Trigonoderopsis enduwaensis, Mitroiu, 2016

Mitroiu, Mircea-Dan, 2016, Pireninae (Hymenoptera: Pteromalidae) of Papua New Guinea: key to genera and description of new taxa, Mémoires du Muséum national d'Histoire naturelle 209, pp. 307-320 : 318-320

Trigonoderopsis enduwaensis n. sp.

(Figures 11-12)

TYPE MATERIAL — Female holotype, Papua New Guinea. PNG, Mount Wilhelm 3200m, 19- 20.10.2012, PNG MAL- MW3200 B-04/16-d04, P3357 / 8871 [on rectangular card] ( MNHN). Allotype ♂: Papua New Guinea, PNG, Mount Wilhelm 3200m, 29- 30.10.2012, PNG MAL-MW3200D-14/16-d14, P3399 / 9158 ( MNHN). Additional paratypes: 1♀, Papua New Guinea, PNG, Mount Wilhelm 3200m, 30- 31.10.2012, PNG MAL-MW3200B-15/16-d15, P3368 / 9165( MICO).

ETYMOLOGY — The specific name is derived from the word Enduwa, part of Enduwa Kombuglu, the name of Mount Wilhelm in the local Kuman language.

DIAGNOSIS — Both sexes. Body colour mainly coppery-green, with slight to distinct violet reflections; tibiae dark except yellow apices (Figures 11-12). Female. Toruli above lower margins of eyes; fl1 and fl2 approximately equal in length; fl3 as wide as pedicel; fl5-fl6 transverse. Inner eye orbits moderately diverging in lower part. Frenal area with numerous extremely fine longitudinal striae. Propodeal spiracle separated from metanotum by distinctly more than its great diameter. Gaster petiolate, petiole transverse (Figure 11). Male. Scape dark, length about 4.3× width (Figure 12). Combined length of pedicel plus flagellum about 1.3× head width.

DESCRIPTION — FEMALE: Holotype (Figure 11). Body length. 2 mm. Color. Body mainly coppery-green. Head and scutellum coppery, the latter with slight violet reflections. Eyes pale red, ocelli whitish. Mandibles yellowish-brown, teeth darker. Petiole dorsally metallic, laterally yellow. Antenna with scape mostly yellow, but darker dorsally; pedicel and flagellum dark brown. Legs with coxae brown, with slight metallic reflections, stronger on hind coxae; trochanters and femora brown, the latter lighter on extreme apices; tibiae brown except yellow apices; tarsi yellow, last tarsal segment brown. Wings hyaline; tegulae and venation brown (Figure 11). Head in frontal view with inner eye orbits moderately diverging in lower part. Toruli well above lower eye margins. Clypeal margin shallowly emarginate, lateral sides of clypeus slightly indicated. Clypeus and a narrow band reaching toruli smooth. Face mostly finely alutaceous except smooth along inner eye orbits and scrobes. Area around ocelli coarsely coriaceous-alutaceous.Vertex short, carinate behind posterior ocelli. Temple short, finely alutaceous. Gena almost smooth, malar sulcus inconspicuous. Eyes bare. Antenna (Figure 11) with scape reaching upper edge of median ocellus; both fl1 and fl2 (anelli) strongly transverse and without longitudinal sensilla, about equal in length; fl3-fl8 large, with one row of longitudinal sensilla each; fl3 slightly longer than wide; fl4-fl6 about as long as wide, almost round; fl7 and fl8 transverse; claval sutures slightly oblique due to large ventral micropilosity area. Relative measurements: head length: 23, width: 54, height: 42; POL: 12; OOL: 10; eye height: 22, length: 18; IOD: 34; eye length dorsally: 17; temple length dorsally: 3; malar space: 14.5; scape length: 22, width: 4; pedicel length: 7, width: 4; pedicel plus flagellum: 55; fl3 length: 5, width: 4; fl8 length: 4, width: 7; clava length: 15, width: 8. Mesosoma in lateral view strongly arched dorsally (Figure 11). Pronotum almost smooth; mesoscutum and axillae mostly finely transversely striateimbricate in anterior part and coriaceous in posterior part. Notauli complete and deep, their posterior ends wider apart than scutello-axillar sutures. Scutellum dorsally coarsely but finely reticulate except smooth transverse band anterior to frenal line; frenal line deep, crenulated and sinuous; frenal area not raised in the middle, with numerous extremely fine longitudinal striae; axillulae smooth. Dorsellum smooth. Propodeum smooth except strong and complete median carina, posteriorly meeting a transverse carina that borders a short adpetiolar strip; propodeal spiracle raised, broadly oval, separated from metanotum by distinctly more than its great diameter. Mesopleura with mesepisternum almost smooth; mesepimeron mainly smooth except deep oval fovea almost in middle, continued ventrally with a shallow groove, and slightly alutaceous area near anterior edge of metapleura. Metapleura smooth. Hind tibial pecten not developed. Fore wing (Figure 11) with basal cell bare, basal vein with fives setae; ventral side of costal cell with several setae distally; speculum progressively getting narrower towards stigmal vein and reaching it; ventral side of the wing with a regular row of admarginal setae behind marginal vein, and a few additional ones near base of marginal vein; disc uniformly pilose. Relative measurements: mesosomal length: 65, width: 46, height: 46; mesoscutum length: 30; scutellum length: 27, width: 29; dorsellum length: 6; propodeum length: 13; hind tibia length: 45; fore wing length: 180, width: 70; M: 54; S: 9; P: 32. Metasoma rhomboidal in dorsal view, almost flat dorsally, about as long as head plus mesosoma (Figure 11). Petiole transverse, smooth. Gt1 largest, gt2-gt4 approximately subequal in length; gt5-gt7 much shorter. Hypopygium about in the middle of gaster. Ovipositor sheaths very short, barely visible in dorsal view (Figure 11). Relative measurements: petiole length: 5, width: 8.5; gastral length (including ovipositor sheaths): 82, width: 42; ovipositor sheaths length: 11.

MALE: Allotype (Figure 12).

Differs from the female holotype mainly as follows. Body length: 1.8 mm. Scape entirely dark brown (Figure 12), length about 4.3× width.Combined length of pedicel plus flagellum about 1.3× head width. Flagellum with seven large segments and a 2-segmented clava, the latter lacking a distinct micropilosity area on ventral side (Figure 12). Metasoma gradually widening towards apex, length about 2.4× width, about as long as and narrower than mesosoma (Figure 12).

VARIATION — FEMALES. Body length: 2.0- 2.2 mm. Violet reflections on head and mesosoma from slight to very distinct.

REMARK — This is the first description of a Trigonoderopsis male. According to Bouček (1988) and Noyes (2015), there are two described species of Trigonoderopsis in the Australasian region: T. silvensis Girault, 1915 and T. petiolata Bouček, 1988 . I did not examine any of these species and thus the following differences are based on Bouček’s comments ( Bouček 1988). The females of T. enduwaensis can be separated from those of the above mentioned species as follows: from T. silvensis mainly by the petiolate gaster and higher toruli; from T. petiolata mainly by the colour of body and legs, larger size, greater length of pedicel plus flagellum compared with head width, and longer gaster. In the PNG material examined I found a female that is apparently extremely close to T. petiolata , and differing only in the shape of anelli (the first one being shorter than the second) and longer gaster. I decided to postpone its description until the examination of the T. petiolata type. A third possibly new species was found (one female and one male), but the female lacks the head, thus preventing its description. The male is close to that of T. enduwaensis , but the scape is thinner.