of Wotu). length 40–41, abdomen 31–32, Fw 31–34; Hw 31–34 (surroundings of Gn. Watuwila ).

Differential diagnosis

Males are distinguished from other species of Celebargiolestes by the combination of the following characters: (1) labrum yellow; (2) postclypeus and frons yellow-brown; (3) genital ligula with a shallow apical cleft; (4) lower appendages long, about half the length of the upper appendages and, when seen from lateral, with the tip reaching the level of the upper appendages.