Nyctiophylax (Paranyctiophylax) lancelot, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435, Zootaxa 2435 (1), pp. 1-63 : 48-50

Nyctiophylax (Paranyctiophylax) lancelot , new species

Figures 87–89

Diagnosis: This species has slender, elongate, unbranched gonopods and no sclerotized dorsal paraproctal processes; sternite IX is strongly produced into a median lobe. Similar genitalic structures are found in N. (P.) icelus (Neboiss) from Sulawesi; N. (P.) nepenthes , new species, from Sabah; N. (P.) padangensis Malicky from Sumatra; N. (P.) pongdiatensis Malicky & Chantaramongkol from Thailand; N. simaritensis Malicky & Chantaramongkol from Sumatra; and N. (P.) temburongensis Malicky from Brunei. The new species is most similar to the latter species, from which it is separated by the presence of oval cerci; gonopods arching uniformly in ventral view, not irregularly bending more abruptly at mid-length; and a lance-shaped spine present on the tip of the ventromesal lobe of each gonopod.

Description: Male. Body pale brown, lighter on less-sclerotized parts. Maxillary palp formula (I,II)-IV- III-V. Forewings light brown with weakly pronounced hyaline windows (1) along crossveins r -m, (2) along crossvein m -cu and (3) on arculus (junction of A1+2+3 with wing margin). Forewing length 4.2 mm. Forewing A1, A2, and A3 looping.

Male genitalia. Sternite IX robust, obliquely elongated-ovoid in lateral view ( Fig. 87 ), with welldeveloped apicoventral mesal lobe; tergum IX membranous except for small sclerotized rudiment anterior of fulcrum on each side where sternite IX, paraproctal complex, and cercus meet. Segment X membranous, with single, digitate, hyaline process with terminal, small seta. Cerci large, setose, drop-shaped in lateral view; almost as long as ventral paraproctal processes. Paraproctal complex well-developed, highly sclerotized; each composed of narrow, paraproctal base andlarge ventral paraproctal process horizontal, slightly curving ventrad at apex; dorsal paraproctal process absent. Gonopods elongate, slender, slightly S-shaped in lateral view ( Fig. 87 ) evenly curving mesad in ventral view ( Fig. 88 ); each with small ventrobasal “elbow” and with mesal lobe bearing strong, straight spine ( Figs. 87, 88 ). Phallic apparatus ( Fig. 89 ) located dorsally in genitalia, guided by ventral processes of paraproct complexes and membranous segment X. Phallotheca thick, horizontal, convex dorsally to dorsal constriction, then more slender and slightly downcurved; apex of membranous, retracted membranes covered with granules.

Holotype male: MALAYSIA: Sabah: Tawau: Maliau Basin, Nepenthes Camp , crossing stream, 4 ° 43 ’ 58.9 ’’ N, 116 ° 52 ’ 40.7 ’’ E, 994 m, 9.xii.–14.xii.2007, Mega Malaise Trap (6 m long) [B. Viklund & N. Jönsson] ( NHMS).

Distribution: Malaysia (Sabah).

Etymology: Lancelot, named after the presence of a straight lance-like, long spine on the ventromesal lobe of each gonopod. In the Arthurian and Holy Grail legends, Sir Lancelot is the greatest of King Arthur's knights of the Round Table.

Figures

FIGURES 87–89. Nyctiophylax (Paranyctiophylax) lancelot, new species, holotype. 87—genitalia, left lateral; 88— right gonopod, ventral; 89—phallic apparatus, left lateral.