Bolbelasmus, Boucomont, 1911

Sommer, David, Hillert, Oliver, Hrůzová, Lucie & Král, David, 2021, Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus, Zootaxa 4920 (3), pp. 380-394 : 392-393

publication ID

https://doi.org/ 10.11646/zootaxa.4920.3.4

publication LSID

lsid:zoobank.org:pub:845CF8B2-07CD-45DC-8128-2EA56C825356

DOI

https://doi.org/10.5281/zenodo.4531757

persistent identifier

https://treatment.plazi.org/id/039387DB-FF97-AD2F-D8DA-8351FA3AFDC3

treatment provided by

Plazi

scientific name

Bolbelasmus
status

 

Key to the subgenus Bolbelasmus species from the western Palaearctic Region

(modified from Hillert et al. 2016)

1. Posterior margin of pronotum incompletely bordered, interrupted laterally of scutellum. …........................... 2

- Posterior margin of pronotum completely bordered. …........................................................ 6

2. Lateral margin of pronotum nearly semicircular between anterior angles and midline in dorsal view, anterior angles almost rounded; frontal horn and medial hornlike apophysis of pronotum distinct; scutellum triangular, approximately as long as wide; body length 11.0– 13.5 mm; southern parts of central Europe, France (Alsace), Italy, Balkan Peninsula, Poland, Russia and Ukraine; Asian part of Turkey...................................... … B. (B.) unicornis (Schrank von Paula, 1789)

- Lateral margin of pronotum broadly rounded, regularly bent from anterior angle to middle, anterior corners of pronotum distinctly acuminate, lateral margin straight in front of anterior angles; frontal horn and medial hornlike apophysis of pronotum always indistinct; scutellum triangular, longer than wide; body length 9.5–12.0 mm; Cyprus, Greece (Rhodes), Iran, Iraq, Syria and Turkey. …....................................................................................... 3

3. Eye cantus weakly acuminate and distinctly exceeding eye laterally; scutellum rounded apically; Asian part of Turkey. ….............................................................................. B. (B.) tauricus Petrovitz, 1973

- Eye cantus regularly rounded and not exceeding eye laterally; scutellum acuminate apically. …....................... 4

4. Frontal horn weakly spaced from fronto-clypeal carina (viewed from above), pronotal punctation double and dense, small punctures sparser; frontal horn directed upwards to weakly backwards ( Figs. 3A, 3 View FIGURE 3 C–D, 4A–B, 4D).................… 5

- Frontal horn close to fronto-clypeal carina (viewed from above); pronotal punctation double and dense, small punctures denser; frontal horn directed obliquely forewards ( Figs. 3B View FIGURE 3 , 4C View FIGURE 4 ); aedeagus as in Fig. 5B View FIGURE 5 ; Cyprus..... B. (B.) makrisi Miessen, 2011

5. Border of posterior margin of pronotum weakly interrupted laterobasally, but with only several single punctures; frontal horn directed weakly backwards ( Figs. 3 View FIGURE 3 C–D, 4D); aedeagus as in Fig. 5C View FIGURE 5 ; Greece (Rhodes), Iraq, Syria and Asian part of Turkey............................................................................… B. (B.) nireus ( Reitter, 1895)

- Border of posterior margin of pronotum distinctly interrupted laterobasally, single punctures absent; frontal horn directed upwardly ( Figs. 3A View FIGURE 3 , 4 View FIGURE 4 A–B); aedeagus as in Fig. 5A View FIGURE 5 ; Iran..................................................................................................... …. B. (B.) zagrosensis Sommer, Hillert, Hrůzová & Král , new species

6. Scutellum impunctate or finely, shallowly punctate; frontal horn simple or furcate apically. …........................ 7

- Scutellum densely punctate; apex of frontal horn simple..................................................... 11

7. Frontal horn simple; genae evenly rounded; oblique keel above eye longer than length of eye; head strongly constricted behind eye. ….............................................................................................. 8

- Frontal horn furcate apically; genae more or less angulate anterolaterally; oblique keel above eye reaching to anterior part of eye; head not strongly constricted behind eye. …............................................................ 9

8. Body surface blackish; pronotal punctation dense and distinct, separated by distance of 1–2 punctures combined; Algeria, Italy (Sicily) and Tunisia. …................................................ B. (B.) vaulogeri (Abeille de Perrin, 1898)

- Body surface brownish; pronotal punctation sparse but distinct, separated by distance of 3–4 punctures combined; Greece (Crete, Kasos). …....................................................... B. (B.) keithi Miessen & Trichas, 2011

9. Frontal horn short (not longer than wide), apex of horn approximately as wide as base of horn; pronotal punctation denser with exception of basal part, puncture separated by diameter of puncture; pronotal base with fine and regularly spaced punctures, puctures in shape of pinned points; elytral striae well impressed, completely developed; elytral intervals moderately convex; Spain. …............................................................... B. (B.) brancoi Hillert & Král, 2016

- Frontal horn long (longer than wide), tapering apically; pronotal punctation sparser, puncture separated approximately by three their diameters; pronotal base with fine, poorly visible and well separated punctures, punctures in shape of pinned points; elytral striae not impressed, punctures in row visible; elytral intervals flat; northern Africa. …............................. 10

10. Frontal horn parallel to anterior part of eyes; horn sharply keeled at basal half; middle hornlike apophysis of pronotum directed upwards, subapical protrusion situated basally; lateral hornlike apophyses of pronotum directed forwards; Algeria and Morocco. ….................................................................. B. (B.) bocchus (Erichson, 1841)

- Base of frontal horn parallel to level of posterior part of eyes; horn broadly keeled apically; middle hornlike apophysis of pronotum directed obliquely, subapical protrusion situated at apex; lateral hornlike apophyses of pronotum directed forwardly; Egypt, Libya and Tunisia.................................. … B. (B.) nikolajevi Hillert, Arnone, Král & Massa, 2016

11. Apex of frontal horn tapering apically, acute, weakly flattened in anterior part of horn; Spain and Gibraltar................................................................................... … B. (B.) howdeni Hillert & Král, 2016

- Apex of frontal horn roughly as wide as base, apex broadly acute, strongly flattened in anterior part of horn; France (including Corsica), Portugal and Spain................................................. … B. (B.) gallicus (Mulsant, 1842)

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